Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Adephaga
Familia: Carabidae
Subfamilia: Harpalinae
Supertribus: Harpalitae
Tribus: Harpalini
Subtribus (7): Amblystomina - Anisodactylina - Bradybaenina - Ditomina - Harpalina - Pelmatellina - Stenolophina
Overview of genera (178)
Acinopus – Acupalpus – Afromizonus – Agonidus – Agonoleptus – Allendia – Allocinopus – Allosiopelus – Amblygnathus – Amblystomus – Amerinus – Amphasia – Angionychus – Anisocnemus – Anisodactylus – Anisostichus – Anomostomus – Anoplogenius – Anthracus – Athrostictus – Axinotoma – Aztecarpalus – Barysomus – Batoscelis – Bleusea – Boeomimetes – Bottchrus – Bradybaenus – Bradycellus – Bradycidus – Bronislavia – Carenochyrus – Carterus – Cenogmus – Chilotomus – Chydaeus – Coleolissus – Crasodactylus – Cratacanthus – Cratognathus – Cratosoma – Criniventer – Cyptomicrus – Daptus – Diachromus – Dicheirotrichus – Dicheirus – Dioryche – Discoderus – Ditomus – Dixus – Dregus – Ectinothorax – Egadyla – Eocarterus – Eriophonus – Eucarterus – Euryderus – Euthenarus – Gaioxenus – Geodromus – Geopinus – Gnathaphanus – Goniocellus – Graniger – Gugheorites – Gynandromorphus – Hakaharpalus – Haplaner – Haplanister – Harpalinus – Harpalobrachys – Harpalodiodes – Harpalomimetes – Harpalomorphus – Harpalus – Harpathaumas – Harponixus – Hartonymus – Hemiaulax – Heteracantha – Hippoloetis – Hyphaereon – Hypharpax – Idiomelas – Indiophonus – Kaffovatus – Kenyacus – Kiwiharpalus – Kupeharpalus – Lecanomerus – Liochirus – Liodaptus – Lioholus – Loxoncus – Machozetus – Maoriharpalus – Meroctenus – Microderes – Nemaglossa – Neoaulacoryssus – Neodiachipteryx – Neohyparpalus – Neophygas – Nesarpalus – Nipponoharpalus – Nornalupia – Nothodaptus – Notiobia – Notospeophonus – Odontocarus – Oedesis – Oesyperus – Omostropus – Ooidius – Oosoma – Ophoniscus – Ophonus – Orthogenium – Oxycentrus – Pachycarus – Pachytrachelus – Panagrius – Pangus – Parabaris – Paramecus – Parasiopelus – Paraulacoryssus – Paregaploa – Parophonus – Paulianoscirtus – Pelmatellus – Penthophonus – Penthus – Phanagnathus – Phobophorus – Pholeodytes – Phorticosomus – Phyrometus – Piosoma – Platymetopsis – Platymetopus – Pogonodaptus – Polpochila – Prakasha – Proditomus – Progonochaetus – Pseudanisotarsus – Pseudaristus – Pseudodiachipteryx – Pseudognathaphanus – Pseudohyparpalus – Pseudorhysopus – Pseudoselenophorus – Psychristus – Rhabidius – Rhysopus – Scybalicus – Selenophorus – Selenotichnus – Siopelus – Stenolophidius – Stenolophus – Stenomorphus – Syllectus – Trachysarus – Trichopselaphus – Trichotichnus – Trichoxycentrus – Triplosarus – Tropicoritus – Tschitscherinellus – Tuiharpalus – Typsiharpalus – Uenanthracus – Veradia – Xenodochus – Xenophonus – Xestonotus – ?Diaphoromerus
Name
Harpalini Bonelli, 1810
References
Bonelli, F.A. 1810. Observations Entomologiques, Première partie. Mémoires de l’Academie des Sciences de Turin 18: 21–78, Tabula Synoptica. BHL Reference page.
Martínez-Navarro, E.M.; Galián, J.; Serrano, J. 2005: Phylogeny and molecular evolution of the tribe Harpalini (Coleoptera, Carabidae) inferred from mitochondrial cytochrome-oxidase I. Molecular phylogenetics and evolution, 35(1): 127–146. DOI: 10.1016/j.ympev.2004.11.009
Noonan, G.R. 1976: Synopsis of the supra-specific taxa of the tribe Harpalini (Coleoptera: Carabidae). Quaestiones entomologicae, 12: 3–87.
Shpeley, D., Hunting, W.M. & Ball, G.E. 2017. A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography. ZooKeys 690: 1—195. DOI: 10.3897/zookeys.690.13751. Reference page.
Links
Carabidae of the World
Harpalini is a tribe of a diverse group of ground beetles belonging to the subfamily Harpalinae within the broader family Carabidae. The tribe contains more than 2500 species.[1]
Trichotichnus dichrous
Genera
These 120 genera belong to Harpalini:[2][3][4][5][6]
Subtribe Amblystomina Fauvel, 1889
Amblystomus Erichson, 1837
Anomostomus LaFerté-Sénectère, 1853
Barysomus Dejean, 1829
Oosoma Nietner, 1857
Subtribe Anisodactylina (Lacordaire, 1854)
Amphasia Newman, 1838
Anisodactylus Dejean, 1829
Dicheirus Mannerheim, 1843
Geopinus LeConte, 1847
Notiobia Perty, 1830
Subtribe Ditomina Bonelli, 1810
Bottchrus Jedlibcka, 1935
Bronislavia Semenov, 1891
Carenochyrus Solsky, 1874
Carterus Dejean, 1830
Chilotomus Chaudoir, 1842
Ditomus Bonelli, 1810
Dixus Billberg, 1820
Eocarterus Stichel, 1923
Eucarterus Reitter, 1900
Graniger Motschulsky, 1864
Liochirus Tschitscherine, 1897
Machozetus Chaudoir, 1850
Meroctenus Gemminger & Harold, 1868
Odontocarus
Odotoncarus Solier, 1835
Oedesis Motschulsky, 1850
Orthogenium Chaudoir, 1835
Pachycarus Solier, 1835
Panagrius Andrewes, 1933
Penthus Chaudoir, 1843
Phobophorus
Phorticosomus Schaum, 1863
Proditomus Schauberger, 1934
Pseudaristus Reitter, 1900
Tschitscherinellus Csiki, 1906
Veradia
Xenophonus Müller, 1942
Subtribe Harpalina Bonelli, 1829
Acinopus Dejean, 1821
Afromizonus Basilewsky, 1947
Allosiopelus N.Ito, 1995
Amblygnathus Dejean, 1829
Anisocnemus Chaudoir, 1843
Athrostictus Bates, 1878
Aulacoryssus
Axinotoma Dejean, 1829
Aztecarpalus Ball, 1970
Bleusea Bedel, 1897
Boeomimetes Péringuey, 1896
Bradybaenus Dejean, 1829
Coleolissus Bates, 1892
Cratacanthus Dejean, 1829
Cratognathus Dejean, 1829
Daptus Fischer von Waldheim, 1823
Dioryche W.S.MacLeay, 1825
Discoderus LeConte, 1853
Dregus Motschulsky, 1864
Ectinothorax Alluaud, 1941
Eriophonus Tschitscherine, 1901
Euryderus LeConte, 1846
Geodromus Dejean, 1829
Harpalinus Jeannel, 1946
Harpalobrachys Tschitschérine, 1899
Harpalodiodes Bousquet, 2002
Harpalomorphus Péringuey, 1896
Harpalus Latreille, 1802
Harpathaumas Basilewsky, 1947
Harponixus Basilewsky, 1950
Hartonymus Casey, 1914
Heteracantha Brullé, 1835
Hyphaereon W.S.MacLeay, 1825
Indiophonus N.Ito, 1996
Laparhetes Jeannel, 1946
Liodaptus Bates, 1889
Microderes Faldermann, 1836
Neoaulacoryssus Noonan, 1985
Neodiachipteryx Noonan, 1985
Neohyparpalus Clarke, 1981
Neophygas Noonan, 1976
Nesacinopus Tschitscherine, 1900
Nesarpalus Bedel, 1897
Nipponoharpalus Habu, 1973
Nothodaptus Maindron, 1906
Oesyperus Andrewes, 1923
Omostropus Péringuey, 1896
Ooidius Chaudoir, 1847
Ophoniscus Bates, 1892
Ophonus Dejean, 1821
Oxycentrus de Chaudoir, 1854
Pangus Dejean, 1821
Parasiopelus Basilewsky, 1946
Parophonus Ganglbauer, 1891
Paulianoscirtus Basilewsky, 1976
Penthophonus Reitter, 1900
Phyrometus
Piosoma LeConte, 1847
Platymetopsis Ball & Maddison, 1987
Platymetopus Dejean, 1829
Prakasha Andrewes, 1919
Pseudodiachipteryx Burgeon, 1936
Pseudohyparpalus Basilewsky, 1946
Pseudoselenophorus Péringuey, 1896
Selenophorus Dejean, 1829
Selenotichnus Kataev, 1999
Siopelus Murray, 1859
Stenomorphus Dejean, 1831
Trichopselaphus Chaudoir, 1843
Trichotichnus Morawitz, 1863
Trichoxycentrus N.Ito, 2000
Typsiharpalus Tschitscherine, 1901
Xenodochus Andrewes, 1941
Subtribe Pelmatellina
Pelmatellus Bates, 1882
Subtribe Stenolophina Kirkby, 1837
Acupalpus Latreille, 1829
Agonoleptus Casey, 1914
Amerinus Casey, 1884
Anoplogenius
Bradycellus Erichson, 1837
Dicheirotrichus Jacquelin du Val, 1855
Philodes LeConte, 1861
Pogonodaptus G. Horn, 1881
Polpochila Solier, 1849
Stenolophus Dejean, 1821
References
A. Larochelle and M. C. Lariviere (2005). "Harpalini (Insecta: Coleoptera: Carabidae: Harpalinae". Fauna of New Zealand. 53: 160 pp. Retrieved 5 June 2020.
"Carabidae". GBIF, Global Biodiversity Information Facility. Retrieved 13 November 2021.
"Harpalini Bonelli, 1810". Catalogue of Life. Retrieved 13 November 2021.
"Harpalini tribe Information". BugGuide.net. Retrieved 13 November 2021.
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.
"Harpalini Report". Integrated Taxonomic Information System. Retrieved 13 November 2021.
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