Harpalus

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Adephaga

Familia: Carabidae
Subfamilia: Harpalinae
Supertribus: Harpalitae
Tribus: Harpalini
Subtribus: Harpalina
Genus: Harpalus

Subgenera (11): Baeticoharpalus – Cephalophonus – Cryptophonus – Glanodes – Harpalus – Loboharpalus – Opadius – Plectralidus – Pseudoophonus – Semiophonus – Zangoharpalus

[source: Carabidae of the World]
Overview of species

H. abolitus – H. actiosus – H. acupalpoides – H. adenticulatus – H. aeneipennis – H. aenigma – H. aequicollis – H. affinis – H. agakhaniantzi – H. agnatus – H. akinini – H. alajensis – H. albanicus – H. alexandrae – H. alexeevi – H. alienus – H. alpivagus – H. amarellus – H. amariformis – H. amplicollis – H. amputatus – H. anactus – H. anatolicus – H. angulatus – H. angustipennis – H. angustitarsis – H. animosus – H. anisodactyliformis – H. antonowi – H. anxioides – H. anxius – H. aogashimensis – H. apache – H. araraticus – H. arcuatus – H. arnoldii – H. ascetes – H. asemus – H. aterrimus – H. atratus – H. atrichatus – H. atripes – H. attenuatus – H. australasiae – H. autumnalis – H. azumai – H. babai – H. badakschanus – H. baleensis – H. balli – H. basanicus – H. basilewskyi – H. basuto – H. bellieri – H. beneshi – H. bitinctus – H. brachypterus – H. brevicornis – H. brevis – H. breviusculus – H. bruckmanni – H. brunnipes – H. bucharicus – H. bungii – H. burmeisteri – H. caeruleatus – H. caiphus – H. calathoides – H. calceatus – H. caliginosus – H. capicola – H. capito – H. cardoni – H. caspius – H. cautus – H. cephalotes – H. chalcentus – H. chalcopterus – H. chasanensis – H. chengjiangensis – H. chiloschizontus – H. chobautianus – H. chrysopus – H. cilihumerus – H. circumpunctatus – H. cisteloides – H. cohni – H. colchicus – H. compar – H. compressus – H. conditus – H. constrictus – H. contemptus – H. convexus – H. cordifer – H. cordifer – H. coreanus – H. corporosus – H. corpulentus – H. corrugatus – H. crates – H. cupreus – H. cursorius – H. cyanopterus – H. cyclogonus – H. cyrenaicus – H. danieli – H. davidi – H. davidianus – H. decipiens – H. defector – H. deletus – H. demelti – H. desertus – H. dilatatus – H. diligens – H. dimidiatus – H. disaogashimensis – H. disimuciulus – H. dispar – H. dissitus – H. distinguendus – H. diversicollis – H. dubius – H. dudkoi – H. durangoensis – H. ebeninus – H. egenus – H. egorovi – H. ellipsis – H. eous – H. erraticus – H. erosus – H. erythropus – H. euchlorus – H. ewaldi – H. excavatus – H. exiguus – H. famelicus – H. farkaci – H. faunus – H. ferghanensis – H. fimetarius – H. flavescens – H. flavicornis – H. fokienensis – H. foveiger – H. franzi – H. frater – H. fraternus – H. froelichii – H. fukiensis – H. fulgens – H. fulvicornis – H. fulvilabris – H. fulvipennis – H. fulvus – H. fuscicornis – H. fuscipalpis – H. fuscipennis – H. fuscoaeneus – H. ganssuensis – H. giacomazzoi – H. gilgil – H. gisellae – H. glasunovi – H. gravis – H. gregoryi – H. grilli – H. griseus – H. hartmanni – H. hatchi – H. hauserianus – H. heeri – H. herbivagus – H. heyrovskyi – H. hiekei – H. hirtipes – H. honestus – H. hospes – H. huachuca – H. hybridus – H. hypogeomysis – H. idiotus – H. imerinae – H. impressicollis – H. impressus – H. inconcinnus – H. indianus – H. indicola – H. indicus – H. indigens – H. inexspectatus – H. ingenuus – H. innocuus – H. italus – H. janinae – H. jeanneli – H. jordanus – H. jureceki – H. kabakianus – H. kabakovi – H. kadleci – H. kadyrbekovi – H. kagyzmanicus – H. kailiensis – H. kandaharensis – H. karamani – H. karakorum – H. katiae – H. kazanensis – H. kaznakovi – H. kibonoti – H. kirgisicus – H. kiritshenkoi – H. klapperichi – H. kmecoi – H. knorri – H. koshiensis – H. kozlovi – H. kryzhanovskii – H. kunarensis – H. laetus – H. laevipes – H. lama – H. lampronotus – H. lateralis – H. laticeps – H. latus – H. lederi – H. lethierryi – H. lewisii – H. lheritieri – H. liasinus – H. liobasis – H. litigiosus – H. longihornus – H. longipalmatus – H. lopezi – H. lugubris – H. lumbaris – H. luteicornis – H. lutshniki – H. maceratus – H. macronotus – H. madagascariensis – H. mairei – H. major – H. makhekensis – H. manas – H. marginellus – H. mariae – H. martini – H. masoreoides – H. massarti – H. mauritanicus – H. medvedevi – H. megacephalus – H. meghalayensis – H. melancholicus – H. melaneus – H. meridianus – H. merkli – H. metallinus – H. metarsius – H. meteorus – H. michaili – H. michailovi – H. microdemas – H. microthorax – H. miles – H. minutulus – H. mitridati – H. mlynari – H. modestus – H. morvani – H. muciulus – H. nanniscus – H. natalensis – H. natalicus – H. neglectus – H. nevadensis – H. nigrans – H. nigritarsis – H. numidicus – H. nuperus – H. nyassicus – H. obliquus – H. oblitus – H. obnixus – H. obtusiusculus – H. obtusus – H. ochropus – H. oodioides – H. opacipennis – H. optabilis – H. oustaleti – H. ovtshinnikovi – H. pachys – H. pallidipennis – H. pallipes – H. parasinuatus – H. paratus – H. parvulus – H. pastor – H. pecinai – H. penglainus – H. pensylvanicus – H. periglabellus – H. petri – H. picipennis – H. pilosus – H. plancyi – H. platynotus – H. pleistocenicus – H. plenalis – H. politus – H. polyglyptus – H. poncei – H. potanini – H. poussereaui – H. praecurrens – H. praticola – H. procognatus – H. progrediens – H. prosperus – H. protractus – H. providens – H. pseudoasemus – H. pseudohauserianus – H. pseudophonoides – H. pseudoserripes – H. pseudotinctulus – H. pterostichus – H. puetzi – H. pulchrinulus – H. pulvinatus – H. pumilus – H. punctatostriatus – H. puncticeps – H. punctipennis – H. pusillus – H. pygmaeus – H. quadratus – H. raphaili – H. redivivus – H. reflexus – H. reitteri – H. remboides – H. retractus – H. reversus – H. rhemboides – H. rivalsi – H. roninus – H. rotundus – H. rougemonti – H. rubefactus – H. rubripes – H. rufipalpis – H. rufipes – H. rufiscapus – H. rufitarsoides – H. rufomarginatus – H. rumelicus – H. salinulus – H. salinus – H. sanctaehelenae – H. saxicola – H. scabripectus – H. schaumii – H. schlotheimi – H. semenowi – H. semipunctatus – H. sericatus – H. serripes – H. servus – H. setiporus – H. seyrigi – H. siculus – H. signaticornis – H. silipes – H. simplicidens – H. singularis – H. sinicus – H. sinis – H. sinuatipennis – H. sinuatus – H. smaragdinus – H. smyrnensis – H. solitaris – H. somereni – H. somnulentus – H. spadiceus – H. spretus – H. spurius – H. stephani – H. stevenii – H. stevensi – H. stierlini – H. stoetznerianus – H. strenuus – H. stricticollis – H. stygius – H. subaeneus – H. subcylindricus – H. subtruncatus – H. suensoni – H. sulphuripes – H. sundaicus – H. sushenicus – H. suturangulus – H. szalliesi – H. tabidus – H. taciturnus – H. tadorcus – H. tangutorum – H. tardigradus – H. tardus – H. tarsalis – H. tenebrosus – H. tenuestriatus – H. terrestris – H. texanus – H. tjanschanicus – H. tibeticus – H. tichonis – H. tinctulus – H. tiridates – H. tithonus – H. torosensis – H. torridoides – H. trichophorus – H. tridens – H. triseriatus – H. turcicus – H. turmalinus – H. udege – H. ulomaeformis – H. uniformis – H. ussuricus – H. ussuriensis – H. vagans – H. vanemdeni – H. venator – H. ventralis – H. vereschaginae – H. vernicosus – H. veterum – H. viridanus – H. viridicupreus – H. vittatus – H. wadiensis – H. whitfieldi – H. wohlberedti – H. xanthopus – H. xinjiangensis – H. yinchuanensis – H. zabroides – H. zhdankoi

[source: Lorenz, W. 2018. CarabCat, 2018-12]
Name

Harpalus Latreille, 1802

Source: Latreille 1802: 98 ZooBank: act:CF0F53C5-86F9-4676-9599-67D9AFB5F04E
Genus: masculine
Type species: Carabus proteus Paykull, 1790 = Harpalus affinis (Schrank, 1781)
Fixation: subsequent designation by Andrewes, 1935: 19

References
Primary references

Latreille, P.A. 1802. Histoire naturelle, générale et particulière des crustacés et des insectes. Ouvrage faisant suite à l’histoire naturelle générale et particulière, composée par Leclerc de Buffon, et rédigée par C.S. Sonnini, membre de plusieurs sociétés savantes. Familles naturelles des genres. Tome troisième. F. Dufart, Paris, xii + pp. 13–467 + [1 (errata)]. BHL Reference page. : 98

Additional references

Andrewes, H. E. 1935: On the genotypes of British Carabidae. — II. Annals and Magazine of Natural History (10), 16: 12–25.

Links

Harpalus – Taxon details on Biological Library (BioLib).
Harpalus – Taxon details on Global Biodiversity Information Facility (GBIF).
Lorenz, W. 2018. CarabCat: Global database of ground beetles (version Oct 2017). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P.M., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2018. Species 2000 & ITIS Catalogue of Life, 20th December 2017. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on December 24, 2018]

Vernacular names
čeština: Kvapník
Deutsch: Schnelläufer
lietuvių: Vikriažygiai
русский: Бегун Настоящий / Жужелица зерноядная

Harpalus is a genus of ground beetle first described by Pierre André Latreille in 1802.[1]
Species

Harpalus contains the following 409 species:[2]

Harpalus actiosus Casey, 1914
Harpalus acupalpoides Reitter, 1900
Harpalus aeneipennis (Faldermann, 1836)
Harpalus aenigma (Tschitscherine, 1897)
Harpalus aequicollis Motschulsky, 1844
Harpalus affinis (Schrank, 1781)
Harpalus agakhaniantzi (Mikhailov, 1972)
Harpalus agnatus Reiche, 1849
Harpalus akinini Tschitscherine, 1895
Harpalus alajensis Tschitscherine, 1898
Harpalus albanicus Reitter, 1900
Harpalus alexandrae Kataev, 1990
Harpalus alexeevi Kataev, 1990
Harpalus alienus Bates, 1878
Harpalus alpivagus Tschitscherine, 1899
Harpalus amarellus Bates, 1891
Harpalus amariformis Motschulsky, 1844
Harpalus amplicollis Menetries, 1848
Harpalus amputatus Say, 1830
Harpalus anatolicus Tschitscherine, 1898
Harpalus angulatus Putzeys, 1878
Harpalus angustipennis Boheman, 1848
Harpalus angustitarsis Reitter, 1887
Harpalus animosus Casey, 1924
Harpalus anisodactyliformis Solsky, 1874
Harpalus antonowi (Tschitscherine, 1898)
Harpalus anxioides Kataev, 1991
Harpalus anxius (Duftschmid, 1812)
Harpalus aogashimensis (Habu, 1957)
Harpalus apache Kataev, 2010
Harpalus araraticus Mlynar, 1979
Harpalus arcuatus Tschitscherine, 1898
Harpalus arnoldii Kataev, 1988
Harpalus ascetes Kataev & Wrase, 1997
Harpalus asemus Basilewsky, 1946
Harpalus asphaltinus Roth, 1851
Harpalus aterrimus Casey, 1914
Harpalus atratus Latreille, 1804
Harpalus atrichatus Hatch, 1949
Harpalus atripes Casey, 1914
Harpalus attenuatus Stephens, 1828
Harpalus australasiae Dejean, 1829
Harpalus autumnalis (Duftschmid, 1812)
Harpalus azumai Habu, 1968
Harpalus babai Habu, 1973
Harpalus badakschanus Jedlicka, 1956
Harpalus baleensis Clarke, 1973
Harpalus balli Noonan, 1991
Harpalus basanicus J.R. Sahlberg, 1911
Harpalus basilewskyi Facchini, 2003
Harpalus basuto Basilewsky, 1958
Harpalus bellieri Reiche, 1861
Harpalus beneshi Kataev & Wrase, 1997
Harpalus brachypterus Tschitcherine, 1898
Harpalus brevicornis Germar, 1824
Harpalus brevis Motschiulsky, 1844
Harpalus breviusculus Chaudoir, 1846
Harpalus brunnipes Dejean, 1829
Harpalus bucharicus Tschitscherine, 1898
Harpalus bungii Chaudoir, 1844
Harpalus caeruleatus Bates, 1878
Harpalus caiphus Reiche & Saulcy, 1855
Harpalus calathoides Motschulsky, 1844
Harpalus calceatus (Duftschmid, 1812)
Harpalus caliginosus (Fabricius, 1775)
Harpalus capicola Dejean, 1829
Harpalus capito A.Morawitz, 1862
Harpalus cardoni Antoine, 1922
Harpalus caspius (Steven, 1806)
Harpalus cautus Dejean, 1829
Harpalus cephalotes Fairmaire & Laboulbene, 1854
Harpalus chalcentus Bates, 1873
Harpalus chalcopterus Jeannel, 1948
Harpalus chasanensis Lafer, 1989
Harpalus chobautianus Lutshnik, 1922
Harpalus chrysopus Reitter, 1887
Harpalus circumpunctatus Chaudoir, 1846
Harpalus cisteloides Motschulsky, 1844
Harpalus clarkei Kataev & Schmidt, 2020
Harpalus cohni Ball, 1972
Harpalus colchicus Motschulsky, 1850
Harpalus compar Leconte, 1848
Harpalus compressus Motschulsky, 1844
Harpalus contemptus Dejean, 1829
Harpalus convexus Faldermann, 1836
Harpalus cordifer Notman, 1919
Harpalus coreanus (Tschitscherine, 1895)
Harpalus corporosus (Motschulsky, 1861)
Harpalus corpulentus (Casey, 1914)
Harpalus corrugatus Basilewsky, 1958
Harpalus crates Bates, 1883
Harpalus cupreus Dejean, 1829
Harpalus cursorius Peringuey, 1896
Harpalus cyanopterus (Tschitscherine, 1897)
Harpalus cyclogonus Chaudoir, 1844
Harpalus cyrtonotoides Notman, 1919
Harpalus danieli Reitter, 1900
Harpalus davidi (Tschitscherine, 1897)
Harpalus davidianus Tschitscherine, 1903
Harpalus decipiens Dejean, 1829
Harpalus defector Peringuey, 1896
Harpalus desertus Leconte, 1859
Harpalus diligens Tschitscherine, 1898
Harpalus dimidiatus (P. Rossi, 1790)
Harpalus disimuciulus Huang; Lei; Yan & Hu, 1996
Harpalus dispar Dejean, 1829
Harpalus dissitus Antoine, 1931
Harpalus distinguendus (Duftschmid, 1812)
Harpalus diversicollis Basilewsky, 1958
Harpalus dubius Boheman, 1848
Harpalus dudkoi Kataev, 2011
Harpalus durangoensis Bates, 1891
Harpalus ebeninus Heyden, 1870
Harpalus egenus Dejean, 1829
Harpalus egorovi Lafer, 1989
Harpalus ellipsis Leconte, 1848
Harpalus elliptipennis Facchini, 2003
Harpalus eous Tschitscherine, 1901
Harpalus eraticus Say, 1823
Harpalus erosus Mannerheim, 1825
Harpalus erythropus Dejean, 1829
Harpalus euchlorus Menetries, 1839
Harpalus exiguus Boheman, 1848
Harpalus famelicus Tschitscherine, 1898
Harpalus farkaci Kataev & Wrase, 1995
Harpalus faunus Say, 1823
Harpalus ferghanensis Tschitscherine, 1899
Harpalus fimetarius Dejean, 1829
Harpalus flavescens (Piller & Mitterpacher, 1783)
Harpalus flavicornis Dejean, 1829
Harpalus fokienensis Schauberger, 1930
Harpalus foveiger Tschitscherine, 1895
Harpalus franzi Mateu, 1954
Harpalus frater Chaudoir, 1876
Harpalus fraternus Leconte, 1852
Harpalus froelichii Sturm, 1818
Harpalus fukiensis Jedlicka, 1956
Harpalus fulvicornis Thunberg, 1806
Harpalus fulvilabris Mannerheim, 1853
Harpalus fulvipennis Chaudoir, 1843
Harpalus fulvus Dejean, 1829
Harpalus fuscicornis Menetries, 1832
Harpalus fuscipalpis Sturm, 1818
Harpalus fuscipennis Wiedemann, 1825
Harpalus fuscoaeneus Dejean, 1829
Harpalus ganssuensis Semenov, 1889
Harpalus giacomazzoi Kataev & Wrase, 1996
Harpalus gilgil Basilewsky, 1946
Harpalus gisellae Csiki, 1932
Harpalus glasunovi Kataev, 1987
Harpalus gravis Leconte, 1858
Harpalus gregoryi Alluaud, 1917
Harpalus grilli Kataev, 2002
Harpalus griseus (Panzer, 1796)
Harpalus hartmanni Kataev, 2001
Harpalus hatchi Ball & Anderson, 1962
Harpalus hauserianus Schauberger, 1929
Harpalus herbivagus Say, 1823
Harpalus heyrovskyi Jedlicka, 1928
Harpalus hiekei Kataev & Wrase, 2010
Harpalus hirtipes (Panzer, 1796)
Harpalus honestus (Dufischmid, 1812)
Harpalus hospes Sturm, 1818
Harpalus huachuca Ball, 1972
Harpalus hybridus Boheman, 1848
Harpalus idiotus Bates, 1889
Harpalus impressus Roth, 1851
Harpalus inconcinnus Chaudoir, 1876
Harpalus indianus Csiki, 1932
Harpalus indicola Bates, 1878
Harpalus indicus Bates, 1891
Harpalus indigens Casey, 1924
Harpalus inexspectatus Kataev, 1989
Harpalus ingenuus Tschitscherine, 1898
Harpalus innocuus Leconte, 1863
Harpalus italus Schaum, 1860
Harpalus janinae Jeanne, 1984
Harpalus jeanneli Basilewsky, 1946
Harpalus jordanus Jedlicka, 1964
Harpalus jureceki (Jedlicka, 1928)
Harpalus kabakianus Kataev, 1988
Harpalus kabakovi Kataev, 1987
Harpalus kadleci Kataev & Wrase, 1995
Harpalus kadyrbekovi Kataev, 1988
Harpalus kagyzmanicus Kataev, 1984
Harpalus kandaharensis Jedlicka, 1956
Harpalus karamani Apfelbeck, 1902
Harpalus karokorum Jedlicka, 1958
Harpalus katiae F.Battoni, 1985
Harpalus kazanensis Jedlicka, 1958
Harpalus kaznakovi Kataev & Wrase, 1997
Harpalus kibonoti Alluaud, 1926
Harpalus kirgisicus Motschulsky, 1844
Harpalus kiritshenkoi Kataev, 1990
Harpalus klapperichi Jedlicka, 1956
Harpalus kmecoi Facchini, 2003
Harpalus koshiensis Kirschenhofer, 1992
Harpalus kozlovi Kataev, 1993
Harpalus kryzhanovskii Kataev, 1988
Harpalus kunarensis Kataev, 1993b: 129
Harpalus laetus Reiche, 1843
Harpalus laevipes Zetterstedt, 1828
Harpalus lama Kataev & Wrase, 1997
Harpalus lateralis Dejean, 1829
Harpalus laticeps Leconte, 1850
Harpalus latus (Linne, 1758)
Harpalus lederi Tschitscherne, 1899
Harpalus lethierryi Reiche, 1860
Harpalus lewisii Leconte, 1865
Harpalus liobasis Chaudoir, 1868
Harpalus litigiosus Dejean, 1829
Harpalus longicollis Leconte, 1848
Harpalus longipalmatus Mordkovitsh, 1969
Harpalus lopezi Serrano & Lencina, 2008
Harpalus lugubris Boheman, 1848
Harpalus lumbaris Mannerheim, 1825
Harpalus luteicornis (Dufischmid, 1812)
Harpalus lutshniki Schauberger, 1932
Harpalus macronotus Tschitscherine, 1893
Harpalus mairei Peyerimhoff, 1928
Harpalus major (Motschulsky, 1850)
Harpalus makhekensis Basilewsky, 1958
Harpalus manas Kataev, 1990c: 400
Harpalus marginellus Gyllenhal, 1827
Harpalus mariae Kataev, 1997
Harpalus martini Vandyke, 1926
Harpalus masoreoides Bates, 1878
Harpalus massarti Burgeon, 1935
Harpalus mauritanicus Gaubil, 1844
Harpalus medvedevi Kataev, 2006
Harpalus megacephalus Leconte, 1848
Harpalus meghalayensis Kataev, 2001
Harpalus melancholicus Dejean, 1829
Harpalus melaneus Bates, 1878
Harpalus meridianus Andrewes, 1923
Harpalus merkli Kataev, 2021
Harpalus metallinus Menetries, 1836
Harpalus metarsius Andrewes, 1930
Harpalus meteorus Basilewsky, 1946
Harpalus michaili Kataev, 1990
Harpalus michailovi Kataev, 1987
Harpalus microdemas Schauberger, 1932
Harpalus microthorax (Motschulsky, 1849)
Harpalus miles Peringuey, 1896
Harpalus minutissimus Facchini, 2015
Harpalus minutulus Kataev & Liang, 2004
Harpalus mitridati Pliginskiy, 1915
Harpalus mlynari Kataev, 1990
Harpalus modestus Dejean, 1829
Harpalus morvani Kataev, 2001
Harpalus nanniscus Peringuey, 1896
Harpalus natalensis Boheman, 1848
Harpalus natalicus Peringuey, 1896
Harpalus neglectus Audinet-Serville, 1821
Harpalus nevadensis K. Daniel & J. Daniel, 1898
Harpalus nigrans A. Morawitz, 1862
Harpalus nigritarsis C.R. Sahlberg, 1827
Harpalus numidicus Bedel, 1893
Harpalus nyassicus Basilewsky, 1946
Harpalus obliquus G.Horn, 1880
Harpalus oblitus Dejean, 1829
Harpalus obnixus Casey, 1924
Harpalus ochropus Kirby, 1837
Harpalus oodioides Dejean, 1829
Harpalus opacipennis (Haldemann, 1843)
Harpalus optabilis Dejean, 1829
Harpalus ovtshinnikovi Kataev, 1990
Harpalus paivanus Wollaston, 1867
Harpalus pallidipennis A. Morawitz, 1862
Harpalus pallipes Chaudoir, 1837
Harpalus parallelocollis Facchini, 2003
Harpalus parasinuatus Kataev & Liang, 2007
Harpalus paratus Casey, 1924
Harpalus parvulus Dejean, 1829
Harpalus pastor Motschulsky, 1844
Harpalus pecinai Hovorka, 2006
Harpalus pensylvanicus (Degeer, 1774)
Harpalus perlucens Bates, 1878
Harpalus petri Tschitscherine, 1902
Harpalus picipennis (Duftschmid, 1812)
Harpalus plancyi Tschitscherine, 1897
Harpalus platynotus Bates, 1873
Harpalus plenalis Casey, 1914
Harpalus politus Dejean, 1829
Harpalus polyglyptus Schaum, 1862
Harpalus poncei Will, 2001
Harpalus potanini Tschitscherine, 1906
Harpalus poussereaui Facchini & Giachino, 2011
Harpalus praecurrens Schauberger, 1934
Harpalus praticola Bates, 1891
Harpalus procognatus Lorenz, 1998
Harpalus progrediens Schauberger, 1922
Harpalus prosperus Basilewsky, 1972
Harpalus protractus Casey, 1914
Harpalus providens Casey, 1914
Harpalus pseudoasemus Kataev, 2021
Harpalus pseudohauserianus Kataev, 2001
Harpalus pseudophonoides Schauberger, 1930
Harpalus pseudoserripes Reitter, 1900
Harpalus pseudotinctulus Schauberger, 1932
Harpalus pterostichus (Reitter, 1900)
Harpalus puetzi Kataev & Wrase, 1997
Harpalus pulchirinulus Reitter, 1900
Harpalus pulvinatus Menetries, 1848
Harpalus pumilus Sturm, 1818
Harpalus punctatostriatus Dejean, 1829
Harpalus puncticeps (Casey, 1914)
Harpalus punctipennis Mulsant, 1852
Harpalus pusillus (Motschulsky, 1850)
Harpalus pygmaeus Dejean, 1829
Harpalus quadratus Chaudoir, 1846
Harpalus raphaili Kataev, 1997
Harpalus reflexus Putzeys, 1878
Harpalus regressus (Casey, 1914)
Harpalus remboides Solsky, 1874
Harpalus retractus Leconte, 1863
Harpalus reversus Casey, 1924
Harpalus rivalsi Jeannel, 1948
Harpalus roninus Bates, 1873
Harpalus rotundus Facchini, 2003
Harpalus rougemonti Clarke, 1973
Harpalus rubefactus Bates, 1873
Harpalus rubripes (Dufischmid, 1812)
Harpalus rufipalpis Sturm, 1818
Harpalus rufipes (Degeer, 1774)
Harpalus rufiscapus Gebler, 1833
Harpalus rufitarsoides Schauberger, 1934
Harpalus rufomarginatus (Boheman, 1848)
Harpalus rumelicus Apfelbeck, 1904
Harpalus salinulus Reitter, 1900
Harpalus salinus Dejean, 1829
Harpalus sanctaehelenae Basilewsky, 1972
Harpalus saxicola Dejean, 1829
Harpalus schaumii Wollaston, 1864
Harpalus semenowi Tschitscherine, 1901
Harpalus semipunctatus Dejean, 1829
Harpalus sericatus Tschitscherine, 1906
Harpalus serripes (Quensel, 1806)
Harpalus servus (Duftschmid, 1812)
Harpalus setiporus (Reitter, 1894)
Harpalus siculus Dejean, 1829
Harpalus signaticornis (Duftschmid, 1812)
Harpalus silipes Dejean, 1831
Harpalus simplicidens Schauberger, 1929
Harpalus singularis Tschitscherine, 1906
Harpalus sinicus Hope, 1845
Harpalus sinuaticollis Facchini, 2015
Harpalus sinuatus Tschitscherine, 1893
Harpalus smaragdinus (Duftschmid, 1812)
Harpalus smyrnensis Heyden, 1888
Harpalus solitaris Dejean, 1829
Harpalus somereni Basilewsky, 1946
Harpalus somnulentus Dejean, 1829
Harpalus spadiceus Dejean, 1829
Harpalus spretus Peringuey, 1896
Harpalus spurius Peringuey, 1896
Harpalus stephani Ball, 1972
Harpalus stevenii Dejean, 1829
Harpalus stevensi Kataev, 2011
Harpalus stoetznerianus Schauberger, 1932
Harpalus strenuus Tschitscherine, 1898
Harpalus subaeneus Boheman, 1848
Harpalus subcylindricus Dejean, 1829
Harpalus subphaedrus Basilewsky, 1946
Harpalus subtruncatus Chaudoir, 1846
Harpalus suensoni Kataev, 1997
Harpalus sulphuripes Germar, 1824
Harpalus sushenicus Kataev, 1990
Harpalus suturangulus Reitter, 1887
Harpalus szalliesi Kataev & Wrase, 1995
Harpalus taciturnus Dejean, 1829
Harpalus tadorcus Ball, 1972
Harpalus tangutorum Kataev, 1993
Harpalus tardus (Panzer, 1796)
Harpalus tarsalis Mannenheim, 1825
Harpalus tenebrosus Dejean, 1829
Harpalus terrestris (Motschulsky, 1844)
Harpalus texanus Casey, 1914
Harpalus tibeticus Andrewes, 1930
Harpalus tichonis Jakobson, 1907
Harpalus tinctulus Bates, 1873
Harpalus tiridates Reitter, 1900
Harpalus tithonus Reitter, 1900
Harpalus tjanschanicus Semenov, 1889
Harpalus torosensis Jedlicka, 1961
Harpalus torridoides Reitter, 1900
Harpalus trichophorus Tschitscherine, 1897
Harpalus tridens A.Morawitz, 1862
Harpalus triseriatus A. Fleischer, 1897
Harpalus turcicus Jedlicka, 1958
Harpalus turmalinus Erichson, 1847
Harpalus udege Lafer, 1989
Harpalus uniformis (Motschulsky, 1844)
Harpalus ussuricus Mlynar, 1979
Harpalus vanemdeni Schauberger, 1932
Harpalus venator Boheman, 1848
Harpalus ventralis Leconte, 1848
Harpalus vereschaginae Kataev, 1988
Harpalus vernicosus Kataev & Liang, 2007
Harpalus viridanus Motschulsky, 1844
Harpalus viridicupreus Reiche, 1843
Harpalus vittatus Gebler, 1833
Harpalus wadiensis Jedlicka, 1964
Harpalus wohlberedti Emden & Schauberger, 1932
Harpalus xanthopus Gemminger & Harold, 1868
Harpalus yinchuanensis Huang, 1993
Harpalus zabroides Dejean, 1829
Harpalus zhdankoi Kataev, 1990

References

Latreille, Pierre André (1802). "Histoire naturelle, générale et particulière des crustacés et des insectes". 3. F. Dufart. doi:10.5962/bhl.title.15764. OCLC 5050710.
"Harpalus Latreille, 1802". Carabidae of the World. 2022. Retrieved 23 April 2022.