Classification System: APG IV
Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Lamiales
Familia: Lamiaceae
Subfamiliae: Ajugoideae – Callicarpoideae – Cymarioideae – Lamioideae – Nepetoideae – Peronematoideae – Premnoideae – Prostantheroideae – Scutellarioideae – Symphorematoideae – Tectonoideae – Viticoideae
Overview of genera
Acanthomintha – Acanthoprasium – Achyrospermum – Acrotome – Acrymia – Aegiphila – Aeollanthus – Agastache – Ajuga – Ajugoides – Alajja – Alvesia – Amasonia – Amethystea – Anisochilus – Anisomeles – Asterohyptis – Ballota – Basilicum – Benguellia – Betonica – Blephilia – Brachysola – Brazoria – Bystropogon – Callicarpa – Cantinoa – Capitanopsis – Caryopteris – Catoferia – Cedronella – Chaiturus – Chamaesphacos – Chaunostoma – Chelonopsis – Chloanthes – Cleonia – Clerodendrum – Clinopodium – Colebrookea – Coleus – Collinsonia – Colquhounia – Comanthosphace – Condea – Congea – Conradina – Cornutia – Craniotome – Cuminia – Cunila – Cyanocephalus – Cyanostegia – Cyclotrichium – Cymaria – Dauphinea – Dicerandra – Dicrastylis – Discretitheca – Dorystaechas – Dracocephalum – Drepanocaryum – Drymosiphon – Elsholtzia – Endostemon – Eplingiella – Equilabium – Eremostachys – Eriope – Eriophyton – Eriopidion – Eriothymus – Eurysolen – Fabricia – Fuerstia – Galeopsis – Garrettia – Glechoma – Glechon – Glossocarya – Gmelina – Gomphostemma – Gontscharovia – Gumira – Gymneia – Hanceola – Haplostachys – Haumaniastrum – Hedeoma – Hemiandra – Hemigenia – Hemiphora – Hemistegia – Hesperozygis – Heterolamium – Hoehnea – Holmskioldia – Holocheila – Holostylon – Horminum – Hosea – Hoslundia – Huxleya – Hymenocrater – Hymenopyramis – Hypenia – Hypogomphia – Hyptidendron – Hyptis – Hyssopus – Isodon – Isoleucas – Kalaharia – Karomia – Killickia – Kudrjaschevia – Kurzamra – Lachnostachys – Lagochilus – Lagopsis – Lallemantia – Lamium – Lavandula – Leonotis – Leonurus – Lepechinia – Leptohyptis – Leucas – Leucosceptrum – Lophanthus – Loxocalyx – Lycopus – Macbridea – Madlabium – Marmoritis – Marrubium – Marsypianthes – Martianthus – Matsumurella – Medusantha – Meehania – Melissa – Melittis – Mentha – Meriandra – Mesosphaerum – Metastachydium – Microcorys – Micromeria – Microtoena – Minthostachys – Moluccella – Monarda – Monardella – Monochilus – Mosla – Neoeplingia – Nepeta – Newcastelia – Notochaete – Obtegomeria – Ocimum – Ombrocharis – Oncinocalyx – Oocephalus – Origanum – Orthosiphon – Otostegia – Ovieda – Oxera – Panzerina – Paralamium – Paraphlomis – Paravitex – Peltodon – Pentapleura – Perilla – Perillula – Peronema – Petitia – Petraeovitex – Phlomidoschema – Phlomis – Phlomoides – Phyllostegia – Physominthe – Physopsis – Physostegia – Piloblephis – Pityrodia – Platostoma – Plectranthus – Pogogyne – Pogostemon – Poliomintha – Prasium – Premna – Prostanthera – Prunella – Pseuderemostachys – Pseudocarpidium – Pseudocaryopteris – Pseudochamaesphacos – Pseudodictamnus – Pseudomarrubium – Pycnanthemum – Pycnostachys – Renschia – Rhabdocaulon – Rhaphiodon – Rhododon – Rosmarinus – Rostrinucula – Rotheca – Roylea – Rubiteucris – Rydingia – Saccocalyx – Salvia – Satureja – Schnabelia – Scutellaria – Sideritis – Siphocranion – Solenostemon – Spartothamnella – Sphenodesme – Stachydeoma – Stachyopsis – Stachys – Stenogyne – Sulaimania – Suzukia – Symphorema – Symphostemon – Synandra – Syncolostemon – Tectona – Teijsmanniodendron – Tetradenia – Teucridium – Teucrium – Thorncroftia – Thuspeinanta – Thymbra – Thymus – Tinnea – Trichostema – Tripora – Tsoongia – Vitex – Viticipremna – Volkameria – Warnockia – Wenchengia – Westringia – Wrixonia – Zataria – Zhumeria – Ziziphora – × Clinomicromeria
Name
Lamiaceae Martinov Tekhno-Bot. Slovar: 355. (1820), nom. cons. & Labiatae Juss., Gen. Pl.: 110. (1789) nom. cons. et nom. alt.
Type genus: Lamium L. (1753)
Synonyms
Heterotypic
Aegiphilaceae Raf., Sylva Tellur.: 161 (1838)
Ajugaceae Döll, Rhein. Fl.: 375 (1843)
Chloanthaceae Hutch., Fam. Fl. Pl., ed. 2: 396 (1959)
Glechomaceae Martinov, Tekhno-Bot. Slovar: 288 (1820)
Labiatae Juss., Gen. Pl.: 110 (1789), nom. cons.
Melissaceae Bercht. & J.Presl, Prir. Rostlin: 245 (1820)
Melittidaceae Martinov, Tekhno-Bot. Slovar: 390 (1820)
Menthaceae Burnett, Outlines Bot.: 969, 1095, 1106 (1835)
Monardaceae Döll, Fl. Baden 2: 661 (1858)
Nepetaceae Bercht. & J.Presl, Prir. Rostlin: 245 (1820)
Salazariaceae F.A.Barkley, Phytologia 32: 304 (1975)
Salviaceae Bercht. & J.Presl, Prir. Rostlin: 245 (1820)
Saturejaceae Döll, Fl. Baden 2: 664. med (1858)
Scutellariaceae Döll, Rhein. Fl.: 373 (1843)
Siphonanthaceae Raf., Fl. Tellur. 4: 87 (1838)
Stachydaceae Döll, Rhein. Fl.: 366 (1843)
Symphoremataceae Wight, Icon. Pl. Ind. Orient. 4(3): 13 (1849)
Viticaceae Juss., Gen. Pl.: 106 (1789)
References
Primary references
Martinov, I.I. 1820. Техно-ботаническiй словарь, на латинскомъ и россiйскомъ языкахъ, составленный […]. [Techno-Botanical Dictionary, in Latin and Russian, compiled by [...]]. i-vi, 682 p. St.-Petersburg: v tipografii Imperatorskoj Rossijskoj Akademii [Nauk] (Russian) DOI: 10.5962/bhl.title.96260 Reference page.
Additional references
Harley, R.M., Paton, A.J. & Ryding, O. 2003. New synonymy and taxonomic changes in the Labiatae. Kew Bulletin 58(2): 485-489. DOI: 10.2307/4120633 Paywall ResearchGate Reference page.
Harley, R.M., Atkins, S., Budantsev, A.L., Cantino, P.D., Conn, B.J., Grayer, M., Harley, M.M., De Kok, R., Krestovskaya, T., Moralaes, R., Paton, A.J., Ryding, O. & Upson, T. 2004. Labiatae. Pp. 167–275, in Kadereit, J. (ed.), The Families and Genera of Vascular Plants. VII. Flowering Plants: Dicotyledons. Lamiales (except Acanthaceae including Avicenniaceae). Springer, Berlin.
Li, B., Cantino, P.D., Olmstead, R.G., Bramley, G.L.C., Xiang, C.L., Ma, Z.H., Tan, Y.H. & Zhang, D.X. 2016. A large-scale chloroplast phylogeny of the Lamiaceae sheds new light on its subfamilial classification. Scientific Reports 6: 34343. DOI: 10.1038/srep34343 Open access Reference page.
Li, B.O. & Olmstead, R.G. 2017. Two new subfamilies in Lamiaceae. Phytotaxa 313(2): 222-226. DOI: 10.11646/phytotaxa.313.2.9 Paywall ResearchGate Reference page.
Olmstead, R.G. (ed.) 2016. A Synoptical Classification of the Lamiales. Version 2.6.2 (in prog.). 20 pp. PDF (Internet Archive) Reference page. Accessed 15 Jan. 2016.
Paton, A.J., Mwanyambo, M., Govaerts, R.H., Smitha, K., Suddee, S., Phillipson, P.B., Wilson, T.C., Forster, P.I. & Culham, A. 2019. Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera. PhytoKeys 129: 1–158. DOI: 10.3897/phytokeys.129.34988 Open access Reference page.
Zhao, F., Chen, Y.P., Salmaki, Y., Drew, B.T., Wilson, T.C., Scheen, A.C., Celep, F., Bräuchler, C., Bendiksby, M., Wang, Q., Min, D.Z., Peng, H., Olmstead, R.G., Li, B. & Xiang, C.L. 2021. An updated tribal classification of Lamiaceae based on plastome phylogenomics. BMC Biology 19(1): 1-27. DOI: 10.1186/s12915-020-00931-z Open access Reference page.
Links
Govaerts, R. et al. 2022. Lamiaceae in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2022 Mar 14. Reference page.
Hassler, M. 2022. Lamiaceae. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2022. Species 2000 & ITIS Catalogue of Life. Published online. Accessed: 2022 Mar 14. Reference page.
Tropicos.org 2022. Lamiaceae. Missouri Botanical Garden. Published online. Accessed: 14 Mar 2022.
International Plant Names Index. 2022. Lamiaceae. Published online. Accessed: Mar 14 2022.
Stevens, P.F. 2001 onwards. Angiosperm Phylogeny Website. Version 14, July 2017 [and more or less continuously updated since]. Online. Reference page. Accessed 7 June 2014.
Vernacular names
العربية: شفوية
azərbaycanca: Dalamazkimilər
башҡортса: Һаңғырау кесерткән һымаҡтар
беларуская: Ясноткавыя
български: Устноцветни
català: Lamiàcies
čeština: Hluchavkovité
dansk: Læbeblomst-familien
Deutsch: Lippenblütler
ދިވެހިބަސް: ކުދީނާ
Ελληνικά: Χειλανθή
English: Mint family
Esperanto: Lamiacoj
español: Lamiáceas
eesti: Huulõielised
فارسی: نعنائیان
suomi: Huulikukkaiskasvit
Nordfriisk: Lapbloosplaanten
français: Lamiacées, Labiacées, Labiées
galego: Lamiáceas
עברית: שפתניים
hrvatski: Usnače
hornjoserbsce: Cycawkowe rostliny
magyar: Árvacsalánfélék, Ajakosak, Ajakosvirágúak
հայերեն: Շրթնածաղիկներ, Խուլեղինջազգիներ, շրթնածաղկավորներ
íslenska: Varablómaætt
italiano: Lamiacee, Labiate
日本語: シソ科
ქართული: ტუჩოსანნი
қазақша: Ерінгүлділер
ಕನ್ನಡ: ಲಾಮಿಯೇಸಿ
한국어: 꿀풀과
kurdî: Famîleya gezika derewîn
кыргызча: Эрин гүлдүүлөр тукуму
lietuvių: Notreliniai augalai
latviešu: Lūpziežu dzimta
македонски: Усноцветни
മലയാളം: ലാമിയേസി
Nederlands: Lipbloemenfamilie
norsk nynorsk: Lepeblomsterfamilien
norsk: Leppeblomstfamilien
Diné bizaad: Azeeniłchin
ирон: Былдидинон зайæгойтæ
polski: Jasnotowate
پنجابی: تلسی ٹبر
پښتو: د نعنا بوټي
Runa Simi: Muña yura rikch'aq ayllu
русский: Яснотковые, Губоцветные
slovenčina: Hluchavkovité
slovenščina: Ustnatice
српски / srpski: Уснатице
svenska: Kransblommiga växter
తెలుగు: లామియేసి
ไทย: วงศ์กะเพรา
Türkçe: Ballıbabagiller
українська: Глухокропивові
اردو: ریمانیہ
oʻzbekcha/ўзбекча: Yalpizdoshlar
Tiếng Việt: Họ Hoa môi
walon: Polurêyes
中文: 唇形科
The Lamiaceae (/ˌleɪmiˈeɪsiaɪ, -iː/ LAY-mee-AY-see-eye, -ee)[3] or Labiatae are a family of flowering plants commonly known as the mint or deadnettle or sage family. Many of the plants are aromatic in all parts and include widely used culinary herbs like basil, mint, rosemary, sage, savory, marjoram, oregano, hyssop, thyme, lavender, and perilla, as well as other medicinal herbs such as catnip, salvia, bee balm, wild dagga, and oriental motherwort. Some species are shrubs, trees (such as teak), or, rarely, vines. Many members of the family are widely cultivated, not only for their aromatic qualities, but also their ease of cultivation, since they are readily propagated by stem cuttings. Besides those grown for their edible leaves, some are grown for decorative foliage. Others are grown for seed, such as Salvia hispanica (chia), or for their edible tubers, such as Plectranthus edulis, Plectranthus esculentus, Plectranthus rotundifolius, and Stachys affinis (Chinese artichoke). Many are also grown ornamentally, notably coleus, Plectranthus, and many Salvia species and hybrids.
The family has a cosmopolitan distribution.[4] The enlarged Lamiaceae contain about 236 genera[5] and have been stated to contain 6,900[4] to 7,200[5] species, but the World Checklist lists 7,534.[6] The largest genera are Salvia (900), Scutellaria (360), Stachys (300), Plectranthus (300), Hyptis (280), Teucrium (250), Vitex (250), Thymus (220), and Nepeta (200).[5] Clerodendrum was once a genus of over 400 species,[5] but by 2010, it had been narrowed to about 150.[7]
The family has traditionally been considered closely related to the Verbenaceae;[5] in the 1990s, phylogenetic studies suggested that many genera classified in the Verbenaceae should be classified in the Lamiaceae [8][9] or to other families in the order Lamiales.[1]
Salvia jurisicii
The alternative family name Labiatae refers to the flowers typically having petals fused into an upper lip and a lower lip (labia in Latin). The flowers are bilaterally symmetrical with five united petals and five united sepals. They are usually bisexual and verticillastrate (a flower cluster that looks like a whorl of flowers, but actually consists of two crowded clusters). Although this is still considered an acceptable alternative name, most botanists now use the name Lamiaceae in referring to this family. The leaves emerge oppositely, each pair at right angles to the previous one (decussate) or whorled. The stems are frequently square in cross section,[10] but this is not found in all members of the family, and is sometimes found in other plant families.
Genera
The last revision of the entire family was published in 2004.[5] It described and provided keys to 236 genera. These are marked with an asterisk (*) in the list below. A few genera have been established or resurrected since 2004. These are marked with a plus sign (+). Other genera have been synonymised. These are marked with a minus sign (-). The remaining genera in the list are mostly of historical interest only and are from a source that includes such genera without explanation.[11] Few of these are recognized in modern treatments of the family.
Kew Gardens provides a list of genera that includes additional information.[12] A list at the Angiosperm Phylogeny Website is frequently updated.[13]
*Acanthomintha
*Achyrospermum
Acinos
Acrocephalus
*Acrotome
*Acrymia
Adelosa
*Aegiphila
*Aeollanthus
*Agastache
*Ajuga
*Ajugoides
*Alajja
*Alvesia
*Amasonia
*Amethystea
*Anisochilus
*Anisomeles
*Asterohyptis
*Ballota
*Basilicum
Becium
*Benguellia
*Blephilia
*Bostrychanthera
Bovonia
*Brachysola
*Brazoria
*Bystropogon
Calamintha
*Callicarpa
*Capitanopsis
Capitanya
*Caryopteris
*Catoferia
*Cedronella
Ceratanthus
*Chaiturus
*Chamaesphacos
*Chaunostoma
*Chelonopsis
*Chloanthes
*Cleonia
*Clerodendrum
*Clinopodium
*Colebrookea
*Collinsonia
*Colquhounia
*Comanthosphace
*Congea
*Conradina
Coridothymus
*Cornutia
*Craniotome
*Cryphia
*Cuminia
*Cunila
*Cyanostegia
*Cyclotrichium
*Cymaria
*Dauphinea
*Dicerandra
*Dicrastylis
Discretitheca
Dorystoechas
*Dracocephalum
*Drepanocaryum
*Elsholtzia
*Endostemon
Englerastrum
+Eplingiella
*Eremostachys
*Eriope
*Eriophyton
Eriopidion
*Eriothymus
Erythrochlamys
Euhesperida
*Eurysolen
*Faradaya
*Fuerstia
*Galeopsis
*Garrettia
Geniosporum
*Glechoma
*Glechon
*Glossocarya
*Gmelina
*Gomphostemma
*Gontscharovia
*Hanceola
*Haplostachys
*Haumaniastrum
*Hedeoma
*Hemiandra
*Hemigenia
*Hemiphora
*Hemizygia
*Hesperozygis
*Heterolamium
*Hoehnea
*Holmskioldia
*Holocheila
Holostylon
*Horminum
*Hosea
*Hoslundia
*Huxleya
*Hymenocrater
*Hymenopyramis
*Hypenia
*Hypogomphia
*Hyptidendron
*Hyptis
*Hyssopus
Isodictyophorus
*Isodon
*Isoleucas
+Kalaharia
*Karomia
Keiskea
Killickia
Kudrjaschevia
*Kurzamra
*Lachnostachys
*Lagochilus
*Lagopsis
*Lallemantia
*Lamiophlomis
*Lamium
*Lavandula
*Leocus
*Leonotis
*Leonurus
*Lepechinia
*Leucas
Leucophae
*Leucosceptrum
Limniboza
*Lophanthus
*Loxocalyx
*Lycopus
*Macbridea
*Madlabium
*Marmoritis
+Martianthus
*Marrubium
*Marsypianthes
*Matsumurella
*Meehania
*Melissa
*Melittis
*Mentha
*Meriandra
Mesona
*Metastachydium
*Microcorys
*Micromeria
*Microtoena
*Minthostachys
*Moluccella
*Monarda
*Monardella
*Monochilus
*Mosla
*Neoeplingia
Neohyptis
Neorapinia
*Nepeta
*Newcastelia
Nosema
*Notochaete
*Obtegomeria
*Ocimum
Octomeron
*Ombrocharis
*Oncinocalyx
*Origanum
*Orthosiphon
*Otostegia
+Ovieda
*Oxera
*Panzerina
*Paralamium
*Paraphlomis
*Paravitex
*Peltodon
*Pentapleura
*Perilla
*Perillula
*Peronema
-Perovskia
Perrierastrum
Petitia
*Petraeovitex
*Phlomidoschema
*Phlomis
*Phlomoides
*Phyllostegia
*Physopsis
*Physostegia
*Piloblephis
Pitardia
*Pityrodia
*Platostoma
*Plectranthus
*Pogogyne
*Pogostemon
*Poliomintha
*Prasium
*Premna
*Prostanthera
*Prunella
*Pseuderemostachys
*Pseudocarpidium
*Pseudocaryopteris
*Pseudomarrubium
Puntia
*Pycnanthemum
*Pycnostachys
Rabdosiella
*Renschia
*Rhabdocaulon
*Rhaphiodon
*Rhododon
-Rosmarinus
*Rostrinucula
*Rotheca
*Roylea
*Rubiteucris
+Rydingia
Sabaudia
*Saccocalyx
Salazaria
*Salvia
*Satureja
*Schizonepeta
*Schnabelia
*Scutellaria
*Sideritis
*Siphocranion
Solenostemon
*Spartothamnella
*Sphenodesme
*Stachydeoma
*Stachyopsis
*Stachys
*Stenogyne
*Sulaimania
*Suzukia
*Symphorema
Symphostemon
*Synandra
*Syncolostemon
*Tectona
*Teijsmanniodendron
+Tetraclea
*Tetradenia
*Teucridium
*Teucrium
*Thorncroftia
*Thuspeinanta
*Thymbra
*Thymus
*Tinnea
*Trichostema
*Tripora
*Tsoongia
*Vitex
*Viticipremna
+Volkameria
*Warnockia
*Wenchengia
*Westringia
Wiedemannia
*Wrixonia
Xenopoma
*Zataria
*Zhumeria
*Ziziphora
Recent changes
The circumscription of several genera has changed since 2004. Tsoongia, Paravitex, and Viticipremna have been sunk into synonymy with Vitex.[14] Huxleya has been sunk into Volkameria.[7] Kalaharia, Volkameria, Ovieda, and Tetraclea have been segregated from a formerly polyphyletic Clerodendrum.[7] Rydingia has been separated from Leucas.[15] The remaining Leucas is paraphyletic over four other genera.[16]
Subfamilies and tribes
In 2004, the Lamiaceae were divided into seven subfamilies, plus 10 genera not placed in any of the subfamilies.[5] The unplaced genera are: Tectona, Callicarpa, Hymenopyramis, Petraeovitex, Peronema, Garrettia, Cymaria, Acrymia, Holocheila, and Ombrocharis. The subfamilies are the Symphorematoideae, Viticoideae, Ajugoideae, Prostantheroideae, Nepetoideae, Scutellarioideae, and Lamioideae. The subfamily Viticoideae is probably not monophyletic.[14] The Prostantheroideae and Nepetoideae are divided into tribes. These are shown in the phylogenetic tree below.
Phylogeny
Most of the genera of Lamiaceae have never been sampled for DNA for molecular phylogenetic studies.[17] Most of those that have been are included in the following phylogenetic tree. The phylogeny depicted below is based on seven different sources.[5][9][7][14][18][19][20]
| Lamiaceae |
|
References
Stevens, P. F. (July 2012). "Lamiales (Lamiaceae Family)". Angiosperm Phylogeny Website. Archived from the original on 21 March 2015. Retrieved 25 March 2015.
Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III". Botanical Journal of the Linnean Society. 161 (2): 105–121. doi:10.1111/j.1095-8339.2009.00996.x.
"Pronunciation of lamiaceae". Archived from the original on 2015-09-24. Retrieved 2014-11-25.
Heywood, Vernon H.; Brummitt, Richard K.; Seberg, Ole; Culham, Alastair (2007). Flowering Plant Families of the World. Ontario, Canada: Firefly Books. ISBN 978-1-55407-206-4.
Raymond M. Harley, Sandy Atkins, Andrey L. Budantsev, Philip D. Cantino, Barry J. Conn, Renée J. Grayer, Madeline M. Harley, Rogier P.J. de Kok, Tatyana V. Krestovskaja, Ramón Morales, Alan J. Paton, and P. Olof Ryding. 2004. "Labiatae" pages 167-275. In: Klaus Kubitzki (editor) and Joachim W. Kadereit (volume editor). The Families and Genera of Vascular Plants volume VII. Springer-Verlag: Berlin; Heidelberg, Germany. ISBN 978-3-540-40593-1
"World Checklist of Selected Plant Families". Archived from the original on 2017-07-09. Retrieved 2022-02-21.
Yuan, Yao-Wu; Mabberley, David J.; Steane, Dorothy A.; Olmstead, Richard G. (2010). "Further disintegration and redefinition of Clerodendrum (Lamiaceae): Implications for the understanding of the evolution of an intriguing breeding strategy". Taxon. 59 (1): 125–133. doi:10.1002/tax.591013.
Cantino, P.D., Harley, R.M. & Wagstaff, S.J. 1992. Genera of Labiatae: status and classification. Pp. 511-522. In: Raymond M. Harley and Tom Reynolds (editors). Advances in Labiate Science. Richmond, Royal Botanic Gardens, Kew.
Wagstaff, Steven J.; Hickerson, Laura; Spangler, Russ; Reeves, Patrick A.; Olmstead, Richard G. (1998). "Phylogeny in Labiatae s.l., inferred from cpDNA sequences". Plant Systematics and Evolution. 209 (3–4): 265–274. doi:10.1007/bf00985232. S2CID 601658.
Parnell, J. and Curtis, T. 2012. Webb's An Irish Flora. Cork University Press ISBN 978-185918-4783
"List of genera in Lamiaceae". In: "Lamiaceae". In: "List of families". In: "Families and genera in GRIN. (see External links below)
List of Genera in Lamiaceae. At: Vascular Plant Families and Genera. At: World Checklist of Selected Plant Families. At: Electronic Plant Information Center. At: Website of Royal Botanic Gardens, Kew. (see External Links below).
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