Coccophagus

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Hymenopterida
Ordo: Hymenoptera
Subordo: Apocrita
Superfamilia: Chalcidoidea

Familia: Aphelinidae
Subfamilia: Coccophaginae
Genus: Coccophagus
Species (262):
A

C. acanthosceles – C. adumbratus – C. adustus – C. aethiopis – C. aethochreus – C. afrangiatus – C. africanus – C. albiapicella – C. albicoxa – C. albifuniculatus – C. albusus – C. amblydon – C. anchoroides – C. angolensis – C. anthracinus – C. apricus – C. argenteus – C. argentifascia – C. argentiscutellum – C. argocoxa – C. ashmeadi – C. assamensis – C. asterolecanii – C. aterrimus – C. atratus – C. aurantifrons – C. aureonotus – C. auricaput – C. avetianae –
B

C. baldassarii – C. bartletti – C. basalis – C. berzeliae – C. biguttatus – C. bimaculatus – C. bivittatus – C. bogoriensis – C. brachypterus – C. brasiliensis – C. brethesi – [[Coccophagus breviclavulus {{{3}}}|C. {{{3}}}]] – C. breviscapus – C. brevisetus – C. brunneus – C. burksi –
C

C. candidus – C. caophongi – C. capensis – C. caridei – C. catherinae – C. caudatus – C. ceroplastae – C. chaetosus – C. chengtuensis – C. chilensis – C. chloropulvinariae – C. cinguliventris – C. clavatus – C. clavellatus – C. coccidarum – C. comperei – C. concinnus – C. conditus – C. cooperatus – C. copernicus – C. coracinus – C. cowperi – C. crenatus – C. croconotus – C. crucigerus – C. cryptus – C. cubaensis –
D

C. debachi – C. desantisi – C. desertus – C. diabolicus – C. diachraceus – C. differens – C. dilatatus – C. diminutus – C. distinctus – C. dius –
E

C. eleaphilus – C. emersoni – C. equifuniculatus – C. eusaissetiae – C. euxanthodes – C. excelsus – C. exiguiventris –
F

C. fallax – C. fasciatus – C. femoralis – C. flavescens – C. flaviceps – C. flavicorpus – C. flavidus – C. flavifrons – C. flavoscutellum – C. fletcheri – C. formicariae – C. fraternus – C. fumadus – C. funeralis –
G

C. gahani – C. ghesquierei – C. gigas – C. gilvus – C. gondolae – C. gonzalezi – C. gossypariae – C. graminis – C. gregarius – C. grenadensis – C. gurneyi –
H

C. hanoiensis – C. hawaiiensis – C. hemera – C. hibisci – C. hispaniolae –
I

C. immaculatus – C. impensus – C. indochraceus – C. inkermani – C. insidiator – C. insignis – C. ishiii – C. isipingoensis –
J

C. japonicus – C. jasnoshae – C. javensis –
K

C. kabulensis – C. kvavadze –
L

C. lepidus – C. leptospermi – C. lii – C. longiclavatus – C. longicornis – C. longifasciatus – C. longipedicellus – C. lucani – C. lucidus – C. luciensis – C. lunai – C. lutescens – C. lycimnia –
M

C. maculipennis – C. malthusi – C. mangiferae – C. margaritatus – C. mariformis – C. matsuyamensis – C. mazatlan – C. meghaianus – C. merceti – C. mexicanus – C. mexicensis – C. minor – C. mixtus – C. modestus – C. multisetae –
N

C. narendrani – C. neocomperei – C. neserorum – C. nigrans – C. nigricorpus – C. nigricoxae – C. nigritus – C. nigropleurum – C. nipponicus – C. nubeculus – C. nubes – C. nympha –
O

C. obscurus – C. ochraceus – C. oculatipennis – C. ophicus –
P

C. palaeolecanii – C. pallidiceps – C. pallidis – C. parlobatae – C. pellucidus – C. perflavus – C. perhispidus – C. perlucidus – C. pernigritus – C. philippiae – C. physokermis – C. piceae – C. pisinnus – C. planus – C. poei – C. princeps – C. prinslooi – C. probus – C. propodealis – C. provisus – C. proximus – C. pseudococci – C. pseudopulvinariae – C. pulchellus – C. pulcher – C. pulcini – C. pulvinariae – C. pumilus – C. purpureus –
Q

C. qenai – C. quaestor –
R

C. redini – C. restionis – C. rjabovi – C. robustus – C. rosae – C. ruizi – C. rusti –
S

C. saintebeauvei – C. saissetiae – C. saltator – C. samarae – C. scutatus – C. scutellaris – C. secamonei – C. semiatratus – C. semicircularis – C. sexvittatus – C. shafeei – C. shakespearella – C. shillongensis – C. sibiricus – C. signatus – C. signus – C. silvestrii – C. sostenesi – C. spartanus – C. specialis – C. speciosus – C. spectabilis – C. spireae – C. srilankensis – C. subflavescens – C. subochraceus – C. subsignus – C. sudhiri –
T

C. tarongaensis – C. teeceeni – C. tenebrisetus – C. terani – C. tetrastichoides – C. thanhoaensis – C. tibialis – C. timberlakei – C. tobiasi – C. triangulatinotus – C. triguttatus – C. tropicanus – C. tschirchii –
U

C. ussuriensis –
V

C. varius – C. vegai – C. viator – C. vietnamicus –
Y

C. yoshidae – C. youngi –
Z

C. zebratus – C. zeyai – C. zinniae
{{Just|Checklist based uncritically on UCD, with addition of C. minor
Check (5): C. argiope – C. clypeator – C. dentatus – C. falderus – C. longisetus
Name

Coccophagus Westwood, 1833: 344

Type species: Entedon scutellaris Dalman
Fixation: subsequent designation by Westwood, 1839

Synonyms

Euxanthellus Silvestri, 1915

References

Westwood, J.O. 1833: Descriptions of several new British forms amongst the parasitic hymenopterous insects. London and Edinburgh Philosophical Magazine and Journal of Science (3) 3: 342-344. BHL Reference page.
Chen, Y. & Li, C-D. 2017. Three new species of Coccophagus (Hymenoptera: Aphelinidae) from China, with new distributional data for three additional species. Zootaxa 4294(2): 256–270. DOI: 10.11646/zootaxa.4294.2.8. Reference page.
Compere, H. 1931: A revision of the species of Coccophagus, a genus of hymenopterous, coccid-inhabiting parasites. Proceedings of the United States National Museum, 78(2850): 1–132. BHL Reference page.
Qin, Y-G., Chen, H-F., Li, C-D. & Chen, Y. 2022. On the genus Coccophagus Westwood (Hymenoptera, Aphelinidae) from Xishuangbanna Rainforest. Contribution I: Two new species of the Coccophagus varius group, with an identification key and phylogenetic analysis. Zookeys 1091ː 119–138. DOI: 10.3897/zookeys.1091.80065 Open access Reference page.

Coccophagus is a large genus of chalcid wasps belonging to the family Aphelinidae.
Species

Coccophagus contains the following species:[1]

Coccophagus ablusus Annecke and Insley, 1974
Coccophagus acanthosceles Waterston, 1916
Coccophagus adumbratus Annecke and Insley, 1970
Coccophagus adustus (Annecke and Prinsloo, 1976)
Coccophagus aethiopis Girault, 1935
Coccophagus aethochreus Annecke and Insley, 1974
Coccophagus afrangiatus Viggiani, 1994
Coccophagus africanus Risbec, 1956
Coccophagus albiapicella De Santis, 1996
Coccophagus albicoxa Howard, 1911
Coccophagus albifuniculatus (Huang, 1994)
Coccophagus amblydon Compere, 1937
Coccophagus amoenus Hayat, 2015
Coccophagus anchoroides (Huang, 1994)
Coccophagus angolensis Annecke and Insley, 1974
Coccophagus anthracinus Compere, 1925
Coccophagus apricus Annecke and Insley, 1970
Coccophagus argenteus Girault, 1915
Coccophagus argentifascia Girault, 1915
Coccophagus argentiscutellum (Girault, 1915)
Coccophagus argocoxa Annecke and Insley, 1974
Coccophagus assamensis Hayat, 1993
Coccophagus asterolecanii (Dozier, 1932)
Coccophagus atratus Compere, 1926
Coccophagus aurantifrons (Compere, 1936)
Coccophagus aureonotus (Howard, 1897)
Coccophagus auricaput Girault, 1915
Coccophagus avetianae Yasnosh and Herthevtzian, 1972
Coccophagus baldassarii Compere, 1931
Coccophagus bartletti Annecke and Insley, 1974
Coccophagus basalis Compere, 1939
Coccophagus berzeliae Annecke and Insley, 1974
Coccophagus biguttatus Girault, 1915
Coccophagus bimaculatus Myartseva, 2004
Coccophagus bivittatus Compere, 1931
Coccophagus bogoriensis (Koningsberger, 1897)
Coccophagus brachypterus Sugonjaev, 2011
Coccophagus brasiliensis (Compere, 1936)
Coccophagus brethesi De Santis, 1967
Coccophagus breviscapus Li and Chen, 2017
Coccophagus brevisetus Huang, 1980
Coccophagus brunneus Provancher, 1887
Coccophagus burksi Hayat, 1971
Coccophagus candidus Hayat, 1993
Coccophagus caophongi Sugonjaev, 1996
Coccophagus capensis Compere, 1931
Coccophagus caridei (Brethes, 1918)
Coccophagus catherinae Annecke, 1964
Coccophagus caudatus (Huang, 1994)
Coccophagus ceroplastae (Howard, 1895)
Coccophagus chaetosus Sugonjaev, 1995
Coccophagus chengtuensis Sugonjaev and Peng, 1960
Coccophagus chilensis De Santis, 1988
Coccophagus chloropulvinariae Hayat, 1974
Coccophagus cinguliventris Girault, 1909
Coccophagus clavatus Husain and Agarwal, 1982
Coccophagus clavellatus Compere, 1931
Coccophagus coccidarum (Ghesquiere, 1949)
Coccophagus comperei Mercet, 1932
Coccophagus concinnus De Santis, 1963
Coccophagus conditus Annecke and Insley, 1974
Coccophagus cooperatus Sugonjaev and Ren, 1993
Coccophagus copernicus Husain and Agarwal, 1982
Coccophagus coracinus Compere, 1940
Coccophagus cowperi Girault, 1917
Coccophagus crenatus Huang, 1980
Coccophagus croconotus (Waterston, 1917)
Coccophagus crucigerus Girault, 1931
Coccophagus cryptus Annecke and Insley, 1974
Coccophagus cubaensis Compere, 1931
Coccophagus debachi Myartseva and Ruiz, 2005
Coccophagus desantisi (Fidalgo, 1981)
Coccophagus desertus Sugonjaev and Myartseva, 1984
Coccophagus diabolicus (Girault, 1915)
Coccophagus diachraceus Annecke and Insley, 1974
Coccophagus differens Yasnosh, 1966
Coccophagus dilatatus (Huang, 1994)
Coccophagus diminutus Sugonjaev, 1995
Coccophagus distinctus Li and Chen, 2017
Coccophagus dius Hayat, 1998
Coccophagus divisus Hayat, 2015
Coccophagus eleaphilus Silvestri, 1915
Coccophagus emersoni Girault, 1917
Coccophagus equifuniculatus (Huang, 1994)
Coccophagus eusaissetiae Ozdikmen, 2011
Coccophagus euxanthodes Hayat, Schroer and Pemberton, 2009
Coccophagus excelsus Erdos, 1956
Coccophagus exiguiventris Girault, 1929
Coccophagus fallax Compere, 1939
Coccophagus fasciatus Annecke and Insley, 1970
Coccophagus femoralis Myartseva, 2006
Coccophagus flavescens Howard, 1896
Coccophagus flaviceps Compere, 1931
Coccophagus flavicorpus Husain and Agarwal, 1982
Coccophagus flavidus Compere, 1940
Coccophagus flavifrons Howard, 1885
Coccophagus flavoscutellum Ashmead, 1881
Coccophagus fletcheri Howard, 1897
Coccophagus fraternus Howard, 1881
Coccophagus fumadus Hayat, 2010
Coccophagus funeralis Girault, 1913
Coccophagus funiculatus Mysartseva, 2016
Coccophagus gahani Annecke and Insley, 1974
Coccophagus ghesquierei Hayat, 1987
Coccophagus gigas Erdos, 1956
Coccophagus gilvus Hayat, 1971
Coccophagus gondolae (Castel-Branco, 1951)
Coccophagus gonzalezi Myartseva, 2006
Coccophagus gossypariae Gahan, 1927
Coccophagus graminis Annecke and Insley, 1974
Coccophagus gregarius Compere, 1931
Coccophagus grenadensis Hayat, 1994
Coccophagus gurneyi Compere, 1929
Coccophagus hanoiensis Sugonjaev, 1996
Coccophagus hawaiiensis Timberlake, 1926
Coccophagus hemera (Walker, 1839)
Coccophagus hibisci Sugonjaev and Ren, 1993
Coccophagus hispaniolae (Dozier, 1932)
Coccophagus immaculatus Howard, 1881
Coccophagus impensus Annecke and Insley, 1974
Coccophagus indefinitus Myartseva, 2014
Coccophagus indochraceus Hayat, 1998
Coccophagus inkermani Girault, 1926
Coccophagus insidiator (Dalman, 1826)
Coccophagus insignis Hayat and Zeya, 1993
Coccophagus ishiii Compere, 1931
Coccophagus isipingoensis Compere, 1931
Coccophagus japonicus Compere, 1924
Coccophagus jasnoshae Sugonjaev, 1978
Coccophagus javensis Girault, 1916
Coccophagus kabulensis Sugonjaev, 1985
Coccophagus kvavadze Japoshvili and Karaca, 2002
Coccophagus lepidus Compere, 1931
Coccophagus leptospermi Girault, 1917
Coccophagus lii (Huang, 1994)
Coccophagus longiclavatus Shafee, 1972
Coccophagus longicornis Hayat, 1971
Coccophagus longifasciatus Howard, 1907
Coccophagus longipedicellus Shafee, 1972
Coccophagus lucani Girault, 1922
Coccophagus lucidus Ishihara, 1977
Coccophagus luciensis Annecke and Insley, 1974
Coccophagus lunai Myartseva, 2006
Coccophagus lutescens Compere, 1931
Coccophagus lycimnia (Walker, 1839)
Coccophagus maculipennis Yasnosh, 1966
Coccophagus malthusi Girault, 1917
Coccophagus mangiferae (Dozier, 1932)
Coccophagus margaritatus Compere, 1931
Coccophagus mariformis Compere, 1931
Coccophagus matsuyamensis Ishihara, 1977
Coccophagus mazatlan Myartseva, 2006
Coccophagus meghaianus Hayat, 2010
Coccophagus merceti Hayat, 1994
Coccophagus mexicanus Girault, 1915
Coccophagus mexicensis Girault, 1917
Coccophagus minor Myartseva, 2014
Coccophagus mixtus (Girault, 1915)
Coccophagus modestus Silvestri, 1915
Coccophagus multisetae Girault, 1931
Coccophagus narendrani Hayat and Zeya, 1993
Coccophagus neocomperei Myartseva and Ruiz, 2005
Coccophagus neserorum (Annecke and Mynhardt, 1979)
Coccophagus nesiotes Hayat, 2015
Coccophagus nigrans Myartseva, 2006
Coccophagus nigricorpus Shafee, 1972
Coccophagus nigricoxae Karam and Ramadan, 2013
Coccophagus nigritus Compere, 1931
Coccophagus nigropleurum Girault, 1917
Coccophagus nipponicus (Ishihara, 1977)
Coccophagus nubeculus Brethes, 1913
Coccophagus nubes Compere, 1928
Coccophagus nympha (Girault, 1915)
Coccophagus obscurus Westwood, 1833
Coccophagus ochraceus Howard, 1895
Coccophagus oculatipennis (Girault, 1916)
Coccophagus ophicus Husain and Agarwal, 1982
Coccophagus palaeolecanii Yasnosh, 1957
Coccophagus pallidiceps (Compere, 1939)
Coccophagus pallidis Huang, 1980
Coccophagus parlobatae Hayat, 2007
Coccophagus pellucidus (Huang, 1994)
Coccophagus perflavus Girault, 1916
Coccophagus perhispidus Girault, 1926
Coccophagus pernigritus De Santis, 1948
Coccophagus philippiae (Silvestri, 1915)
Coccophagus physokermis Sugonjaev and Pilipyuk, 1972
Coccophagus piceae Erdos, 1956
Coccophagus pisinnus Annecke and Insley, 1974
Coccophagus planus (Sugonjaev, 1969)
Coccophagus poei Girault, 1915
Coccophagus princeps Silvestri, 1915
Coccophagus prinslooi Hayat, 1998
Coccophagus probus Annecke and Mynhardt, 1979
Coccophagus propodealis Myartseva, 2004
Coccophagus provisus Sugonjaev and Ren, 1993
Coccophagus proximus Yasnosh, 1966
Coccophagus pseudococci Compere, 1933
Coccophagus pseudopulvinariae Li, 1996
Coccophagus pulchellus Westwood, 1833
Coccophagus pulcher (Girault, 1914)
Coccophagus pulcini Girault, 1926
Coccophagus pulvinariae Compere, 1931
Coccophagus pumilus Sugonjaev, 1994
Coccophagus purpureus Ashmead, 1886
Coccophagus qenai Abd-Rabou, 2003
Coccophagus quaestor Girault, 1917
Coccophagus redini Girault, 1924
Coccophagus restionis Annecke and Insley, 1970
Coccophagus rjabovi Yasnosh, 1966
Coccophagus robustus Compere, 1931
Coccophagus rosae Sugonjaev and Pilipyuk, 1972
Coccophagus ruizi Myartseva, 2004
Coccophagus rusti Compere, 1928
Coccophagus saintebeauvei Girault, 1917
Coccophagus saissetiae Gahan, 1922
Coccophagus saltator Sugonjaev, 1994
Coccophagus samarae Hayat, 1998
Coccophagus scutatus Howard, 1895
Coccophagus scutellaris (Dalman, 1826)
Coccophagus secamonei Risbec, 1957
Coccophagus semiatratus De Santis, 1947
Coccophagus semicircularis (Foerster, 1841)
Coccophagus sexvittatus Hayat, 1974
Coccophagus shafeei Hayat, 1974
Coccophagus shakespearella Girault, 1929
Coccophagus shillongensis Hayat and Singh, 1989
Coccophagus sibiricus Sugonjaev, 1976
Coccophagus signatus Yasnosh, 1966
Coccophagus signus Girault, 1920
Coccophagus silvestrii Compere, 1931
Coccophagus sostenesi Myartseva, 2006
Coccophagus spartanus Japoshvili and Karaca, 2002
Coccophagus specialis Compere, 1931
Coccophagus speciosus Compere, 1931
Coccophagus spectabilis Compere, 1931
Coccophagus spireae Nikolskaya and Yasnosh, 1966
Coccophagus srilankensis Hayat, 1988
Coccophagus subflavescens Hayat, 1971
Coccophagus subochraceus Howard, 1907
Coccophagus subsignus Girault, 1932
Coccophagus sudhiri Hayat, 1993
Coccophagus tamaulipecus Myartseva, 2014
Coccophagus tarongaensis Compere, 1931
Coccophagus teeceeni Myartseva, 2004
Coccophagus tenebrisetus Li and Chen, 2017
Coccophagus terani De Santis, 1993
Coccophagus tetrastichoides Sugonjaev, 2006
Coccophagus thanhoaensis Sugonjaev, 2011
Coccophagus tibialis Compere, 1931
Coccophagus timberlakei Compere, 1931
Coccophagus tobiasi Myartseva, 2004
Coccophagus triangulatinotus Girault, 1926
Coccophagus triguttatus Girault, 1915
Coccophagus tropicanus Sugonjaev and Ren, 1993
Coccophagus tschirchii Mahdihassan, 1923
Coccophagus ussuriensis Sugonjaev, 1979
Coccophagus varius (Silvestri, 1915)
Coccophagus vegai Girault, 1932
Coccophagus viator Sugonjaev, 1960
Coccophagus vietnamicus Sugonjaev, 1998
Coccophagus yoshidae Nakayama, 1921
Coccophagus youngi (Girault, 1917)
Coccophagus zebratus Howard, 1907
Coccophagus zeyai Hayat, 1998
Coccophagus zinniae De Santis, 1979
Coccophagus formicariae Hayat, 1998

References

Noyes, John S. (March 2019). "Universal Chalcidoidea Database". Natural history museum. Retrieved 1 April 2022.