Ichneumonidae

Ichneumon bucculentus

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Hymenopterida
Ordo: Hymenoptera
Subordo: Apocrita
Superfamilia: Ichneumonoidea

Familia: Ichneumonidae
Subfamiliae (42 + 3†): Acaenitinae – Adelognathinae – Agriotypinae – Anomaloninae – Banchinae – Brachycyrtinae – Brachyscleromatinae – Campopleginae – Collyriinae – Cremastinae – Cryptinae – Ctenopelmatinae – Cylloceriinae – Diacritinae – Diplazontinae – Eucerotinae – Hybrizoninae – Ichneumoninae – Labeninae – Lycorininae – Masoninae – Mesochorinae – Metopiinae – Microleptinae – Neorhacodinae – Nesomesochorinae – Nonninae – Ophioninae – Orthocentrinae – Orthopelmatinae – Oxytorinae – Pedunculinae – Pherombinae – Phrudinae – Pimplinae – Poemeniinae – Rhyssinae – Stilbopinae – Tatogastrinae – Tersilochinae – Townesitinae – Tryphoninae – Xoridinae – †Labenopimplinae – †Palaeoichneumoninae – †Tanychorinae

'Overview of genera (1554 + 6†)
A

Absyrtus – Abzaria – Acaenitellus – Acaenitus – Acanthobenyllus – Acantholabus – Acerastes – Acerataspis – Achaius – Acidnus – Aclastus – Aclosmation – Acolobus – Aconias – Acorystus – Acourtia – Acrodactyla – Acrolyta – Acromia – Acropimpla – Acroricnus – Acrotaphus – Acrotomus – Aculichneumon – Aculicoxa – Adelognathus – Adelotropis – Adelphion – Aderaeon – Aechmeta – Aeglocryptus – Aeliopotes – Aeneonaenaria – Aethecerus – Aethiamblys – Aethianoplis – Aethioplites – Aethioplitops – Afrectopius – Afrephialtes – Afretha – Afrobystra – Afrocoelichneumon – Afrolongichneumon – Afromelanichneumon – Afromevesia – Afrophion – Afrotrogus – Agasthenes – Agathilla – Agathophiona – Aglaodina – Aglaojoppa – Agonocryptus – Agriotypus – Agrothereutes – Agrypon – Ahilya – Aiura – Akymichneumon – Alaothyris – Alcima – Alcochera – Alexeter – Algathia – Allomacrus – Allonotus – Allophatnus – Allophroides – Allophrys – Alloplasta – Allotheca – Alomya – Alophophion – Alophopimpla – Alophosternum – Alystria – Amauromorpha – Amazopimpla – Ambloplisus – Amblyaclastus – Amblyaeneus – Amblyjoppa – Amblysmenus – Amblyteles – Amphibulus – Amphirhachis – Amrapalia – Amydraulax – Anacis – Anastelgis – Anepomias – Aneuclis – Aniseres – Anisobas – Anisopygus – Anisotacrus – Ankylophon – Anomalon – Anoncus – Anupama – Anurotropus – Aoplus – Apaeleticus – Apatetor – Apatetorides – Apechoneura – Apechthis – Aperileptus – Aphanistes – Aplomerus – Apoclima – Apocryptus – Apoglutus – Apophua – Apophysius – Apotemnus – Aprix – Apsilops – Aptesis – Araucacis – Aravenator – Arbelus – Archboldiella – Archoprotus – Arearia – Arenetra – Areyonga – Arhytis – Aridella – Aritranis – Arotes – Arotrephes – Artherola – Arthula – Asmenophylax – Asperpunctatus – Aspidon – Asthenara – Asthenolabus – Astigmaton – Astiphromma – Astomaspis – Astrenis – Atabulus – Atanyjoppa – Ateleute – Atithasus – Atopotrophos – Atractodes – Atractogaster – Atrometoides – Atrometus – Atropha – Auberteterus – Aubertiana – Aucklandella – Augerella – Aulojoppa – Auritus – Australochus – Australoglypta – Austriteles – Austropion – Avnia – Azelus –
B

Baeosemus – Baltazaria – Bambuscopus – Banchogastra – Banchopsis – Banchus – Baranisobas – Barichneumon – Barichneumonites – Barronia – Baryceros – Barycnemis – Barylypa – Barytarbes – Barytarbodes – Barytatocephalus – Barythixis – Basileucus – Batakomacrus – Bathyplectes – Bathythrix – Bathyzonus – Belesica – Benjaminia – Bentyra – Benyllus – Bicryptella – Bicurta – Bilira – Bimentum – Bioblapsis – Blapsidotes – Bodedia – Boethella – Boethus – Boleslawia – Boloderma – Bonthainiella – Bothromus – Bovijoppa – Bozakites – Brachedra – Brachychroa – Brachycyrtus – Brachynervus – Brachypimpla – Brachyscleroma – Brachyzapus – Bremiella – Breviterebra – Brevitubulus – Brussinocryptus – Buathra – Buodias – Bureschias – Buysmania – Bystra –
C

Cacomisthus – Cadarca – Caenocryptoides – Caenocryptus – Caenopelte – Calaminus – Calcaneum – Callajoppa – Calleupalamus – Callidora – Calliephialtes – Calosphyrum – Camera – Campocraspedon – Campoctonus – Campodorus – Campoletis – Campoplex – Camptotypus – Carinityla – Carinodes – Carlsonia – Carria – Casinaria – Caspipina – Castrosion – Catachora – Catadacus – Catadelphops – Catadelphus – Cataptygma – Catastenus – Catucaba – Cechenodes – Cecidopimpla – Celata – Celebarches – Celebichneumon – Celebijoppa – Celor – Centeterichneumon – Centeterus – Cephalobaris – Cephaloglypta – Ceratocryptus – Ceratodolius – Ceratogastra – Ceratojoppa – Ceratomansa – Ceratophenax – Ceratophygadeuon – Certonotus – Cestrus – Chablisea – Chamula – Charitopes – Charmedia – Charmops – Charops – Chasmias – Chasmopygium – Chauvinia – Cheesmanella – Chiaglas – Chilecryptus – Chilelabus – Chilhoplites – Chilocyrtus – Chiloplatys – Chineater – Chirotica – Chlorocryptus – Chorinaeus – Chriodes – Chrionota – Chromocryptus – Chromoplex – Chrysocryptus – Cidaphus – Cisaris – Cladeutes – Clasis – Clatha – Clistopyga – Clitiga – Clydonium – Clypeocampulum – Clypeocava – Clypeodromus – Clypeolabus – Clypeoplex – Clypeoteles – Cnastis – Cnemojoppa – Cobunus – Coccygodes – Coelapatetor – Coelichneumon – Coelichneumonops – Coelojoppa – Coeloleptops – Coelorhachis – Coesula – Colaulus – Coleocentrus – Collyria – Colocnema – Colpognathus – Colpotrochia – Compsocryptus – Compsophorus – Conocalama – Conopyge – Coptomystax – Cordeleboea – Cormobius – Cornuprocerus – Cornutoplisus – Corsoncus – Corymbichneumon – Cosmiocryptus – Cosmiojoppa – Cosmoconus – Costifrons – Cotiheresiarches – Crathiorada – Cratichneumon – Cratocryptus – Cratojoppa – Cratolabus – Creagrura – Cremastus – Cremnocryptus – Cremnodes – Cressonia – †Crusopimpla – Cryptanura – Crypteffigies – Cryptohelcostizus – Cryptojoppa – Cryptophion – Cryptopimpla – Cryptoplites – Cryptopteryx – Cryptus – Crytea – Ctenichneumon – Ctenichneumonops – Cteniscus – Ctenocalops – Ctenocalus – Ctenochares – Ctenochira – Ctenopelma – Cubocephalus – Cubus – Cumatocinetus – Cushmaniella – Cyanodolius – Cyanopelor – Cycasis – Cyclaulus – Cyclolabellus – Cyclolabus – Cylloceria – Cymodusa – Cymodusopsis – Cyrtorhyssa –
D

Dagathia – Daggoo – Dammermaniella – Dammermaniellops – Darachosia – Darymna – Daschia – Debilos – Debophanes – Degithina – Deleboea – Delomerista – Demopheles – Deniya – Denticeria – Dentichasmias – Dentichasmiops – Denticrytea – Dentilabus – Dentimachus – Depressopyga – Dessartodontus – Deuterolabops – Deuterotypus – Deuteroxorides – Diacantharius – Diacritus – Diadegma – Diadromus – Diaglyptella – Diaglyptellodes – Diaglyptidea – Dialipsis – Diaparsis – Diapetimorpha – Diaschisaspis – Diatora – Dicaelodontus – Dicaelognathus – Dicaelotus – Dicamixus – Dicamptus – Dichrogaster – Dictyonotus – Dictyopheltes – Digonocryptus – Dihelus – Dilleria – Dilleritomus – Diloa – Dilopharius – Dimaetha – Dimophora – Dimorphonyx – Dineotropica – Dinocryptus – Diphyus – Diplazon – Diplohimas – Diracela – Diradops – Dirophanes – Dismodix – Distathma – Distictus – Dolichomastix – Dolichomitus – Dolichopselephus – Dolichotrochanter – Dolophron – Dothenia – Dotocryptus – Dreisbachia – Drepanoctonus – Dusona – Dyspetes –
E

Earobia – Ecaepomia – Eccoptosage – Echthromorpha – Echthronomas – Echthrus – Eclytus – Ecphysis – Ectomocolax – Ectopimorpha – Ectopius – Ectopoides – Eiphosoma – Eknomia – Eleebichneumon – Elysioreiga – Encardia – Enclisis – Encrateola – Endasys – Endromopoda – Enicospilus – Enizemum – Entypoma – Enytus – Eparces – Epelaspis – Ephialtes – Epirhyssa – Episemura – Epistathmus – Epitheronia – Epitomus – Eremocinetus – Eremodolius – Eremotylus – Eremura – Eriborus – Eridolius – Erigorgus – Eriostethus – Eriplatys – Eristicus – Erromenus – Eruga – Erythrodolius – Etha – Ethelurgus – Eubaeus – Eucaphila – Euceros – Euchalinus – Eucremastoides – Eucremastus – Eudelus – Eugalta – Euheresiarches – Eupalamus – Eurycryptus – Eurydacus – Eurygenys – Eurylabus – Euryophion – Euryproctus – Eurypterna – Eurytyloides – Eusterinx – Eutanyacra – Eutanygaster – Evirchomella – Excavarus – Exenterus – Exephanes – Exeristes – Exestuberis – Exetastes – Exochus – Extenuosodalis – Exyston –
F

Facydes – Fenixia – Ferrocryptus – Fianoniella – Fileanta – Fitatsia – Flacopimpla – Flavopimpla – Formocryptus – Formostenus – Forrestopius – Fortipalpa – Fotsiforia – Foveosculum – Fractipons – Fredegunda – Friona –
G

Gabia – Gabunia – Gahus – Gambroides – Gambrus – Ganodes – Gareila – Gathetus – Gauldianus – Gavrana – Gelanes – Gelis – Genaemirum – Geniceris – Genotropis – Geraldus – Gerdius – Gessia – Ghilaromma – Ghilarovites – Gibbobystra – Gibbosoplites – Giraudia – Glabridorsum – Glodianus – Glyphaclastus – Glyphicnemis – Glypta – Glyptopimpla – Glyptorhaestus – Gnamptopelta – Gnathochorisis – Gnotus – Gnypetomorpha – Goedartia – Golbachiella – Gonotypus – Goryphus – Gotra – Grasseiteles – Gravenhorstia – Gregopimpla – Groenlabus – Grotea – Grypocentrus – Gunomeria – Guptella – Gyrodonta – Gyrodontichneumon – Gyrolaba – Gyropyga –
H

Habrocampulum – Habronyx – Hackerocryptus – Hadeleboea – Hadrocryptus – Hadrodactylus – Hadrostethus – Hallocinetus – Hanauella – Handaoia – Hapsinotus – Harpura – Hedycryptus – Hedyjoppa – Hedylus – Heinrichia – Helcostizus – Helenanomalon – Helictes – Hellwigia – Hellwigiella – Helotorus – Hemibystra – Hemibystrops – Hemichneumon – Hemigaster – Hemihoplis – Hemimetopius – Hemiphaisura – Hemiphanes – Hemisphragia – Hemiteles – Hepialichneumon – Hepiopelmus – Hercana – Hercus – Heresiarches – Herpestomus – Heterischnus – Heterocola – Heteropelma – Hiatensor – Hidryta – Hieroceryx – Himerta – Himertosoma – Hintelmannia – Hiorada – Hirtolabus – Hodostates – Hoeocryptus – Holcichneumon – Holcojoppa – Holojoppa – Homaspis – Homotherus – Hoplismenus – Hoplocryptus – Hoplophaeogenes – Hoploplatystylus – Horstmannolochus – Humbert – Hybophorellus – Hybrizon – Hylesicida – Hylophasma – Hymenoepimecis – Hymenura – Hypamblys – Hyparcha – Hyperacmus – Hyperallus – Hyperbatus – Hypomecus – Hypopheltes – Hyposoter – Hypsanacis – Hypsantyx – Hypsicera – Hytophatnus –
I

Iaria – Ibornia – Icariomimus – Ichneumon – Idiogramma – Idiolispa – Ileanta – Ileantella – Imeria – Inbioia – Indagrypon – Indovia – Insulcus – Irabatha – Isadelphus – Isandria – Ischnoceros – Ischnojoppa – Ischnus – Ischyrocnemis – Isdromas – Iseropus – Ishigakia – Isotima – Itamuton – Ithaechma – Itoplectis –
J

Jacotitypus – Janzophion – Javra – Jethsura – Jezarotes – Jomine – Jonathania – Joppa – Joppidium – Joppocryptus – Jorgeus – Junctivena –
K

Kasparyania – Kerrichia – Khalaimia – Klutiana – Kokujewiella – Kriegeria – Kristotomus –
L

Labena – Labium – Labrossyta – Lachmetha – Laderrica – Lagarosoma – Lagarotis – Lagavula – Lagoleptus – Lamachus – Lamnatibia – Lamprocryptidea – Lamprocryptus – Lanugo – Lapton – Lareiga – Larischia – Larpelites – Lathiponus – Lathrolestes – Lathroplex – Lathrostizus – Latibulus – Laticoleus – Latilumbus – Latosculum – Lavinifia – Ledora – Legnatia – Leipula – Lemophagus – Lentocerus – Lepidura – Lepiscelus – Leptacoenites – Leptarthron – Leptixys – Leptobatopsis – Leptocampoplex – Leptocryptoides – Leptojoppa – Leptomalaisia – Leptoperilissus – Leptophatnus – Leptophion – Leptopimpla – Leptops – Leptotogea – Lethades – Leurus – Levansa – Levibasis – Leviculus – Liaodontus – Liaoichneumon – Lichmeres – Lienella – Limerodes – Limerodops – Limonethe – Linycus – Liojoppa – Liojoppites – Liopterna – Liotryphon – Lipoprion – Lissaspis – Lissocaulus – Lissolaboides – Lissolongichneumon – Lissonota – Lissonotidea – Lissophadnus – Lissopimpla – Lissosculpta – Listrocryptus – Listrodromus – Listrognathus – Lithoserix – Lithotorus – Litochila – Livipurpurata – Lobaegis – Lochetica – Longichneumon – Longitibia – Lophoglutus – Lophojoppa – Lophyroplectus – Lorio – Losgna – Loxodocus – Loxopus – Lusius – Lusiuxta – Luteocoelius – Lycorina – Lygurella – Lygurus – Lymantrichneumon – Lymeon – Lynteria – Lysibia – Lytarmes –
M

Macrojoppa – Macromalon – Macrulus – Macrus – Madagasgavrana – Madagasichneumon – Madastenus – Magwengiella – Malaisichneumon – Malaycromia – Mallochia – Mamelia – Mansa – Maraces – Marjorietta – Marlisia – Masona – Mastrulus – Mastrus – Matara – Matinangarches – Maxodontus – Mecistum – Mecotes – Medophron – Megacara – Megaceria – Megaetaira – Megajoppa – Megalomya – Megaplectes – Megarhyssa – Megastylus – Megatryphon – Meggoleus – Melalophacharops – Melanichneumon – Melanocryptus – Melanodolius – Melanoplex – Melcha – Meloboris – Menaforia – Menaka – Meniscomorpha – Menkokia – Meringops – Meringopus – Merolides – Mesochorella – Mesochorus – Mesoclistus – Mesoleius – Mesoleptidea – Mesoleptus – Mesophadnus – Mesophragis – Mesostenus – Messatoporus – Metachorischizus – Metallichneumon – Metoa – Metopheltes – Metopius – Mevesia – Micrandria – Micraris – Microcharops – Microleptes – Microlongichneumon – Micromonodon – Microsage – Microstenus – Miolyta – Misetus – Mnioes – Mokajoppa – Monganella – Monoblastus – Monodontichneumon – Monontos – Monothela – Myllenyxis – Myocious – Myrmecacis – Myrmeleonostenus –
N

Naenaria – Naenarides – Naenarosculum – Nanium – Narthecura – Neamblyaeneus – Neamblymorpha – Neaprix – Nebostenus – Necolio – Neeurylabia – Neischnus – Neleges – Neleothymus – Neliopisthus – Nelophia – Nematocryptus – Nematomicrus – Nematopodius – Nemeritis – Neobuodias – Neocratojoppa – Neocryptopteryx – Neodiphyus – Neodontocryptus – Neoexetastes – Neofacydes – Neohabronyx – Neoheresiarches – Neolareiga – Neolevansa – Neolinycus – Neolophron – Neopimpla – Neoproclitus – Neorhacodes – Neostroblia – Neotheronia – Neotypus – Neoxorides – Nepiesta – Nesolinoceras – Nesostenodontus – Netanyacra – Netelia – Neurateles – Neurogenia – Nippocryptus – Nipponaetes – Nomosphecia – Nonnus – Nonpropodeum – Notacma – Notaulites – Nothischnus – Nothocremastus – Notocampsis – Notophasma – Notophrudus – Notoplatylabus – Notopygus – Notosemus – Notostilbops – Noxocremastus –
O

Obisiphaga – Obtusodonta – Occapes – Occia – Odinophora – Odontocolon – Odontocryptus – Odontojoppa – Odontoneura – Odontopimpla – Oecetiplex – Oecotelma – Oedemopsis – Oedicephalus – Oetophorus – Oezdemirus – Ogulnia – Oiorhinus – Ojuelos – Olesicampe – Olethrodotis – Onarion – Opheltes – Ophiogastrella – Ophion – Ophionellus – Ophiopterus – Oreohoplis – Oresbius – Orgichneumon – Orientohemiteles – Orientospilus – Oriphatnus – Oronotus – Orotylus – Ortezia – Orthizema – Orthocentrus – Ortholaba – Orthomiscus – Orthopelma – Osprynchotus – Otlophorus – Otoblastus – Owenus – Oxyrrhexis – Oxytaenia – Oxytorus – Ozlabium –
P

Pachyjoppa – Pachymelos – Pachysomoides – Pagarenes – Palaeometopius – Palmerella – Palpator – Palpostilpnus – Pamophion – Panteles – Pantisarthrus – Pantorhaestes – Parablastus – Paracoelichneumon – Paracollyria – Paraditremops – Paraethecerus – Paragambrus – Paraglyptus – Paranacis – Parania – Paraperithous – Paraphylax – Parapimpla – Paratanychora – Parema – Parmortha – Parvaoplus – Parvipimpla – Patrocloides – Patroclus – Paxylommites – Peakelestes – Pectinorex – Pedinopa – Pedinopelte – Pedunculus – Pegaoplex – Pentelophus – Pepsijoppa – Pergaphaga – Perilimicron – Perilissus – Perinetia – Periope – Periplasma – Perisphincter – Perispuda – Peritasis – Perithous – Perjiva – Petilium – Petilochus – Peucobius – Peumocryptus – Phaeneumon – Phaenolabrorychus – Phaenolobus – Phaeogenes – Phaestacoenitus – Phaestus – Phairichneumon – Phaisura – Phaisurella – Phaisurellops – Phalgea – Pharetrophora – Pharzites – Pherhombus – Philogalleria – Philositus – Phobetes – Phobocampe – Phorotrophus – Photocryptus – Phradis – Phrudus – Phthorima – Phycitiplex – Phygadeuon – Physotarsus – Phytodietus – Piambia – Piasites – Picacharops – Picardiella – Picrocryptoides – Picrostigeus – Pimpla – Pimplaetus – Pimplomorpha – Piogaster – Pion – Pionpherta – Pithotomus – Plagiotrypes – Planochorus – Platybirmania – Platyjoppa – Platylabops – Platylabus – Platymischos – Platymystax – Platyrhabdus – Plectiscidea – Plectiscus – Plectochorus – Plectocryptus – Pleolophus – Pleurogyrus – Podeleboea – Podogaster – Podoschistus – Poecilocryptus – Poecilodromops – Poecilodromus – Poecilojoppa – Poecilojoppoides – Poecilostictus – Poemenia – Polyaulon – Polyblastus – Polyconus – Polycyrtidea – Polycyrtus – Polyphrix – Polysphincta – Polytribax – Porizon – Porizonopteron – Prethophion – Prionacis – Priopoda – Priotomis – Pristicerops – Pristiceros – Pristomerus – Probles – Probolus – Procerochasmias – Procestus – Prochas – Procinetus – Proclitus – Proeliator – Projoppa – Promethes – Prosacron – Prosthoporus – Protarchus – Protichneumon – Protoleptops – Protopelmus – Provancherides – Pseudagrypon – Pseudalomya – Pseudanomalon – Pseuderipternus – Pseudeupalamus – Pseudevirchoma – Pseudischnojoppa – Pseudoamblyteles – Pseudocillimops – Pseudocillimus – Pseudocoelichneumon – Pseudocremastus – Pseudomaraces – Pseudometopius – Pseudopimpla – Pseudoplatylabops – Pseudoplatylabus – Pseudorhyssa – Pseudotogea – Pseudotricapus – Psilomastax – Pterinopus – Pterocryptus – Ptilobaptus – Ptychopsis – Punctileptops – Pycnocryptodes – Pycnoflatsor – Pycnophion – Pygmaeolus – Pygocryptus – Pyracmon – Pyramidamblys – Pyramidophorus –
Q

Quandrus – Queequeg – Quillonota –
R

Rambites – Raninia – Rasnitsynites – Reclinervellus – Regius – Reptatrix – Retalia – Rhabdosis – Rhachioplex – Rhadinodonta – Rhadinodontoplisus – Rhadinodontops – Rhadinomastrus – Rhaestus – Rhembobius – Rhimphoctona – Rhinium – Rhinotorus – Rhopalophion – Rhorodes – Rhorus – Rhynchocryptus – Rhynchophion – Rhysaspis – Rhyssa – Rhyssella – Rhytura – Ribasia – Ricrena – Rictichneumon – Riekophion – Rimbusia – Rodrigama – Romaniella – Rossemia – Rothneyia – Rubicundiella – Rugascuta – Rugosculpta – Rynchobanchus –
S

Sachtlebenia – Saltagenes – Saotis – Saranaca – Sathropterus – Satous – Satrius – Savolia – Scambus – Scapnetes – Scenocharops – Schachticraspedon – Schelocentrus – Schenkia – Schizopyga – Schreineria – Sciocryptus – Sciron – Scirtetes – Sclerophion – Scolobates – Scolomus – Scopesis – Scrobiculus – Scudderopsis – Scutellator – Seleucus – Semimesoleius – Semitogea – Sericopimpla – Serratosculum – Sesioplex – Setanta – Setantops – Seticornuta – Seyrighoplites – Seyrigichneumon – Seyrigiella – Shortia – Sialocara – Sicophion – Silsila – Simophion – Sinarachna – Singalissaspis – Sinophorus – Siphimedia – Sjostedtiella – Skeatia – Skiapus – Sliochia – Smicrolius – Smicroplectrus – Solitosculum – Sozites – Spanophatnus – Spathacantha – Sphaeromanus – Sphecoctonus – Sphecophaga – Sphelodon – Sphinctus – Sphingozona – Spilichneumon – Spilopimpla – Spilopteron – Spilothyrateles – Spinallonotus – Spinamblys – Spinellamblys – Spinolochus – Spolas – Spudaeus – Stangepelma – Stauropoctonus – Stenaoplus – Stenapatetor – Stenarches – Stenarchops – Stenarella – Stenichneumon – Stenichneumonopsis – Stenobarichneumon – Stenobenyllus – Stenodontus – Stenogynaia – Stenolonche – Stenomacrus – Stenotes – Stethantyx – Stetholophus – Stethoncus – Stibeutes – Stictolissonota – Stictopisthus – Stilbops – Stilpnus – Stipomoles – Stirexephanes – Stirojoppa – Stiromesostenus – Stomacis – Strabotes – Strongylopsis – Sulcarius – Surculus – Sussaba – Sustenus – Sycaonia – Sympherta – Symplecis – Syndipnus – Synechocryptus – Synetaeris – Synodites – Synoecetes – Synomelix – Synosis – Syntactus – Syntrips – Syrphidepulo – Syrphoctonus – Syrphophilus – Syspasis – Syzeuctus –
T

Takastenus – Tamaulipeca – Tanepomidos – Tanychela – Tanychora – Tanychorella – Tanyloncha – Tashtego – Tasmabates – Tatogaster – Teleutaea – Teluncus – Temelucha – Terebraella – Terminator – Tersilochus – Tersoakus – Tetractenion – Tetragonochora – Tetrambon – Thamester – Thascia – Thaumatocephalus – Thaumatogelis – Thaumatoplites – Thaumatoplitops – Thelodon – Therion – Theronia – Theroscopus – Thibetoides – Thrybius – Thymariodes – Thymaris – Thymebatis – Thyrateles – Thyreodon – Thyridoplites – Ticapimpla – Tilgidopsis – Tmetogaster – Tobiasites – Toechorychus – Togea – Togeella – Tolonus – Tomagotra – Torbda – Torquaoplus – Torquinsha – Tossinola – Tossinolodes – Townesia – Townesion – Townesites – Townostilpnus – Trachaner – Trachyarus – Trachyglutus – Trachysphyrus – Trafana – Tranosema – Tranosemella – Trathala – Trematopygodes – Trematopygus – Tretobasis – Triancyra – Tricapus – Tricentrum – Tricholabus – Tricholinum – Trichomma – Triclistus – Tricyphus – Trieces – Trihapsis – Triptognathops – Triptognathus – Trogichneumon – Trogomorpha – Trogopyga – Trogus – Tromatobia – Trophophion – Tropicolabus – Tropistes – Trychosis – Tryonocryptus – Trypha – Tryphon – Tsirambia – Tumeclypeus – Tycherus – Tymmophorus –
U

Uchidella – Ulesta – Uloola – Umanella – Urancyla – Urotryphon – Urvashia –
V

Vagenatha – Validentia – Varnado – Venturia – Vernamalon – Vestibulum – Virgichneumon – Vulgichneumon –
W

Wahlamia – Wahliodontus – Westwoodia – Whymperia – Woldstedtius – Wuda –
X

Xanthephialtes – Xanthocampoplex – Xanthocryptus – Xanthophenax – Xanthopimpla – Xanthosomnium – Xenarthron – Xenolytus – Xenoschesis – Xenothyris – Xestojoppa – Xestopelta – Xiomara – Xiphonychidion – Xiphosomella – Xiphulcus – Xorides – Xoridesopus – Xylacis – Xylophion – Xylophrurus – Xylostenus –
Y

Yamatarotes – Yeppoona – Yezoceryx –
Z

Zabrachypus – Zaglyptomorpha – Zaglyptus – Zagryphus – Zambion – Zanthojoppa – Zaplethocornia – Zatypota – Zealochus – Zemiophora – Zonocryptus – Zonopimpla – Zoophthorus – Zophoplites – †Agapia – †Agapteron – †Eubaeus – †Rovenosa – †Urotryphon

[source: Catalogue of Life: 2013 Annual Checklist, except fossils, plus (8) Bicurta – Carinityla – Eknomia – Helenanomalon – Khalaimia – Kibalus – Ojuelos – †Rovenosa
]

Check (1): Longitergus
Name

Ichneumonidae Latreille, 1802

References

Latreille, P.A. 1802. Histoire naturelle, générale et particulière des crustacés et des insectes. Ouvrage faisant suite à l’histoire naturelle générale et particulière, composée par Leclerc de Buffon, et rédigée par C.S. Sonnini, membre de plusieurs sociétés savantes. Familles naturelles des genres. Tome troisième. F. Dufart, Paris, xii + pp. 13–467 + [1 (errata)]. BHL Reference page.
Aquino, D.A.; Gaddi, A.L.; Hernández, E.P.; Martínez, J.J. 2010: The types of Braconidae and Ichneumonidae (Hymenoptera: Ichneumonoidea) in the Museo de La Plata, Argentina. Zootaxa, 2487: 43–51. Preview
Bennett, A.M.R. 2009: The Ichneumonidae (Hymenoptera) of Fiji: keys to subfamilies and genera with a review of the species of Anomaloninae, Banchinae, Brachycyrtinae and Diplazontinae. In: Evenhuis, N.L. & Bickel, D.J. (eds.), Fiji Arthropods XIV. Bishop Museum occasional papers, 105: 3–68. PDF
Broad, G.R. 2016. Checklist of British and Irish Hymenoptera - Ichneumonidae. Biodiversity Data Journal 4: e9042. DOI: 10.3897/BDJ.4.e9042. Reference page.
Fitton, M.G.; Gauld, I.D. 1976: The family-group names of the Ichneumonidae (excluding the Ichneumoninae). Systematic entomology, 1: 247–258.
Fitton, M.G.; Gauld, I.D. 1978: Further notes on family-group names of Ichneumonidae (Hymenoptera). Systematic entomology, 3: 245–247.
Gauld, I.D. 1984: An introduction to the Ichneumonidae of Australia. British Museum (Natural History).
Gauld, I.D.†; Fernández-Triana, J.L. 2010: Type condition and generic placement of Cuban species of Ichneumonidae described by Cresson and collected by Gundlach. Zootaxa, 2394: 41–50. Preview
González-Moreno, A.; Bordera, S. 2011: New records of Ichneumonidae (Hymenoptera: Ichneumonoidea) from Mexico. Zootaxa, 2879: 1–21. Preview
González-Moreno, A.; Bordera, S. 2012: The Ichneumonidae (Hymenoptera: Ichneumonoidea) of Ría Lagartos Biosphere Reserve, Yucatán, Mexico. Zootaxa, 3230: 1–51. Preview
Kopylov, D.S. 2010: A new subfamily of ichneumon wasps (Insecta: Hymenoptera: Ichneumonidae) from the Upper Cretaceous of the Russian Far East. Paleontological journal, 44(4): 422-??? DOI: 10.1134/S003103011004009X
Kopylov, D.S., Spasojevic, T. & Klopfstein, S. 2018. New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East. Zootaxa 4442(2): 319–330. DOI: 10.11646/zootaxa.4442.2.8 Paywall Reference page.
Perkins, J.F., 1962: On the type species of Foerster's genera (Hymenoptera: Ichneumonidae). Bulletin of the British Museum (Natural History) Entomology series 11 (8): 383–483.
Quicke, D.L.J.; Fitton, M.G.; Notton, D.G.; Broad, G.R.; Dolphin, K.; Austin, A.D. 2000: Phylogeny of the subfamilies of Ichneumonidae (Hymenoptera): a simultaneous molecular and morphological analysis. pp. 74–83 in Austin, A. & Dowton, M. (eds) Hymenoptera: Evolution, Biodiversity and Biological Control. CSIRO Publishing, Collingwood.
Quicke, D.L.J.; Laurenne, N.M.; Fitton, M.G.; Broad, G.R. 2009: A thousand and one wasps: a 28S rDNA and morphological phylogeny of the Ichneumonidae (Insecta: Hymenoptera) with an investigation into alignment parameter space and elision. Journal of natural history, 43: 1305–1421. DOI: 10.1080/00222930902807783
Quicke, D.L.J.; Lopez-Vaamonde, C.; Belshaw, R. 1999: The basal Ichneumonidae (Insecta, Hymenoptera): 28S D2 rDNA considerations of the Brachycyrtinae, Labeninae, Paxylommatinae and Xoridinae. Zoologica scripta, 28: 203–210. DOI: 10.1046/j.1463-6409.1999.00010.x
Rodríguez-Berrío, A.; Bordera, S.; Sääksjärvi, I.E. 2009: Checklist of Peruvian Ichneumonidae (Insecta, Hymenoptera). Zootaxa, 2303: 1–44. Abstract & excerpt
Sullivan, G.T.; Karaca, I.; Ozman-Sullivan, S.K.; Kolarov, J. 2010: Ichneumonid (Hymenoptera) parasitoids of overwintering Hyphantria cunea (Drury) (Lepidoptera: Arctiidae) pupae in hazelnut plantations of the central Black Sea region of Turkey. Zootaxa, 2608: 63–68. Preview
Tereshkin, A. 2009: Illustrated key to the tribes of subfamilia Ichneumoninae and genera of the tribe Platylabini of world fauna (Hymenoptera, Ichneumonidae). Linzer biologische Beiträge, 41(2): 1317–1608. PDF
Wahl, D.B.; Gauld, I.D. 1998: The cladistics and higher classification of the Pimpliformes (Hymenoptera: Ichneumonidae). Systematic entomology, 23: 265–298. DOI: 10.1046/j.1365-3113.1998.00057.x

Additional references

Bennett, A.M.R., Cardinal, S., Gauld, I.D. & Wahl, D.B. 2019. Phylogeny of the subfamilies of Ichneumonidae (Hymenoptera). Journal of Hymenoptera Research 71: 1–156. DOI: 10.3897/jhr.71.32375 Reference page.
di Giovanni, F., Reshchikov, A., Riedel, M., Diller, E. & Schwarz, M. 2015. New records of Ichneumonidae (Hymenoptera) for the Italian fauna. Biodiversity Data Journal 3: e5057. DOI: 10.3897/BDJ.3.e5057. Reference page.
Klopfstein, S., Riedel, M. & Schwarz, M. 2019. Checklist of ichneumonid parasitoid wasps in Switzerland (Hymenoptera, Ichneumonidae): 470 species new for the country and an appraisal of the alpine diversity. Alpine Entomologiy, 3: 51–81. DOI: 10.3897/alpento.3.31613 Reference page.

Vernacular names
suomi: Ahmaspistiäiset
日本語: ヒメバチ科

The Ichneumonidae, also known as the ichneumon wasps, Darwin wasps, or ichneumonids, are a family of parasitoid wasps of the insect order Hymenoptera. They are one of the most diverse groups within the Hymenoptera with roughly 25,000 species currently described.[2] However, this likely represents less than a quarter of their true richness as reliable estimates are lacking, along with much of the most basic knowledge about their ecology, distribution, and evolution.[1] Ichneumonid wasps, with very few exceptions, attack the immature stages of holometabolous insects and spiders, eventually killing their hosts.[3] They thus fulfill an important role as regulators of insect populations, both in natural and semi-natural systems, making them promising agents for biological control.[4]
Yellow and black flying insect lacking an ovipositor... male Ichneumonid Wasp
Male Ichneumonid Wasp

The distribution of the ichneumonids was traditionally considered an exception to the common latitudinal gradient in species diversity, since the family was thought to be at its most species-rich in the temperate zone instead of the tropics, but numerous new tropical species have now been discovered.

Etymology and history

Insects in the family Ichneumonidae are commonly called ichneumon wasps, or ichneumonids. However, the term ichneumon wasps can refer specifically to the genus Ichneumon within the Ichneumonidae and thus can cause confusion. A group of ichneumonid specialists have proposed Darwin wasps as a better vernacular name for the family.[4] Less exact terms are ichneumon flies (they are not closely related to true flies) and scorpion wasps due to the extreme lengthening and curving of the abdomen (scorpions are arachnids, not insects).

The name is derived from Latin 'ichneumon', from Ancient Greek ἰχνεύμων (ikhneúmōn, "tracker"), from ἴχνος (íkhnos, "track, footstep"). The name first appeared in Aristotle's "History of Animals", c. 343 BC. Aristotle noted that the ichneumon preys upon spiders, is a wasp smaller than ordinary wasps, and carries its prey to a hole which they lay their larvae inside, and that they seal the hole with mud.[5] Aristotle's writing, however, more accurately describes the mud daubers than the true ichneumon wasps, which do not construct mud nests and do not sting.
Description

Adult ichneumonids superficially resemble other wasps. They have a slender waist, two pairs of wings, a pair of large compound eyes on the side of the head and three ocelli on top of the head. Their size varies considerably from a few millimetres to seven or more centimetres.

The ichneumonids have more antennal segments than typical, aculeate wasps (Aculeata: Vespoidea and Apoidea): ichneumonids typically possess 16 or more, while most other wasps have 13 or fewer. Unlike aculeate wasps, which have an ovipositor modified for prey capture and defense, and do not pass their eggs along the stinger, ichneumonid females have an unmodified ovipositor which they use to lay eggs inside or on their host. Ichneumonids generally inject venom along with the egg, but only larger species (some in the genera Netelia and Ophion) with relatively short ovipositors use the ovipositor in defense. Males do not possess stingers or ovipositors in either lineage.

Ichneumonids are distinguished from their sister group Braconidae mainly on the basis of wing venation. The fore wing of 95% of ichneumonids has vein 2m-cu (in the Comstock–Needham system), which is absent in braconids. Vein 1rs-m of the fore wing is absent in all ichneumonids, but is present in 85% of braconids. In the hind wing of ichneumonids, vein rs-m joins Rs apical to (or rarely opposite) the split between veins Rs and R1. In braconids, vein rs-m joins basal to this split. The taxa also differ in the structure of the metasoma: about 90% of ichneumonids have a flexible suture between tergites 2 and 3, whereas these tergites are fused in braconids (though the suture is secondarily flexible in Aphidiinae).[6]
Ichneumonid fore wing (Syzecteus sp.). The presence of vein 2m-cu and absence of vein 1Rs+M distinguish the wing from that of braconids.

Ichneumonid hind wing. Vein rs-m joins Rs after the split between veins Rs and R1.

Braconid hind wing. Vein rs-m joins Rs before the split between veins Rs and R1.
Distribution

Ichneumonids are found on all continents with the exception of Antarctica. They inhabit virtually all terrestrial habitats, wherever there are suitable invertebrate hosts.

The distribution of ichneumonid species richness is subject to ongoing debate. Long believed to be rare in the tropics, and at its most species rich in the temperate region, the family became a classic textbook example of an 'exceptional' latitudinal diversity gradient.[7] Recently this belief has been questioned, after the discovery of numerous new tropical species.[8][9][10]
Reproduction and diet

A very few ichneumonid species lay their eggs in the ground, but the vast majority inject eggs either directly into their host's body or onto its surface, and this may require penetration of substrate around the host, as in wood-boring host larvae that live deep inside of tree trunks, requiring the ichneumon to drill its ovipositor through several centimeters of solid wood (e.g., Megarhyssa species). After hatching, the ichneumonid larva consumes its still living host. The most common hosts are larvae or pupae of Lepidoptera, Coleoptera and Hymenoptera. Some species in the subfamily Pimplinae also parasitise spiders. Hyperparasitoids such as Mesochorinae oviposit inside the larvae of other ichneumonoids. The hosts of some species are agricultural pests, therefore ichneumons are sometimes valuable for biological pest control, but the hosts of most species are unknown; host information has been reviewed and summarized by various researchers, e.g. Aubert,[11][12][13] Perkins.[14][15] and Townes.[16]

Ichneumonids use both idiobiont and koinobiont strategies. Idiobionts paralyze their host and prevent it from moving or growing. Koinobionts allow their host to continue to grow and develop. In both strategies, the host typically dies after some weeks, after which the ichneumonid larva emerges and pupates.[17]
See also: Parasitoid § Basic concepts

Adult ichneumonids feed on a diversity of foods, including plant sap and nectar. They spend much of their active time searching, either for hosts (female ichneumonids) or for emerging females (male ichneumonids).[18] The parasitism pressure exerted by ichneumonids can be tremendous, and they are often one of the major regulators of invertebrate populations.[19][20] It is quite common for 10-20% or more of a host's population to be parasitised (though reported parasitism rates often include non-ichneumonid parasitoids).[21][22]

Taxonomy and systematics
An ichneumonid caught in amber 15-20 million years ago.

The taxonomy of the ichneumonids is still poorly known. The family is highly diverse, containing 24,000 described species. Approximately 60,000 species are estimated to exist worldwide, though some estimates place this number at over 100,000. They are severely undersampled, and studies of their diversity typically produce very high numbers of species which are represented by only a single individual.[23][24] Due to the high diversity, the existence of numerous small and hard to identify species, and the majority of species being undiscovered, it has proven difficult to resolve the phylogeny of the ichneumonids. Even the relationships between subfamilies are unclear. The sheer diversity also means DNA sequence data is only available for a tiny fraction of the species, and detailed cladistic studies require major computing capacity.

Extensive catalogues of the ichneumonids include those by Aubert,[11][12][13] Gauld,[25] Perkins,[14][15] and Townes.[16][26][27][28][29] Due to the taxonomic difficulties involved, however, their classifications and terminology are often confusingly contradictory. Several prominent authors have gone as far as to publish major reviews that defy the International Code of Zoological Nomenclature.[26][27][28][29][30][31]

The large number of species in Ichneumonidae may be due to the evolution of parasitoidism in hymenoptera, which occurred approximately 247 million years ago.[32][33] Ichneumonidae is the basal branch of Apocrita, the lineage in which parasitoidism in hymenoptera evolved, and some ichneumonids are thought to have been in stasis for millions of years and closely resemble the common ancestor in which parasitoidism evolved. This common ancestor was likely an Ectoparasitoid woodwasp that parasitized wood-boring beetle larvae in trees.[34] The family has existed since at least the Early Cretaceous (c. 125 mya),[35][36] but probably appeared already in the Jurassic (c. 181 mya), soon after the appearance of its major host groups.[37] It diversified during the Oligocene.
Subfamilies

In 1999, the extant ichneumonids were divided into 39 subfamilies,[38][39] whose names and definitions have varied considerably. The phylogenetic relationships between the subfamilies are still unclear.

Acaenitinae
Adelognathinae
Agriotypinae
Anomaloninae (= Anomalinae)
Banchinae
Brachycyrtinae (sometimes included in Labeninae)
Campopleginae (= Porizontinae)
Collyriinae
Cremastinae
Cryptinae (= Gelinae, Hemitelinae, Phygadeuontinae)
Xanthocryptus novozealandicus
Ctenopelmatinae (= Scolobatinae)
Cylloceriinae (sometimes included in Microleptinae)
Diacritinae (sometimes included in Pimplinae)
Diplazontinae
Eucerotinae (sometimes included in Tryphoninae)
Hybrizontinae (= Paxylommatinae) (sometimes placed in own family)
Ichneumoninae
Labeninae (= Labiinae)
Labenopimplinae (extinct)
Lycorininae (sometimes included in Banchinae)
Mesochorinae
Metopiinae
Microleptinae
Neorhacodinae (sometimes included in Banchinae)
Nesomesochorinae
Ophioninae
Orthocentrinae (sometimes included in Microleptinae)
Orthopelmatinae
Oxytorinae
Pedunculinae
Phrudinae
Pimplinae (= Ephialtinae)
Poemeniinae (sometimes included in Pimplinae)
Rhyssinae (sometimes included in Pimplinae)
Sisyrostolinae (sometimes included in Phrudinae)
Stilbopinae (sometimes included in Banchinae)
Tatogastrinae (sometimes included in Microleptinae)
Tersilochinae
Xoridinae

Famous ichneumonologists

Famous ichneumonologists include:

Jacques Aubert
Carl Gustav Alexander Brischke
Peter Cameron
Arnold Förster
Johann Ludwig Christian Gravenhorst
Alexander Henry Haliday
Gerd Heinrich
August Emil Holmgren
Joseph Kriechbaumer
Thomas Ansell Marshall
Henry Keith Townes
Constantin Wesmael

Darwin and the Ichneumonidae

The perceived cruelty of the ichneumonids troubled philosophers, naturalists, and theologians in the 19th century, who found the parasitoid life cycle inconsistent with the notion of a world created by a loving and benevolent God.[40] Charles Darwin found the example of the Ichneumonidae so troubling that it contributed to his increasing doubts about the nature and existence of a Creator. In an 1860 letter to the American naturalist Asa Gray, Darwin wrote:

I own that I cannot see as plainly as others do, and as I should wish to do, evidence of design and beneficence on all sides of us. There seems to me too much misery in the world. I cannot persuade myself that a beneficent and omnipotent God would have designedly created the Ichneumonidae with the express intention of their feeding within the living bodies of Caterpillars, or that a cat should play with mice.[41]

Morphology

Megarhyssa greenei female

Megarhyssa greenei female
Morphology of the head and its processes: (А) head capsule; (В) antenna; (С) mandible[42]

Morphology of the head and its processes: (А) head capsule; (В) antenna; (С) mandible[42]
Morphology of the thorax (D)[42]

Morphology of the thorax (D)[42]
Morphology of the abdomen and processes of the thorax: (E) front wing; (F) leg III; (G) abdomen of female[42]

Morphology of the abdomen and processes of the thorax: (E) front wing; (F) leg III; (G) abdomen of female[42]