Harpalinae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Adephaga

Familia: Carabidae
Subfamilia: Harpalinae
Supertribus (9): Anthiitae - Chlaeniitae - Ctenodactylitae - Dryptitae - Harpalitae - Lebiitae - Pentagonicitae - Platynitae - Pterostichitae

Overview of tribus (48)

Abacetini - Amorphomerini - Anthiini - Caelostomini - Calophaenini - Catapieseini - Chaetodactylini - Chaetogenyini - Chlaeniini - Cnemalobini - Corsyrini - Cratocerini - Ctenodactylini - Cuneipectini - Cyclosomini - Dercylini - Drimostomatini - Dryptini - Galeritini - Geobaenini - Ginemini - Graphipterini - Harpalini - Helluonini - Hexagoniini - Idiomorphini - Lachnophorini - Lebiini - Licinini - Loxandrini - Melanchitonini - Microcheilini - Morionini - Odacanthini - Omphreini - Oodini - Orthogoniini - Panagaeini - Peleciini - Pentagonicini - Perigonini - Physocrotaphini - Platynini - Pseudomorphini - Pterostichini - Sphodrini - Zabrini - Zuphiini
Overview of genera (1170 + 1†)
A

Abacaelostus – Abacetodes – Abacetus – Abacidus – Abacillius – Abacillodes – Abacoleptus – Abacomorphus – Abacophrastus – Abacops – Abaris – Abax – Abrodiella – Abropus – Acalathus – Acanthoferonia – Acanthoodes – Achaetocephala – Acinopus – Acrogenys – Actenoncus – Actenonyx – Actodus – Acupalpus – Acutosternus – Adelopomorpha – Adelotopus – Adrimus – Aenigma – Aeolodermus – Aepsera – Afrodromius – Afromizonus – Afrotarus – Agastus – Agelaea – Agonica – Agonidium – Agonidus – Agonobembix – Agonocheila – Agonops – Agonoriascus – Agonorites – Agonum – Agra – Alachnothorax – Alkestis – Allardina – Allendia – Allocinopus – Allocota – Allophanes – Allophanopsis – Allosiopelus – Allotriopus – Altagonum – Amara – Amaroschema – Amblops – Amblycoleus – Amblygnathus – Amblystomus – Amelus – Amerinus – Ametroglossus – Amorphomerus – Amphasia – Amphimasoreus – Amphimenes – Amphimenoides – Amphithasus – Analoma – Anasis – Anatrichis – Anaulacus – Anchista – Anchomenidius – Anchomenus – Anchonoderus – Ancystroglossus – Andrewesia – Andrewesinulus – Andrewesius – Angionychus – Anisocnemus – Anisodactylus – Anisostichus – Anomostomus – Anomotariella – Anomotarus – Anoncopeucus – Anoplogenius – Antarctiola – Anthia – Anthracus – Antimerina – Antipionycha – Aparupa – Apenes – Aporesthus – Apristomimus – Apristus – Apsaustodon – Apsidocnemus – Apterodromites – Arame – Archagonum – Archicolliuris – Archicolpodes – Ardistomopsis – Argutoridius – Arhytinus – Aristochroa – Aristochroodes – Aristolebia – Aristopus – Arsinoe – Asakalaphium – Asios – Asklepia – Aspasiola – Assadecma – Astastus – Athrostictus – Atlantomasoreus – Atongolium – Atractonotus – Atranopsis – Atranus – Atrotus – Auchmerus – Aulacolius – Aulacopodus – Australomasoreus – Australovelinda – Austroglyptolenus – Axinopalpus – Axinotoma – Aztecarpalus –
B

Badister – Baeoglossa – Barrymooreana – Barylaus – Barysomus – Bascanus – Basilewskya – Basistichus – Batoscelis – Beckeria – Blackburnia – Blennidus – Blenniventer – Bleusea – Boeomimetes – Bothrocolpodes – Bothynoproctus – Bottchrus – Brachichila – Brachidius – Brachyctis – Brachygnathus – Brachynopterus – Brachyodes – Bradybaenus – Bradycellus – Bradycidus – Brigalowia – Bronislavia – Broter – Bruskespar – Bruskmoal – Bryanites – Buderes –
C

Caecocaelus – Caelostomus – Cainogenion – Calathidius – Calathocosmus – Calathus – Calleida – Calleidomorpha – Callidadelpha – Callidagonum – Callistomimus – Callistus – Calodromius – Calophaena – Calophaenoidea – Calybe – Camptogenys – Camptoscelis – Camptotoma – Capabatus – Carbonellia – Cardiomera – Carenochyrus – Carterus – Casaleius – Castelnaudia – Catacolpodes – Catadromus – Catapiesis – Catascopellus – Catascopus – Cedrorum – Celioinkosa – Cenogmus – Cephalosdrophus – Cephalostichus – Cerabilia – Chaetagonum – Chaetocrepis – Chaetodactyla – Chaetogenys – Chaetosaurus – Chalcochrous – Chaudoirella – Chaudoirina – Chilotomus – Chlaeminus – Chlaenius – Chydaeus – Cinctagonum – Cintaroa – Cistelagonum – Clarencia – Cnemalobus – Coarazuphium – Colasidia – Coleolissus – Colfax – Collagonum – Colliuris – Colpodes – Colpodichius – Colpoides – Colpomimus – Colposphodrus – Colpostoma – Conchitella – Cophosomorpha – Coptia – Coptocarpus – Coptodera – Coptoglossus – Coptoptera – Coptopterella – Coptopterina – Corsyra – Cosmodiscus – Cosnania – Crasodactylus – Craspedophorus – Crassacantha – Crassagena – Cratacanthus – Cratocerus – Cratoferonia – Cratogaster – Cratognathus – Cratosoma – Creagris – Crenulostrigus – Criniventer – Cryptobatis – Cryptocephalomorpha – Cryptocolliuris – Cteatus – Ctenodactyla – Ctenognathus – Cuneipectus – Cyanotarus – Cycloloba – Cyclosomus – Cyclotrachelus – Cylindrocranius – Cylindronotum – Cylindropectus – Cymenopterus – Cymindis – Cymindoidea – Cynthidia – Cyphocoleus – Cypholoba – Cyptomicrus – Cyrtolaus – Cyrtomoscelis – Cyrtopilus –
D

Dactyleurys – Dactylinius – Daer – Dailodontus – Dalatagonum – Daptus – Darodilia – Dasiosoma – Deipyrodes – Deliaesianum – Delinius – Deltocolpodes – Demetrias – Demetrida – Demetriola – Dendragonum – Dercylinus – Dercylus – Derostichus – Diacanthostylus – Diachipteryx – Diachromus – Diamella – Diaphoromerus – Dicaelindus – Dicaelus – Diceromerus – Dicheirotrichus – Dicheirus – Dicranoglossus – Dicranoncus – Dicraspeda – Dicrochile – Dicrodontus – Dilonchus – Dinocolpodes – Dinopelma – Dioryche – Diplocheila – Diploharpus – Dirotus – Dischissus – Discoderus – Discoptera – Disphericus – Distrigidius – Ditomus – Dixus – Do – Dobodura – Dolichocolpodes – Dolichoctis – Dolichodes – Dolichus – Doliodactyla – Dontolobus – Dregus – Dromidea – Dromiops – Dromistomus – Dromius – Dromoceryx – Drymatus – Drymonaxus – Drypta – Dyschiridium – Dyscolus –
E

Eccoptomenus – Eccoptoptera – Ecnomolaus – Ectenognathus – Ectinothorax – Egadyla – Elliptoleus – Endynomena – Enoicus – Eocarterus – Eosphodrus – Epicolpodes – Epigraphodes – Epigraphus – Epikastea – Epimicodema – Epomis – Erectocolliuris – Eremolestes – Eremosphodrus – Erephognathus – Eriophonus – Eripus – Eucaerus – Eucamptognathus – Eucarterus – Eucheila – Euchroa – Eucolliuris – Eudalia – Eudromus – Eujalmenus – Euleptus – Eumara – Eunostus – Euphorticus – Euplynes – Euproctinus – Euryaptus – Eurycoleus – Eurydera – Euryderus – Eurygnathus – Eurystomis – Euschizomerus – Euthenarus – Eutogeneius – Evolenes –
F

Falsodromius – Feostoma – Feroniascus – Feroniola – Feronista – Formosiella – Fortagonum – Foveocrotaphus –
G

Gaioxenus – Galerita – Galiciotyphlotes – Gallerucidia – Gastragonum – Gastrellarius – Gastrogmus – Geobaenus – Geobius – Geodromus – Geoffreyella – Geopinus – Gestroania – Giachinoana – Gidda – Gigadema – Ginema – Glycia – Glyphodactyla – Glyptolenoides – Glyptolenus – Glyptus – Gnathaphanus – Gnopheroides – Gomerina – Goniocellus – Gonogenia – Gourlayia – Graniger – Graphipterus – Grundmannius – Guatemalteca – Gugheorites – Gunvorita – Guyanemorpha – Gynandromorphus –
H

Habragonum – Habutarus – Hakaharpalus – Hannaphota – Haplaner – Haplanister – Haplobothynus – Haplocolpodes – Haploferonia – Haplopeza – Haptoderidius – Haptoderodes – Harpaglossus – Harpalinus – Harpalobrachys – Harpalodiodes – Harpalomimetes – Harpalomorphus – Harpalus – Harpathaumas – Harponixus – Hartonymus – Helluapterus – Helluarchus – Helluo – Helluobrochus – Helluocolpodes – Helluodema – Helluodes – Helluomorpha – Helluomorphoides – Helluonidius – Helluopapua – Helluosoma – Hemiaulax – Hemiplatynus – Hemitelestus – Henrotius – Henvelik – Herculagonum – Heteracantha – Hexachaetus – Hexagonia – Hikosanoagonum – Himalosphodrus – Hippoloetis – Holcaspis – Holcocoleus – Holcoderus – Holconotus – Hololeius – Holoponerus – Holosoma – Homalomorpha – Homalonesiota – Homethes – Hoplizomenus – Hoplolenus – Hormacrus – Horniulus – Hyboptera – Hybopteroides – Hybothecus – Hyperectenus – Hyperion – Hyphaereon – Hypharpax – Hystrichopus – Hystricosphodrus –
I

Idiagonum – Idiastes – Idiocolpodes – Idiomelas – Idiomorphus – Ifridytes – Ildobates – Incagonum – Indiophonus – Infernophilus – Inkosa – Iridagonum – Ischnagonum –
J

Ja – Jocqueius – Jujiroa –
K

Kaffovatus – Kalchdigor – Kar – Kaszabellus – Kenyacus – Kiwiharpalus – Klapperichella – Klepteromimus – Kuceraianum – Kupeharpalus – Kuschelinus –
L

Labocephalus – Lachnocrepis – Lachnoderma – Lachnolebia – Lachnophorus – Lachnothorax – Lacordairia – Laemostenus – Laevagonum – Lasiocera – Lassalleianum – Lebia – Lebidema – Lebidia – Lebiomorphica – Lebistina – Lebistinida – Lecanomerus – Leiradira – Leleupidia – Leleuporites – Lelis – Lepcha – Leptagonum – Leptocolpodes – Leptosarcus – Leptotrachelon – Leptotrachelus – Lesticus – Lestignathus – Letouzeya – Leuropus – Liagonum – Liamegalonychus – Lichnasthenus – Licinopsis – Licinus – Lindrothius – Liochirus – Liocolpodes – Liodaptus – Lioholus – Lionedya – Lionychus – Liopasa – Lioptera – Lipostratia – Litarthrum – Lithagonum – Lobatodes – Lobobrachus – Lobocolpodes – Lobodontidius – Lobodontulus – Lobodontus – Lonchosternus – Lophidius – Lophoglossus – Lorestania – Lorostema – Loxandrus – Loxocrepis – Loxodactylus – Loxogenius – Loxoncus – Lucicolpodes –
M

Machozetus – Macrocheilus – Macroprotus – Maculagonum – Madapelmus – Madecassellus – Madecassina – Manumorpha – Maoriharpalus – Marsyas – Martyr – Mascarenhia – Masoreus – Matabele – Mateuellus – Mecynognathus – Megadromus – Megalebia – Megalonychus – Megaloodes – Megamorio – Meladroma – Melanchiton – Melanchrous – Meleagros – Meleagros – Menarus – Merizomena – Meroctenus – Meropalpus – Mesocolpodes – Mesolestes – Metabacetus – Metablus – Metacolpodes – Metadromius – Metallanchista – Metallica – Metascopus – Metaxellus – Metaxidius – Metaxymorphus – Metaxys – Metazuphium – Metius – Metoncidus – Mexisphodrus – Micrixys – Microcephalus – Microcheila – Microcosmodes – Microdaccus – Microderes – Microferonia – Microlestes – Microlestodes – Microodes – Microzargus – Miltodes – Mimocolliuris – Mimodromius – Mimophilorhizus – Mimovelindopsis – Minuphloeus – Minuthodes – Miquihuana – Mirachlaenius – Mireius – Miscelus – Mischocephalus – Mizotrechus – Mnuphorus – Mochtheroides – Mochtherus – Molopidius – Molopinus – Molops – Monnea – Monodryxus – Montagonum – Morimotoidius – Morion – Morionidius – Moriosomus – Mormolyce – Mormolycina – Morphaeus – Morphodaclyla – Myas – Myrmecodemus –
N

Nanodiodes – Nebriagonum – Nebrioferonia – Negrea – Negreum – Nelidus – Nemaglossa – Nemotarsus – Neoaulacoryssus – Neobatenus – Neocolpodes – Neocoptodera – Neodendragonum – Neodiachipteryx – Neoeudalia – Neoferonia – Neoglyptus – Neohelluo – Neohyparpalus – Neoleleupidia – Neomegalonychus – Neoodes – Neoorthogonius – Neophygas – Neotalus – Nepalorthogonius – Nesarpalus – Nesiocolpodes – Nesites – Netrodera – Nipponagonum – Nipponoharpalus – Nipponosynuchus – Nirmala – Nornalupia – Notabax – Notagonum – Nothodaptus – Notiobia – Notocolpodes – Notolestus – Notonomus – Notoplatynus – Notopseudomorpha – Notospeophonus – Novillidius – Nurus – Nycteis –
O

Oberthueria – Odacantha – Odontocarus – Odontomasoreus – Oechalius – Oecornis – Oedesis – Oesyperus – Ogmophora – Oilea – Olisthopus – Omestes – Omobrus – Omophagus – Omostropus – Omphra – Omphreoides – Omphreus – Onota – Onotokiba – Onycholabis – Onypterygia – Oodes – Oodinkosa – Oodinus – Ooidius – Oosoma – Ophonichius – Ophoniscus – Ophonus – Oreodicastes – Oribazus – Orionella – Orophicus – Orthobasis – Orthocerodus – Orthogenium – Orthogonius – Orthomus – Orthotrichus – Oscadytes – Otoglossa – Oxoides – Oxycentrus – Oxycrepis – Oxyglychus – Oxygonium – Oxyodontus – Oxypselaphus –
P

Pachybatenus – Pachycaecus – Pachycallida – Pachycarus – Pachyferonia – Pachylebia – Pachylebioides – Pachymorphus – Pachyroxochus – Pachythecus – Pachytrachelus – Panagaeus – Panagrius – Pangus – Paniestichus – Parabaris – Parabroscus – Parachlaenius – Paracolpodes – Paradromius – Paraeutrichopus – Paraglycia – Paraleleupidia – Paraliagonum – Paralophidius – Paramecus – Paramegalonychus – Paranchodemus – Paranchus – Paranurus – Parapionycha – Parascopodes – Parasiopelus – Parazuphium – Paregaploa – Paregraphus – Parena – Pareuryaptus – Pareurydera – Parhypates – Paridiagonum – Parophonus – Parorthomus – Paulianites – Paulianolebia – Paulianoscirtus – Paussotropus – Pawgammm – Pectinitarsus – Pediomorphus – Pedius – Pelecium – Peliocypas – Pelmatellus – Pentagonica – Penthophonus – Penthus – Pephrica – Percolaus – Percus – Pericalus – Perigona – Peripristus – Perissostomus – Perochnoristhus – Peronomerus – Petrimagnia – Peyrieraselus – Phacocerus – Phaenaulax – Phanagnathus – Philophlaeus – Philorhizus – Phloeoxena – Phobophorus – Pholeodytes – Phorticosomus – Phyrometus – Physocrotaphus – Physodera – Physolaesthus – Physomerus – Piesmus – Piezia – Pionycha – Pioprosopus – Piosoma – Plagiopyga – Plagiotelum – Planetes – Platyagonum – Platycoelus – Platyderus – Platyhelluo – Platylytron – Platymetopsis – Platymetopus – Platynodes – Platynus – Platysmodes – Platytarus – Platyxythrius – Plaumannium – Pleuracanthus – Plicagonum – Plocamostethus – Plochionus – Poecilus – Pogonodaptus – Pogonoglossus – Polistichus – Pollicobius – Polpochila – Polyaulacus – Polychaetus – Polydamasium – Ponapagonum – Pontonoa – Porocara – Potamagonum – †Praeanchodemus – Praepristus – Prakasha – Prionognathus – Pristacrus – Pristosia – Procletodema – Procletus – Proditomus – Progonochaetus – Promecoptera – Promegalonychus – Prophenorites – Propionycha – Prosopogmus – Prosphodrus – Prostalomus – Protocolliuris – Protocolpodes – Protopidius – Psammodromius – Psecadius – Psegmatopterus – Pseudabacetus – Pseudabarys – Pseudagonica – Pseudamara – Pseudanchomenus – Pseudanisotarsus – Pseudaptinus – Pseudaristus – Pseudobatenus – Pseudocalleida – Pseudoceneus – Pseudodiachipteryx – Pseudognathaphanus – Pseudohyparpalus – Pseudomasoreus – Pseudomegalonychus – Pseudomenarus – Pseudometabletus – Pseudomonnea – Pseudomorpha – Pseudomyas – Pseudopachylebia – Pseudophorticus – Pseudoplatyderus – Pseudorhysopus – Pseudoselenophorus – Pseudosinurus – Pseudosphaerodes – Pseudotaphoxenus – Pseudotoglossa – Pseudotrechus – Psychristus – Pterostichus – Pterostillichus – Pylartesius –
Q

Quammenis –
R

Rathymus – Reflexisphodrus – Renneria – Rhabdotus – Rhabidius – Rhadine – Rhagadillius – Rhopalomelus – Rhopalostyla – Rhysopus – Rhytiferonia – Rhytisternus – Ripogenites – Rupa –
S

Samiriamorpha – Sarothrocrepis – Sarticus – Scalidion – Schidonychus – Schuelea – Scopodes – Scybalicus – Secatophus – Selenophorus – Selenotichnus – Selina – Sericoda – Serrimargo – Setalidius – Setalimorphus – Setalis – Setolebia – Sfitakantha – Siagonyx – Simodontus – Simous – Singiliomimus – Singilis – †Sinis – Sinocolpodes – Sinurus – Siopelus – Skoeda – Skorlagad – Skouedhirraad – Smeringocera – Sofota – Somalodromius – Somoplatodes – Somoplatus – Somotrichus – Sophroferonia – Speagonum – Speluncarius – Speocolpodes – Speokokosia – Speomolops – Speotarus – Speothalpius – Speozuphium – Sphaerodes – Sphaerodinus – Sphallomorpha – Sphodropsis – Sphodroschema – Sphodrosomus – Sphodrus – Stegazopteryx – Stenidia – Stenocheila – Stenocnemus – Stenocrepis – Stenognathus – Stenolepta – Stenolophidius – Stenolophus – Stenomorphus – Stenoodes – Stenotelus – Stereocerus – Stereostoma – Steropanus – Sthenocranion – Stigmaphorus – Stilboma – Stolonis – Stomis – Stomonaxellus – Straneoa – Straneostichus – Straneotia – Stricklandiana – Stricteripus – Strigia – Strigomerodes – Strigomerus – Stuhlmannium – Styracoderus – Sugimotoa – Syletor – Syllectus – Syntomus – Synuchidius – Synuchus – Systolocranius –
T

Taicona – Takasagoagonum – Tantillus – Tanystoma – Taphoxenus – Tapinopterus – Taridius – Tarsagonum – Tecnophilus – Tefflus – Teiresia – Teradaia – Teratotarsa – Teropha – Tetragonoderus – Tetraleucus – Teukrus – Thermoscelis – Thoasia – Thryptocerus – Thyreochaetus – Thyreopterus – Thysanotus – Tiferonia – Tilius – Tinoderus – Tinognathus – Titaresius – Tostkar – Trachelocyphoides – Trachelocyphus – Trachysarus – Trephionus – Triaenogenius – Tricharnhemia – Trichidema – Trichillinus – Trichis – Trichisia – Trichocoptodera – Trichognathus – Trichopiezia – Trichopselaphus – Trichosternus – Trichotichnus – Trichoxycentrus – Trigonognatha – Trigonothops – Trigonotoma – Triplosarus – Trirammatus – Trogloagonum – Tropicoritus – Trymosternus – Tschitscherinellus – Tuiharpalus – Tuxtlamorpha – Typsiharpalus –
U

Uenanthracus –
V

Vachinius – Valeriaaschero – Velinda – Velindastus – Velindomimus – Velindopsis – Veradia – Vianasia – Violagonum – Vitagonum – Vulcanophilus –
W

Wahlbergiana – Wate –
X

Xanthomelina – Xenion – Xenitenopsis – Xenitenus – Xenodochus – Xenodromius – Xenophonus – Xestagonum – Xestonotus – Xestopus –
Y

Yasunimorpha – Yukihikous –
Z

Zabrus – Zargus – Zariquieya – Zeodera – Zeopoecilus – Zolotarevskyella – Zuphium
Name

Harpalinae Bonelli, 1810

References

Maddison, D.R.; Baker, M.D.; Ober, K.A. 1999: Phylogeny of carabid beetles as inferred from 18S ribosomal DNA (Coleoptera: Carabidae). Systematic entomology, 24: 103–138. DOI: 10.1046/j.1365-3113.1999.00088.x PDF
Ober, K.A. 2002: Phylogenetic relationships of the carabid subfamily Harpalinae (Coleoptera) based on molecular sequence data. Molecular phylogenetics and evolution, 24: 228–248. DOI: 10.1016/S1055-7903(02)00251-8
Ober, K.A., Maddison, D.R. 2008: Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of insect science, 8.63 DOI: 10.1673/031.008.6301

For more multimedia, look at Category:Harpalinae on Wikimedia Commons.
Links

Carabidae of the World

Vernacular names
norsk: Markløpere
русский: Харпалины