Cyana

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Cladus: Macroheterocera
Superfamilia: Noctuoidea

Familia: Erebidae
Subfamilia: Arctiinae
Tribus: Lithosiini
Subtribus: Nudariina
Genus: Cyana
Subgenera: Australisine – Bizone – Caudovulpecula – C. (Chionaema) – C. (Clerckia) – Comorocyana – C. (Cornutivulpecula) – C. (Cryptanaema) – C. (Cyabarda) – C. (Cyana) – C. (Doliche) – Exotrocha – Frankmuelleria – Gigantovulpecula – C. (Idiovulpecula) – Isine – C. (Louisia) – Oblocutora – Paravulpeculella – Sphragidium – C. (Strigivulpecula) – Tomea – Volitivulpecula – Vulpeculella
Overview of species (171)
a

C. aarviki – C. aberrans – C. abrupta – C. abyssinica – C. adelina – C. adita – C. africana – C. alba – C. alborosea – C. alexi – C. amabilis – C. amatura – C. andromeda – C. angkorensis – C. arama – C. arenbergeri – C. ariadne – C. aroa – C. artemis – C. asticta – C. aurantifascia – C. aurantiorufa – C. aurifinis – C. aurorae –
b

C. barbora – C. barisana – C. barlowi – C. basialba – C. basisticta – C. bebas – C. bella – C. bellissima – C. bhatejai – C. bianca – C. bicolor – C. bigutta – C. brunnea – C. butyracea –
c

C. candida – C. capensis – C. cara – C. carmina – C. catorhoda – C. centripuncta – C. cernyi – C. charybdis – C. clama – C. coccinea – C. conclusa – C. consequenta – C. cornutissima – C. costifimbria – C. crasizona – C. croceizona – C. cruentata – C. curioi –
d

C. delicata – C. determinata – C. detrita – C. devriesi – C. distincta – C. divakara – C. dohertyi – C. dudgeoni –
e

C. effracta – C. ellipsis – C. ethiopica – C. euryxantha – C. exprimata –
f

C. fasciata – C. fasciola – C. fascioloides – C. flammeostrigata – C. flavalba – C. flaviplaga – C. fossi – C. fulvia – C. fumea –
g

C. gabica – C. gabriellae – C. garuda – C. gazella – C. gelida – C. geminipuncta – C. grandis – C. griseilinea – C. guttifera –
h

C. hamata – C. harerti – C. hecqi – C. heidrunae – C. horsfieldi – C. hsui –
i

C. ibabaoae – C. ignifera – C. impunctata – C. inconclusa – C. indonesia – C. indosinica – C. infantula – C. innocua – C. intercomma – C. interrogationis – C. inthanonica – C. inusitata
j

C. jakli – C. janinae – C. javanica – C. johani – C. juliettae –
k

C. kasperi – C. katanga – C. khasiana – C. klausruedigerbecki – C. klohsi – C. kubani – C. kucheri –
l

C. lada – C. leechi – C. liboria – C. libulae – C. lobata – C. loloana – C. longmena – C. luchoana – C. lunulata – C. lutipes – C. luzonica –
m

C. maaikeae – C. magnitrigutta – C. maiae – C. malacca – C. malayana – C. malayensis – C. margarethae – C. marshalli – C. martinii – C. melanoplagia – C. metamelas – C. meyi – C. meyricki – C. miles – C. mira – C. molleri – C. moupinensis –
n

C. natalensis – C. nemasisha – C. neopuer – C. nigrilineata – C. nigroplagata – C. nusa – C. nussi – C. nyasica –
o

C. obliquilineata – C. obscura – C. occidentalis – C. ochrata – C. owadai –
p

C. pallida – C. pallidilinea – C. paramargarethae – C. paukstadti – C. pellucida – C. peregrina – C. perornata – C. phaedra – C. phycomata – C. pitana – C. plateni – C. pratti – C. pretoriae – C. pseudoeffracta – C. pseudojavanica – C. pudens – C. puella – C. puer – C. punctistrigosa –
q

C. quadripartia – C. quentini –
r

C. rejecta – C. ridleyi – C. rhodostriata – C. rosabra – C. rubrifasciata – C. rubrifinis – C. rubristriga – C. rufeola – C. ruwenzoriana – C. rwandana –
s

C. saalmuelleri – C. sanguinea – C. saulia – C. securizonis – C. seismogrammica – C. selangorica – C. shakalesha – C. signa – C. sikkimensis – C. smithi – C. soror – C. speideli – C. squalida – C. sublutipes – C. subornata – C. succincta – C. sumatrensis – C. sumbawensis
t

C. takanoi – C. tegyra – C. tettigonioides – C. thoracica – C. titovi – C. togoana – C. torrida – C. transfasciata – C. treadawayi – C. tricolora – C. trigona – C. trigutta – C. tripuncta –
u

C. ueleana – C. ugandana – C. unipunctata – C. usambara –
v

C. venusta – C. verspectata – C. vespertata – C. v-nigrum
w

C. weerawoothi –
x/y

C. yao – C. yaseminae –
z
Name

Cyana Walker, 1854

Type species: Cyana detrita Walker, 1854

Synonyms

Doliche Walker, 1854
Isine Walker, 1854
Bizone Walker, 1854
Chionaema Herrich-Schäffer, 1855
Clerckia Aurivillius, 1882
Exotrocha Meyrick, 1886
Neobrocha Meyrick, 1886
Sphragidium Butler, 1887
Gnophrioides Heylaerts, 1891
Macronola Kirby, 1892

References

Walker, F. 1854. List of the specimens of lepidopterous insects in the collection of the British Museum. Part I. Lepidoptera Heterocera. British Museum (Natural History), London. 1-278. full article (BHL). Reference page.
Baron, T.R. & Karisch, T. 2016. A new species of Cyana Walker, 1854 from Uganda (Lepidoptera: Erebidae: Arctiinae). Metamorphosis 27: 31–33. Reference page.
Bayarsaikhan, U. & Bae, Y-S. 2016. A review of the genus Cyana Walker, 1854 (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with description of new species. Zootaxa 4114(4): 447–463. DOI: 10.11646/zootaxa.4114.4.5. Reference page.
Buchsbaum, U., Chen, D.J. & Chen, M.Y. 2018. Cyana hsui sp. n. from Lanyu Island (Taiwan) (Insecta, Lepidoptera: Erebidae, Arctiinae, Lithosiini) Contribution to the moths of Taiwan 14. Mitteilungen der Münchner Entomologischen Gesellschaft 108: 109-113. Reference page.
Bucsek, K. 2012. Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula. Malaysia Institute of Zoology SAS, Bratislava: 1–170. Reference page.
Černý, K. 2016. A Contribution to the Knowledge of the Genus: Cyana Walker in the South East Asia (Lepidoptera: Erebidae, Arctiinae, Lithosiinae). Entomofauna N.F. 37(26): 421-436. full article (pdf).Reference page.
Dubatolov, V.V., Kishida, Y. & Wang, M. 2012. New records of lichen-moths from the Nanling Mts., Guangdong, South China, with descriptions of new genera and species (Lepidoptera, Arctiidae: Lithosiinae). Tinea 22(1): 25–52. Full article (PDF).Reference page.
Holloway, J.D., 2001. The Moths of Borneo Part 7 Family Arctiidae, subfamily Lithosiinae
Inoue, H., 1988. Three new species and some synonymic notes on the Arctiidae from Japan, Taiwan and Philippines. Tyô to Ga 39 (2): 99–118. Full article: [1].
Karisch, T., 2003. Beitrag zur kenntnis der gattung Cyana Walker, 1854 in Afrika (Lepidoptera: Arctiidae). Atalanta 34 (1-2): 167–178. Abstract: [2]
Karisch, T. 2003. Zwei neue Afrikanische Cyana-arten (1) (Lepidoptera, Arctiidae). Lambillionea 103(1): 119–122. Reference page.
Karisch, T., 2003. Eine neue Cyana-Art vom Nyiragongo. Lambillionea, 103: 416–417.
Karisch, T., 2005. Eine neue Art aus der Cyana amatura (Walker, 1863) Artengruppe (Lepidoptera: Arctiidae: Lithosiinae). Atalanta 36 (1-2): 169–171.
Karisch, T., 2005. Eine neue Cyana-Art aus Ostafrika (Lepidoptera: Arctiidae: Lithosiinae). Atalanta 36 (3/4): 573–574.
Karisch, T., 2013. Taxonomic revision of the African Cyana-species (Lepidoptera: Arctiidae, Lithosiinae). Esperiana, Buchreihe zur Entomologie 18: 39–198. Abstract: [3]
Karisch, T. & U. Dall'asta, 2010. New species and subspecies of Cyana Walker, 1854 (Lepidoptera: Arctiidae: Lithosiinae) from the collection of the Royal Museum for Central Africa. Journal of Afrotropical Zoology 6: 117–127.
Kishida, Y. 1991. The genus Cyana (Lepidoptera, Arctiidae) from the Philippines. Tinea 13(8): 61-70. Reference page.
Lourens, J.H. 2009. A new species of Cyana from Northern Luzon (Philippines) belonging to the lunulata group, with an analysis of differential features and evaluation of elements for group recognition (Lepidoptera, Arctiidae, Lithosiinae). Neue Entomologische Nachrichten 30(3): 147-160. Reference page.
Lourens, J.H. 2011. Six new Philippine species of the genus Cyana Walker, 1854 and a review of the geminipuncta-group, with emphasis on endemic development lines on various islands (Lepidoptera: Arctiidae, Lithosiinae). Neue Entomologische Nachrichten 32(1-2): 69-96. Reference page.
Lourens, J.H. 2017. The Cyana insularis-group (Arctiidae, Lithosiinae, Cyana Walker 1854) with fifteen Insular and Mainland SE Asian taxa (seven nov.sp./two nov.ssp.) and seven habitus look-alikes (two nov.sp.) from likely parallel developing lineage(s) Entomofauna 38(12): 233–300. full article zobodat (pdf).Reference page.
Singh, N., Bhattacharyya, K., Volynkin, A.V. & Chandra, K. 2019. A new species of genus Cyana Walker, 1854 (Lepidoptera: Erebidae: Arctiinae). Zootaxa 4603(2): 365–376. DOI: 10.11646/zootaxa.4603.2.9 Reference page.
Singh Kirti, J. & Joshi, R. & Singh, N. 2013. A new species of the genus Cyana Walker (Lepidoptera: Arctiidae: Lithossinae) from India. Journal of Chemical, Biological and Physical Sciences 3 (2): 1301–1310. Full article: [4]
Spitsyn, V.M. 2020. Cyana pseudojavanica sp. nov. from Thailand and Cambodia (Lepidoptera: Erebidae: Arctiinae). Ecologica Montenegrina 31: 6-9. Reference page.
Volynkin, A.V. 2019f. Cyana nusa, a new species from Indonesia (Lepidoptera, Erebidae, Arctiinae). Ecologica Montenegrina 26: 102-107. Full article (PDF). Reference page.
Volynkin, A.V. 2021. A review of the Cyana phaedra (Leech, 1889) species-group, with description of a new species from China (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4970(3): 558–568. DOI: 10.11646/zootaxa.4970.3.7. Paywall. Reference page.
Volynkin, A.V. 2021a. Cyana takanoi, a new species from Tanzania (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5004(3): 493–497. DOI: 10.11646/zootaxa.5004.3.7. Paywall. Reference page.
Volynkin, A.V. & Černý, K. 2019. Three new species of Cyana Walker, 1854 from Indochina and China (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Zootaxa 4586(2): 338–350. DOI: 10.11646/zootaxa.4586.2.8. Reference page.
Volynkin, A.V. & Černý, K. 2019b. On the correct identification of Cyana bianca (Walker, 1856), with description of a new species and notes on the taxonomy of the Cyana insularis (Draudt, 1914) species-group (Lepidoptera, Erebidae, Arctiinae). Ecologica Montenegrina 21: 53-61. Reference page.
Volynkin, A.V., Černý, K. & Dubatolov, V.V. 2017. A new Cyana Walker, 1854 from Vietnam (Lepidoptera, Erebidae, Arctiinae). Zootaxa 4269(3): 438–447. DOI: 10.11646/zootaxa.4269.3.7. Reference page.
Volynkin, A.V., Černý, K. & Ivanova, M.S. 2018a. A review of the Cyana bianca (Walker, 1856) species-group with descriptions of a new species and a new subspecies from Indochina (Lepidoptera, Erebidae, Arctiinae). Zootaxa 4497(1): 082–098. DOI: 10.11646/zootaxa.4497.1.5. Reference page.
Volynkin, A.V., Huang, S.Y. & Saldaitis, A. 2021. Two new species of the genus Cyana Walker, 1854 from China (Lepidoptera: Erebidae: Arctiinae). Zootaxa 5047(5): 567–574. DOI: 10.11646/zootaxa.5047.5.6. Paywall. Reference page.
Volynkin, A.V., Černý, K., Saldaitis, A. & Ivanova, M.S. 2019. On the correct identification of ‘Cyana dohertyi (Elwes, 1890)’ from northern Indochina and China, with descriptions of two new species and a new subspecies (Lepidoptera, Erebidae, Arctiinae). Zootaxa 4658(1): 155–167. DOI: 10.11646/zootaxa.4658.1.7. Reference page.
Volynkin, A.V. & Ivanova, M.S. 2021. Cyana fossi, a new species from Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4938(5): 595–600. DOI: 10.11646/zootaxa.4938.5.7. Paywall. Reference page.
Volynkin, A.V. & László, G.M. 2019. Cyana smithi a new species from Liberia (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Ecologica Montenegrina 20: 215-221. Reference page.
Volynkin, A.V. & László, G.M. 2020a. Review of the Cyana rejecta (Walker, 1854) species-group, with descriptions of three new species from mainland Africa and a new subspecies from Madagascar (Lepidoptera, Erebidae, Arctiinae, Lithosiini)2020. Zootaxa 4890(3): 330–346. DOI: 10.11646/zootaxa.4890.3.2. Paywall. Reference page.
de Vos, R. 2018. Cyana devriesi spec. nov., a new species from southern Papua (Erebidae, Arctiinae, Lithosiini). Suara Serangga Papua 11(1): 6-9. full article (PDF). Reference page.

Cyana is a genus of moths in the family Erebidae. Species are well distributed in Africa, Madagascar, China, India, Sri Lanka, Myanmar, Sumatra, Java and Borneo. The genus was erected by Francis Walker in 1854.[1]

Description

Palpi slender and upturned. Antennae ciliated. Forewing of the male with a more or less strongly developed fringe of hair from the centre of costa on upperside and lobe on underside, which much distorts the sub-costal nervures. Vein 5 absent in male. Vein 6 usually absent in female. Veins 7 to 9 stalked. Hindwing with veins 3 and stalked. Vein 5 above angle of cell and veins 6 and 7 usually stalked.[2]

The pupae of this genus are typically attached to plant surfaces and surrounded by a characteristic trellis of hairs (see figure).
characteristic pupa
Species

The following are included in BioLib.cz:[3]

Cyana aarviki Karisch, 2013
Cyana aberrans Karisch, 2013
Cyana abyssinica Karisch, 2003
Cyana adelina (Staudinger, 1887)
Cyana adita (Moore, 1859)
Cyana africana (Holland, 1893)
Cyana alba (Moore, 1878)
Cyana alborosea (Walker, 1865)
Cyana alexi (Cerny, 1993)
Cyana amabilis (Moore, 1877)
Cyana amatura (Walker, 1863)
Cyana andromeda (Cerny, 1993)
Cyana arama (Moore, 1859)
Cyana arenbergeri Karisch, 2003
Cyana aroa (Bethune-Baker, 1904)
Cyana arorai Volynkin et al., 2020
Cyana asticta (Hampson, 1909)
Cyana atlanteia N. Singh et al., 2020
Cyana aurifinis (Cerny, 1993)
Cyana aurorae Lourens, 2011
Cyana barbora Volynkin & Saldaitis, 2020
Cyana basialba (Rothschild, 1913)
Cyana basisticta (Hampson, 1914)
Cyana bella Kishida, 1991
Cyana bellissima (Moore, 1878)[4]
Cyana bianca (Walker, 1856)[4]
Cyana bicolor (Rothschild, 1913)
Cyana bigutta (Karisch, 2005)
Cyana boetonensis (Jurriaanse & Lindemans, 1920)
Cyana britomartis N. Singh & Volynkin, 2020
Cyana brunnea (Bethune-Baker, 1904)
Cyana capensis (Hampson, 1903)
Cyana cara Kishida, 1991
Cyana carmina (Cerny, 1993)
Cyana catorhoda Hampson, 1897
Cyana cernyi Lourens, 2011
Cyana charybdis (Bethune-Baker, 1904)
Cyana chrysopeleia N. Singh et al., 2020
Cyana coccinea (Moore, 1878)
Cyana conclusa (Walker, 1862)
Cyana consequenta Lourens, 2011
Cyana cornutissima Volynkin & Lázló, 2020
Cyana costifimbria (Walker, 1862)
Cyana crasizona (Wileman & West, 1928)
Cyana croceizona (Hampson, 1914)
Cyana curioi Lourens, 2011
Cyana delicata (Walker, 1854)
Cyana determinata (Walker, 1862)
Cyana detoulgoeti Volynkin, 2020
Cyana detrita Walker, 1854 - type species
Cyana distincta (Rothschild, 1912)
Cyana dohertyi (Elwes, 1890)
Cyana dryope Volynkin & N. Singh, 2020
Cyana dudgeoni Hampson, 1895
Cyana effracta (Walker, 1854)
Cyana ellipsis Karisch & Dall'asta, 2010
Cyana ethiopica Karisch, 2013
Cyana euryxantha (Hampson, 1914)
Cyana exprimata Karisch, 2013
Cyana fasciata Karisch, 2013
Cyana flammeostrigata Karisch, 2003
Cyana formosana (Hampson, 1909)
Cyana fossi Volynkin & Ivanova, 2021
Cyana fulvia Linnaeus, 1758
Cyana gabriellae (Cerny, 1993)
Cyana gelida Walker, 1854
Cyana geminipuncta (Cerny, 1993)
Cyana gonypetes (Prout, 1919)
Cyana hamata (Walker, 1854)
Cyana hecqi (Karisch & Dall'asta, 2010)
Cyana heidrunae (Hoppe, 2004)
Cyana horsfieldi Roepke, 1946
Cyana ibabaoae Lourens, 2011
Cyana ignifera Karisch, 2013
Cyana inconclusa (Walker, 1862)
Cyana innocua Karisch, 2013
Cyana interrogationis (Poujade, 1886)
Cyana inusitata (Bethune-Baker, 1910)
Cyana janinae Lourens, 2009
Cyana katanga Karisch & Dall’Asta, 2010
Cyana khasiana Hampson, 1897
Cyana klausruedigerbecki (Karisch, 2005)
Cyana klohsi Karisch, 2003
Cyana lada Volynkin, Černý & Saldaitis, 2019
Cyana leechi Volynkin, 2021
Cyana libulae (Cerny, 1993)
Cyana linatula (Swinhoe, 1891)
Cyana lobata Karisch, 2013
Cyana loloana (Strand, 1912)
Cyana luchoana (Karisch, 2003)
Cyana lunulata Semper, 1899
Cyana lutipes (Hampson, 1900)
Cyana luzonica (Wileman & South)
Cyana magnitrigutta Karisch, 2013
Cyana malayensis (Hampson, 1914)
Cyana margarethae (Kiriakoff, 1958)
Cyana marshalli (Hampson, 1900)
Cyana metamelas (Hampson, 1914)
Cyana meyi Karisch, 2013
Cyana meyricki (Rothschild, 1901)
Cyana miles Butler, 1887
Cyana molleri (Elwes, 1890)
Cyana moupinensis (Leech, 1899)
Cyana natalensis Karisch, 2013
Cyana nemasisha (Roesler, 1990)
Cyana nigroplagata (Bethune-Baker, 1910)
Cyana nussi Karisch, 2013
Cyana nyasica (Hampson, 1918)
Cyana obliquilineata (Hampson, 1900)
Cyana obscura (Hampson, 1900)
Cyana occidentalis Volynkin & Lázló, 2020
Cyana ochrata Karisch, 2013
Cyana owadai (Kishida, 1991)
Cyana pallida Karisch, 2013
Cyana pallidilinea (Karisch, 2003)
Cyana paramargarethae Karisch & Dall'asta, 2010
Cyana pauliani (de Toulgoët, 1954)
Cyana peregrina (Walker, 1854)
Cyana perornata (Walker, 1854)
Cyana phycomata (Wileman & West, 1928)
Cyana pratti (Elwes, 1890)
Cyana pretoriae (Distant, 1897)
Cyana pudens (Walker, 1862)
Cyana puella (Drury, 1773)
Cyana punctistrigosa (Rothschild, 1913)
Cyana quadrinotata (Walker, 1869)
Cyana quentini Karisch, 2003
Cyana rawlinsi Karisch, 2013
Cyana rejecta (Walker, 1854)
Cyana rhodostriata (Hampson, 1914)
Cyana rosabra (Wileman, 1925)
Cyana rubrifasciata (Druce, 1883)
Cyana rubrifinis Lourens, 2011
Cyana rubristriga (Holland, 1893)
Cyana rubritermina (Bethune-Baker, 1911)
Cyana rubriterminalis (Strand, 1912 – incertae sedis)
Cyana rudloffi Volynkin & N. Singh, 2020
Cyana rufeola Karisch & Dall’Asta, 2010
Cyana rufifrons (Rothschild, 1912)
Cyana ruwenzoriana (Karisch, 2003)
Cyana rwandana Karisch, 2013
Cyana saalmuelleri (Butler, 1882)
Cyana saulia (Swinhoe, 1901)
Cyana schaeferi (Gaede, 1924 – incertae sedis)
Cyana selangorica (Hampson, 1903)
Cyana signa (Walker, 1854)
Cyana soror (Cerny, 1993)
Cyana speideli Karisch, 2013
Cyana squalida Karisch, 2013
Cyana subalba (Wileman, 1910)
Cyana sublutipes (Kishida, 1991)
Cyana subornata (Walker, 1854)
Cyana suessmuthi Karisch, 2013
Cyana tettigonioides (Heylaerts, 1892)
Cyana thoracica (Rothschild & Jordan, 1901)
Cyana titovi Volynkin, Černý & Ivanova, 2019
Cyana togoana (Strand, 1912)
Cyana torrida (Holland, 1893)
Cyana transfasciata (Rothschild, 1912)
Cyana treadawayi (Cerny, 1993)
Cyana trigona (Rothschild, 1903)
Cyana trigutta Walker, 1854
Cyana tripuncta (de Toulgoet, 1980)
Cyana tripunctata Rothschild, 1936
Cyana ueleana (Karisch, 2003)
Cyana unipunctata (Elwes, 1890)
Cyana usambara Karisch, 2013
Cyana v-nigrum (Cerny, 1993)
Cyana venusta Karisch, 2013
Cyana vespertata (Cerny, 1993)
Cyana watsoni Hampson, 1897
Cyana yao Volynkin & Lázló, 2020

References

Savela, Markku. "Cyana Walker, 1854". Lepidoptera and Some Other Life Forms. Retrieved February 4, 2018.
Hampson, G. F. (1894). The Fauna of British India, Including Ceylon and Burma: Moths Volume II. Taylor and Francis – via Biodiversity Heritage Library.
BioLib.cz: genus Cyana Walker, 1854 (retrieved 4 September 2021)

Singh N, Volynkin AV, Kirti JS, Datta HS, Ivanova, MS (2020) A review of the genus Cyana Walker, 1854 from India, with descriptions of five new species and three new subspecies (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4738 (1): 1-93. DOI

Dubatolov, Vladimir V.; Kishida, Yasunori; Wang, Min (May 2012). "New records of lichen-moths from the Nanling Mts., Guangdong, South China, with descriptions of new genera and species (Lepidoptera, Arctiidae: Lithosiinae)". Tinea. 22 (1): 25–52. S2CID 86037229.
Holloway, J. D. 2001: The Moths of Borneo Part 7 Family Arctiidae, subfamily Lithosiinae
Inoue, H., 1988: Three new species and some synonymic notes on the Arctiidae from Japan, Taiwan and Philippines. Tyô to Ga 39 (2): 99-118. Full article: [1].
Karisch, T. 2003: Beitrag zur kenntnis der gattung Cyana Walker, 1854 in Afrika (Lepidoptera: Arctiidae). Atalanta 34 (1-2): 167–178. Abstract: [2]
Karisch, T. 2005: Eine neue Art aus der Cyana amatura (Walker, 1863) Artengruppe (Lepidoptera: Arctiidae: Lithosiinae). Atalanta 36 (1-2): 169–171.
Karisch, T. 2005: Eine neue Cyana-Art aus Ostafrika (Lepidoptera: Arctiidae: Lithosiinae). Atalanta 36 (3/4): 573–574.
Karisch, T. & Dall'Asta, U., 2010: New species and subspecies of Cyana Walker, 1854 (Lepidoptera: Arctiidae: Lithosiinae) from the collection of the Royal Museum for Central Africa. Journal of Afrotropical Zoology 6: 117–127.
Lourens, J. H. 2009: A new species of Cyana from Northern Luzon (Philippines) belonging to the lunulata group, with an analysis of differential features and evaluation of elements for group recognition (Lepidoptera: Arctiidae: Lithosiinae). Nachrichten des Entomologischen Vereins Apollo, N.F. 30 (3): 147–160.
Lourens, J. H. 2011: Six new Philippine species of the genus Cyana Walker, 1854 and a review of the geminipuncta-group, with emphasis on endemic development lines on various islands (Lepidoptera: Arctiidae, Lithosiinae). Nachrichten des Entomologischen Vereins Apollo, 32 (1-2): 69–96.

External links
Pitkin, Brian & Jenkins, Paul. "Search results Family: Arctiidae". Butterflies and Moths of the World. Natural History Museum, London.
De Prins, J. & De Prins, W. (2018). "Cyana Walker, 1854". Afromoths. Retrieved January 14, 2019.