Athetis

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Cladus: Macroheterocera
Superfamilia: Noctuoidea

Familia: Noctuidae
Subfamilia: Noctuinae
Tribus: Caradrinini
Subtribus: Athetiina
Genus: Athetis
Subgenera: Athetis – Hydrillula
Overview of species assigned to a sub-genus

A. brunneolineosa – A. cristifera – A. hanmiensis – A. hexencauma – A. longivalva – A. martoni – A. mienshana – A. motuoensis – A. pseudofunesta –
Overview of species not assigned to a sub-genus

A. aeneolineata – A. alpina – A. anaeodeixis – A. assecta – A. aurogrisea – A. auronitens – A. bicornuta – A. chionopis – A. collicola – A. debilis – A. euxoa – A. fraus – A. fulgens – A. gracilis – A. instrata – A. intricata – A. lyrophora – A. melanephra – A. nigristriata – A. rhigognostis – A. robusta – A. sordida – A. splendidula – A. tristigmatica – A. umbrigera
Name

Athetis Hübner, [1821] 1816

Type species: Noctua dasychira Hübner, [1817] (= Noctua furvula Hübner, 1808)
Synonyms

Hydrilla Boisduval, 1840
Proxenus Herrich-Schäffer, 1845
Hydrilla Guenée, 1852 (preoccupied by Hydrilla Boisduval, 1840)
Elydna Walker, 1858
Radinacra Butler, 1878
Dadica Moore, 1881
Strepselydna Warren, 1911
Hydrillula Tams, 1938 (replacement name )

References

Han, H.-L.; Kononenko, V. 2011: Twelve new species of Athetis Hübner, [1821] 1816 from China (Lepidoptera, Noctuidae). Zootaxa, 3068: 49–68. Preview
Han, H.-L. & C.-D. Li 2008: Descriptions of two new species of the genus Athetis (Lepidoptera: Noctuidae: Xyleninae) from Xizang, China. Acta Zootaxonomica Sinica 33 (4): 696–701. Abstract and full article: [1].
Hreblay, M. & L. Ronkay, 2000: New Noctuidae species and subspecies from Taiwan and the Adjacent Areas II (Lepidoptera). Insects Koreana 17 (1-2): 1–38.
Kononenko, V. 2005: Two new species of Athetis lineosa (Moore, 1888) species group from Taiwan and China (Lepidoptera: Noctuidae: Hadeninae s.l.). Entomologische Zeitschrift 115 (2): 51–54.
Kononenko, V. 2005: Two new species of Athetis (Hübner, 1821) from Taiwan (Lepidoptera: Noctuidae: Hadeninae s.l.). Mitt. Internat. Entomol. 30 (1/2): 15–19.
Kononenko, V. 2005: Three new species of Athetis Hübner, [1821] 1816 (Lepidoptera: Noctuidae: Hadeninae s.l.) from China. Entomologische Zeitschrift 115 (5): 201–204.
Krüger, M. 2000. The genus Athetis Hübner, [1821], in Lesotho (Lepidoptera, Noctuidae: Amphipyrinae). Annals of the Transvaal Museum 37: 139-161. Full article (PDF) Reference page.
Krüger, M. 2001b. A revision of the montane Athetis aurogrisea and A. perplexa species groups (Lepidoptera, Noctuidae: Amphipyrinae). Annals of the Transvaal Museum 38: 19-42. Reference page.
Krüger, M., 2005: New genera and species of noctuid moths from Lesotho (Lepidoptera : Noctuoidea : Noctuidae). African Entomology 13 (1): 97–142.
Lafontaine, J.D. & Schmidt, B.C. 2013. Comments on differences in classification of the superfamily Noctuoidea (Insecta, Lepidoptera) between Eurasia and North America. ZooKeys 264: 209–217. DOI: 10.3897/zookeys.264.4441 Reference page.

Athetis is a genus of moths of the family Noctuidae. The genus was erected by Jacob Hübner in 1821.[1][2][3]
Species

Athetis absorbens (Walker, [1857])
Athetis acutipennis (Dognin, 1919)
Athetis aeschria Hampson, 1909
Athetis aeschrioides Berio, 1940
Athetis alacra (Prout, 1928)
Athetis albanalis (Warren, 1912)
Athetis albilineola Prout, 1928
Athetis albipuncta (Hampson, 1902)
Athetis albirena (Hampson, 1902)
Athetis albisignata (Oberthür, 1879)
Athetis albispinosa (Saalmüller, 1880)
Athetis albosignata (Oberthür, 1879)
Athetis alternata (Janse, 1938)
Athetis andriai Viette, 1963
Athetis anomoeosis Hampson, 1909
Athetis atriluna (Warren, 1913)
Athetis atripuncta (Hampson, 1910)
Athetis atrispherica Hampson, 1914
Athetis atristicta Hampson, 1918
Athetis autobrunnea (Janse, 1938)
Athetis bicolor (Chrétien, 1913)
Athetis bicornis (Hampson, 1891)
Athetis bicornuta Galsworthy, 1997
Athetis bilimbata Berio, 1976
Athetis bimacula (Walker, 1862)
Athetis bipuncta (Snellen, [1886])
Athetis biserrata Han & Kononenko, 2011[4]
Athetis bisignata (Hampson, 1907)
Athetis biumbrosa (Berio, 1940)
Athetis bremusa (Swinhoe, 1885)
Athetis brunneaplagata (Bethune-Baker, 1911)
Athetis brunneolineosa Kononenko, 2005
Athetis bytinskii (Schawerda, 1934)
Athetis calypta Viette, 1958
Athetis camptogramma (Hampson, 1910)
Athetis carayoni Laporte, 1977
Athetis castaneipars (Moore, 1882)
Athetis cervina (Moore, 1881)
Athetis chionephra (Hampson, 1911)
Athetis chionopis Hampson, 1909
Athetis cineracea Warren, 1914
Athetis cinerascens (Motschulsky, 1860)
Athetis cognata (Moore, 1882)
Athetis collaris (Wallengren, 1856)
Athetis condei Viette, 1963
Athetis conformis (Walker, [1857])
Athetis contorta Berio, 1976
Athetis correpta (Püngeler, 1907)
Athetis corticea Hampson, 1918
Athetis costiloba Sugi, 1982
Athetis costiplaga (Bethune-Baker, 1906)
Athetis cristifera Berio, 1977
Athetis cryptisirias Viette, 1958
Athetis dallolmoi (Berio, 1974)
Athetis delecta (Moore, 1881)
Athetis denisi Viette, 1963
Athetis despecta Viette, 1958
Athetis didy Viette, 1963
Athetis discopuncta Hampson, 1916
Athetis dissimilis (Hampson, 1909)
Athetis divisa (Moore, 1882)
Athetis duplex (Janse, 1938)
Athetis elephantula Berio, 1976
Athetis erigida (Swinhoe, 1890)
Athetis eupsilioides Han & Kononenko, 2011[4]
Athetis excurvata (Janse, 1938)
Athetis expolita (Butler, 1876)
Athetis externa (Walker, 1865)
Athetis farinacea (Moore, 1888)
Athetis fasciata (Moore, 1867)
Athetis flacourti Viette, 1963
Athetis flavicolor Han & Kononenko, 2011[4]
Athetis flavipuncta Hampson, 1909
Athetis fontainei (Berio, 1974)
Athetis foveata Hampson, 1909
Athetis fragosa Viette, 1958
Athetis fumicolor (Janse, 1938)
Athetis funesta (Staudinger, 1888)
Athetis furcatula Han & Kononenko, 2011[4]
Athetis furvula (Hübner, [1808])
Athetis gaedei Berio, 1955
Athetis gemini (Bethune-Baker, 1906)
Athetis glauca (Hampson, 1902)
Athetis glaucoides (Berio, 1960)
Athetis glaucopis (Bethune-Baker, 1911)
Athetis gluteosa (Treitschke, 1835)
Athetis gonionephra Hampson, 1909
Athetis grandidieri Viette, 1963
Athetis griveaudi Viette, 1963
Athetis grjebinei Viette, 1963
Athetis heliastis Hampson, 1909
Athetis hennia (Swinhoe, 1901)
Athetis heringi Viette, 1963
Athetis hoengshana Han & Kononenko, 2011[4]
Athetis hongkongensis Galsworthy, 1997
Athetis hospes (Freyer, 1831)
Athetis humberti Viette, 1963
Athetis hyperaeschra Hampson, 1909
Athetis ignava (Guenée, 1852)
Athetis immixta Warren, 1914
Athetis improbalis Berio, 1966
Athetis inconspicua (Bethune-Baker, 1906)
Athetis inquirenda (Strand, 1917)
Athetis interlata (Walker, 1857)
Athetis interstincta (Moore, 1882)
Athetis jeanneli Viette, 1963
Athetis lapidea Wileman, 1911
Athetis lepigone (Möschler, 1860)
Athetis leuconephra Hampson, 1909
Athetis leucopis (Hampson, 1902)
Athetis linealis Yoshimoto, 1994
Athetis lineosa (Moore, 1881)
Athetis lineosella Sugi, 1982
Athetis linzhi Han & Kononenko, 2011[4]
Athetis longiciliata Hampson, 1909
Athetis longidigitus Berio, 1976
Athetis longiharpe Han & Kononenko, 2011[4]
Athetis longivalva Kononenko, 2005
Athetis lyrophora Boursin, 1970
Athetis maculatra (Lower, 1902)
Athetis magniplagia (Hampson, 1918)
Athetis martoni Kononenko, 2005
Athetis melanephra Hampson, 1909
Athetis melanerges Hampson, 1918
Athetis melanomma Hampson, 1920
Athetis melanopis Hampson, 1909
Athetis melanosema Hampson, 1914
Athetis melanosticta Hampson, 1909
Athetis metis (Janse, 1938)
Athetis micra (Hampson, 1902)
Athetis microtera (Hampson, 1902)
Athetis mienshana Kononenko, 2005
Athetis milloti Viette, 1963
Athetis minivalva Han & Kononenko, 2011[4]
Athetis morbidensis Berio, 1976
Athetis mozambica Hampson, 1918
Athetis multilinea Wileman & South, 1920
Athetis mus (Hampson, 1891)
Athetis nephrosticta Hampson, 1909
Athetis nigra (Janse, 1938)
Athetis nigrifons (Dognin, 1919)
Athetis nigrilla Berio, 1976
Athetis nitens (Saalmüller, 1891)
Athetis nonagrica (Walker, [1863])
Athetis obscura (Wileman & South, 1920)
Athetis obscuroides Poole, 1989
Athetis obtusa (Hampson, 1891)
Athetis ocellata (Janse, 1938)
Athetis ochracea (Hampson, 1894)
Athetis ochreipuncta (Hampson, 1894)
Athetis ochreosignata (Aurivillius, 1910)
Athetis oculatissima Berio, 1955
Athetis orthosioides Han & Kononenko, 2011[4]
Athetis pallescens (Janse, 1938)
Athetis pallicornis (Felder & Rogenhofer, 1874)
Athetis pallidipennis Sugi, 1982
Athetis pallustris (Hübner, [1808])
Athetis pectinatissima Berio, 1976
Athetis pectinifer (Aurivillius, 1910)
Athetis pellicea (Swinhoe, 1903)
Athetis percnopis (Bethune-Baker, 1911)
Athetis perineti Viette, 1963
Athetis perparva Berio, 1966
Athetis perplexa (Janse, 1938)
Athetis pigra (Guenée, 1852)
Athetis pilosissima Berio, 1955
Athetis placida (Moore, [1884])
Athetis plumbescens Wileman & West, 1929
Athetis poliophaea Hampson, 1911
Athetis poliostrota Hampson, 1909
Athetis postdentata (Berio, 1960)
Athetis postpuncta Holloway, 1979
Athetis praetexta (Swinhoe, 1905)
Athetis prochaskai Viette, 1963
Athetis pseudofunesta Kononenko, 2005
Athetis pseudolineosa Yoshimoto, 1992
Athetis pulvisculum Berio, 1976
Athetis radama Viette, 1961
Athetis reclusa (Walker, 1862)
Athetis rectilinea (Hampson, 1910)
Athetis renalis (Moore, [1884])
Athetis reniflava (Berio, 1960)
Athetis restricta (Janse, 1938)
Athetis rionegrensis (Chiarelli de Gahn, 1949)
Athetis roastis Hampson, 1918
Athetis robertsi (Janse, 1938)
Athetis rufipuncta (Hampson, 1902)
Athetis rufistigma Warren, 1914
Athetis satellitia (Hampson, 1902)
Athetis scopsis Berio, 1976
Athetis scotopis (Bethune-Baker, 1911)
Athetis seyrigi Viette, 1963
Athetis sicaria Viette, 1958
Athetis siccata Viette, 1958
Athetis signata (Janse, 1938)
Athetis simplex Han & Kononenko, 2011[4]
Athetis sincera (Swinhoe, 1889)
Athetis singula (Möschler, 1883)
Athetis sinistra Berio, 1976
Athetis smintha (Hampson, 1902)
Athetis sobria Berio, 1955
Athetis spaelotidia (Butler, 1879)
Athetis stellata (Moore, 1882)
Athetis stellatula Chang, 1991
Athetis striolata (Butler, 1886)
Athetis stygia Hampson, 1909
Athetis subpartita (Bethune-Baker, 1906)
Athetis suffusa Yoshimoto, 1994
Athetis tarda (Guenée, 1852)
Athetis tenuis (Butler, 1886)
Athetis terminata (Hampson, 1898)
Athetis tetraglypha Berio, 1939
Athetis thoracica (Moore, [1884])
Athetis tornipuncta Berio, 1976
Athetis transvalensis (Janse, 1938)
Athetis transversa (Walker, 1858)
Athetis transversistriata Strand, 1911
Athetis triangulata Wileman & West, 1929
Athetis tridentata Han & Kononenko, 2011[4]
Athetis trixysta Viette, 1958
Athetis unduloma Strand, 1920
Athetis variana (Swinhoe, 1886)
Athetis vernalis Yoshimoto, 1994
Athetis v-parum (Kozhantschikov, 1923)
Athetis xantholopha (Hampson, 1902)
Athetis zelopha Viette, 1963
Athetis zombitsy Viette, 1963

Former species

Athetis miranda is now Proxenus miranda (Grote, 1873)
Athetis mindara is now Proxenus mindara (Barnes & McDunnough, 1913)
Athetis mendosa is now Proxenus mendosa (McDunnough, 1927)

References

Beccaloni, G.; Scoble, M.; Kitching, I.; Simonsen, T.; Robinson, G.; Pitkin, B.; Hine, A.; Lyal, C., eds. (2003). "Athetis". The Global Lepidoptera Names Index. Natural History Museum. Retrieved October 24, 2020.
Savela, Markku (June 11, 2020). "Athetis Hübner, [1821]". Lepidoptera and Some Other Life Forms. Retrieved October 24, 2020.
Pitkin, Brian & Jenkins, Paul (November 5, 2004). "Athetis Hübner, 1821". Butterflies and Moths of the World. Natural History Museum, London. Retrieved October 24, 2020.
Han, H. L. & Kononenko, V. S. (2011). "Twelve new species of Athetis Hubner, [1821] 1816 from China (Lepidoptera, Noctuidae)". Zootaxa. 3068: 49-68.