Eumetabola

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladi: Endopterygota - Paraneoptera - incertae sedis
[list of cladi after Grimaldi & Engel (2005: 147, table 4.1)]
Overview of orders

Coleoptera - Diptera - Hemiptera - Hymenoptera - Lepidoptera - "Mecoptera" - Megaloptera - Neuroptera - Phthiraptera - "Psocoptera" - Raphidioptera - Siphonaptera - Skleroptera - Strepsiptera - Thysanoptera - Trichoptera - †Glosselytrodea - †Miomoptera
[list of ordines after Grimaldi & Engel (2005: 147, table 4.1)]

References

Grimaldi, D.; Engel, M.S. 2005: Evolution of the insects. Cambridge University Press, New York, USA. limited preview on Google books

Eumetabola is an unranked clade of Neoptera. Two large unities known as the Paurometabola and Eumetabola are probably from the adelphotaxa of the Neoptera after exclusion of the Plecoptera. The monophyly of these unities appears to be weakly justified.[1]

Eumetabola has the highest number of species of any clade. According to recent molecular phylogenetic analyses Eumetabola could have first appeared in Middle Devonian (391 million years ago) to the era of late Pennsylvania (311 million years ago). In addition to this, it would have also appeared in Hemiptera (310 million years ago).
Phylogeny

The phylogeny of Eumetabola is shown in the cladogram according to Kluge 2004, 2010, and 2012:[2][3][4]

Neoptera

Idioprothoraca     Rhipineoptera     Eumetabola Parametabola   Zoraptera (angel insects)   Acercaria Condylognatha   Thysanoptera (thrips)     Hemiptera (bugs)     Panpsocoptera   Psocoptera (bark lice)     Phthiraptera (lice)         Endopterygota   Elytrophora   Coleoptera (beetles) 1887).jpg" decoding="async" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f3/Pseudacrossus_przewalskyi_%28Reitter%2C_1887%29.jpg/60px-Pseudacrossus_przewalskyi_%28Reitter%2C_1887%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f3/Pseudacrossus_przewalskyi_%28Reitter%2C_1887%29.jpg/80px-Pseudacrossus_przewalskyi_%28Reitter%2C_1887%29.jpg 2x" data-file-width="3935" data-file-height="2625" height="27" width="40" />     Strepsiptera (twisted-wing parasites)   Coleopteroidea Neuropterida   Neuroptera (net-winged insects)       Raphidioptera (snakeflies)     Megaloptera (alderflies, dobsonflies, fishflies)     Neuropteroidea       Antliophora   Diptera (true flies)       Mecoptera (scorpionflies)       Boreidae (snow scorpionflies)     Siphonaptera (fleas)         Amphiesmenoptera   Trichoptera (caddisflies)     Lepidoptera (butterflies and moths)         Hymenoptera (sawflies, wasps, ants, bees)

References

Paurometabola — Eumetabola link.springer.com
Kluge, Nikita J. (2004). "Larval/pupal leg transformation and a new diagnosis for the taxon Metabola Burmeister, 1832 = Oligoneoptera Martynov, 1923" (PDF). Russian Entomological Journal. 13 (4): 189–229.
Kluge, Nikita J. (2010). "Circumscriptional names of higher taxa in Hexapoda" (PDF). Bionomina. 1: 15–55. doi:10.11646/bionomina.1.1.3.
Kluge, Nikita J. (2012). "General System of Neoptera with Description of a New Species of Embioptera" (PDF). Russian Entomological Journal. 21 (4): 371–384. doi:10.15298/rusentj.21.4.03.