Copris

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Scarabaeinae
Tribus: Coprini
Genus: Copris
Subgenera (3): C. (Copris) – C. (Paracopris) – C. (Sinocopris)

Invalid Subgenera: C. (Microcopris)

Species (248): C. acutidens – C. afgoi – C. agnus – C. algol – C. amabilis – C. amyntor – C. anceus – C. andrewesi – C. angusticornis – C. angustus – C. anomiopseoides – C. antares – C. arcturus – C. ares – C. arimotoi – C. arizonensis – C. armatus – C. armeniacus – C. armiger – C. arrowi – C. aspericollis – C. atropolitus – C. barclayi – C. basilewskyi – C. basipunctatus – C. bellator – C. bengalensis – C. bidens – C. bootes – C. boucardi – C. bovinus – C. brachypterus – C. brittoni – C. caelatus – C. caliginosus – C. cambeforti – C. cambodiensis – C. camerunus – C. capensis – C. cariniceps – C. carinicus – C. carmelita – C. cassius – C. celebensis – C. cheni – C. chimalapensis – C. colmanti – C. complexus – C. compressipennis – C. confucius – C. coriarius – C. cornifrons – C. corniger – C. corpulentus – C. costaricensis – C. crassus – C. cribratus – C. cribricollis – C. curlettii – C. davidis – C. davisi – C. davisoni – C. decellei – C. delicatus – C. denticulatus – C. diversus – C. draco – C. dracunculus – C. dudleyi – C. eburneus – C. elpheneroides – C. elphenor – C. erratus – C. evanidus – C. excisus – C. fallaciosus – C. fallax – C. felschei – C. fidius – C. frankenbergerianus – C. freyi – C. fricator – C. fukiensis – C. furciceps – C. garambae – C. gazellarum – C. gibbulus – C. gilleti – C. gladiator – C. gopheri – C. gracilis – C. graueri – C. halffteri – C. harrisi – C. hartli – C. hatayamai – C. hispanus – C. howdeni – C. humilis – C. hybridus – C. igualensis – C. illotus – C. imitans – C. inaequabilis – C. incertus – C. inemarginatus – C. inhalatus – C. insidiosus – C. integer – C. interioris – C. iris – C. jacchoides – C. jacchus – C. jahi – C. johkii – C. juanmorai – C. jucundus – C. kachinensis – C. kasagii – C. kasanka – C. katangae – C. keralensis – C. kiuchii – C. klapperichi – C. klugi – C. kusakabei – C. laeviceps – C. laevigatus – C. laiiformis – C. laioides – C. laius – C. lannathai – C. lecontei – C. lugubris – C. lunaris – C. macacus – C. macclevei – C. macer – C. maesi – C. magicus – C. manni – C. marcus – C. martinae – C. martinezi – C. matthewsi – C. medogensis – C. megaceratoides – C. megasoma – C. mesacanthus – C. mexicanus – C. minutus – C. misellus – C. moechus – C. moffartsi – C. mongkhoni – C. montivagus – C. morphaeus – C. mourgliai – C. neglectus – C. nepos – C. nevinsoni – C. novaki – C. nubilosus – C. numa – C. obenbergeri – C. obesus – C. ochus – C. orion – C. orphanus – C. parapecuarius – C. pauliani – C. pecuarius – C. pedarioides – C. phungae – C. phylax – C. poggii – C. potanini – C. prevosti – C. pseudobootes – C. pseudochus – C. pseudomoffartsi – C. pseudosinicus – C. pueli – C. punctatus – C. puncticollis – C. punctipennis – C. punctulatus – C. punjabensis – C. quasilaevigatus – C. ramosiceps – C. rebouchei – C. remotus – C. renwarti – C. repertus – C. ribbei – C. riekoae – C. ritsemae – C. rugosus – C. ruricola – C. sabinus – C. sacontala – C. sallei – C. sarpedon – C. saundersi – C. schoolmeestersi – C. scorpio – C. serius – C. sexdentatus – C. siamensis – C. siangensis – C. sierrensis – C. signatus – C. simonettai – C. simulator – C. singularis – C. sinicus – C. sinon – C. sodalis – C. sorex – C. sphaeropterus – C. spinator – C. subdolus – C. subpunctatus – C. subsidens – C. sumatrensis – C. surdus – C. szechouanicus – C. tetraodon – C. tridens – C. tridentatus – C. tripartitus – C. truncatus – C. tsukamotoi – C. typhoeus – C. uenoi – C. umbilicatus – C. urus – C. usambaricus – C. vankhaii – C. victorini – C. vietnamicus – C. vilhenai – C. vrydaghi – C. warneri – C. wiesei – C. yangi – C. youngai – C. zhangi
Name

Copris Geoffroy, 1762 BHL

Type species: Scarabaeus lunaris Linnaeus, 1758 by subsequent designation by Latreille 1810: 428 BHL.

Synonym:

Copris obliquatus Voet, 1806 = Canthon quadriguttatus (Olivier, 1789)

References
Primary references

Geoffroy, E.L. 1762. Histoire abrégée des insectes qui se trouvent aux environs de Paris; dans laquelle ces animaux sont rangés suivant un ordre méthodique. Tome seconde. Imprimerie de J. Durand, Paris, 690 pp., 22 pls. BHL [original description]

Additional references

Deschodt, C.M., Davis. L.V. & Scholtz, C.H. 2015. A new synonymy in the fidius group of Copris Müller 1764 (Coleoptera: Scarabaeidae: Scarabaeinae) and a new species from the highland grasslands of South Africa. Zootaxa 3949(3): 431–438. DOI: 10.11646/zootaxa.3949.3.8 Preview (PDF). Reference page.
Falahee, S.L.; Angus, R.B. 2010: Chromosomal separation of difficult species of Copris Geoffroy, 1762 and Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae), with discussion of O. massai Baraud as a British Pleistocene fossil. ZooKeys, 34: 17–32. DOI: 10.3897/zookeys.34.256
Latreille, P.A. 1810. Considérations générales sur l'ordre naturel des animaux composant les classes des crustacés, des arachnides, et des insectes : avec un tableau méthodique de leurs genres, disposés en familles. Paris: Schoell, 444 pp. BHL Reference page.
Marchisio, R.; Zunino, M. 2012: Il genere Copris Müller. Tassonomia, filogenesi e note di zoogeografia. World Biodiversity Association monographs, (2) ISBN 978-88-906379-1-9 preview only seen
McCleve, S.; Kohlmann, B. 2005: New species and new records of Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and the United States. Zootaxa, 1096: 17–28. Abstract & excerpt PDF
Ochi, T.; Kon, M. 2010: Two new taxa of Copris (Coleoptera: Scarabaeidae) from Asia. Entomological review of Japan, 65(1): 127–130. [Publication date: 'June 30, 2010', sourced from first page of article]
Ochi, T. & Kon, M. 2021. Notes on the Coprophagous Scarab Beetles (Coleoptera, Scarabaeidae) from Southeast Asia. XXXIV Four new species of Copris Müller from Myanmar, Laos and Cambodia. Giornale Italiano di Entomologia 16(66): 149-158. Reference page.
Ochi, T., Kon, M. & Bai, M. 2009. Three new species of Copris (Coleoptera: Scarabaeidae) from China, with description of a new subgenus. Entomological review of Japan 64(2): 207-216. Reference page.
Ochi, T.; Kon, M.; Kawahara, M. 2011: Four new taxa of Scarabaeoidea (Coleoptera) from Southeast Asia. Special publication of the Japanese Society of Scarabaeoidology, (1): 153–162. [Publication date: 'April 25, 2011', sourced from first page of article]
Ziani, S. 2017. Morphological revision of the western Palaearctic species of the genus Copris Geoffroy, 1762 with three foretibial external teeth (Coleoptera: Scarabaeoidea: Scarabaeidae). Insecta mundi 0528: 1–26. Full text (PDF) Reference page. [Key to Western Palaearctic Copris: p. 14]

Links

Schoolmeesters, P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on March 27, 2017]

Species

References

Scholtz, Clarke H.; Davis, Adrian L. V.; Kryger, Ute (2009). Evolutionary biology and conservation of dung beetles. Sofia-Moscow: Pensoft Pub. ISBN 978-954-642-517-1.

Copris acutidens Motschulsky, 1860
Copris afgoi Cambefort, 1992
Copris agnus Sharp, 1875
Copris algol Nguyen-phung, 1989
Copris amabilis Kolbe, 1914
Copris amyntor Klug, 1855
Copris anceus (Olivier, 1789)
Copris andrewesi Waterhouse, 1891
Copris angusticornis Arrow, 1933
Copris angustus Nguyen-phung & Cambefort, 1986
Copris anomiopseoides Boucomont, 1924
Copris antares Ferreira, 1958
Copris arcturus Gillet, 1907
Copris ares Zelenka, 1993
Copris arimotoi Ochi & Araya, 1992
Copris arizonensis Schaeffer, 1906
Copris armatus Harold, 1869
Copris armeniacus Faldermann, 1835
Copris armiger Gillet, 1910
Copris arrowi Felsche, 1910
Copris aspericollis Gillet, 1910
Copris atropolitus Gillet, 1910
Copris bacchus Thunberg, 1787
Copris barclayi Ochi, Kon & Bai, 2009
Copris basilewskyi Ferreira, 1962
Copris basipunctatus Balthasar, 1942
Copris bellator Chevrolat, 1844
Copris bengalensis Gillet, 1911
Copris bidens Kolbe, 1893
Copris binodis
Copris bootes Klug, 1855
Copris boucardi Harold, 1869
Copris bovinus Gillet, 1908
Copris brachypterus Nomura, 1964
Copris brittoni Ferreira, 1962
Copris caelatus (Fabricius, 1794)
Copris caliginosus Kohlmann, Cano & Delgado, 2003
Copris cambeforti Nguyen-phung, 1988
Copris cambodiensis Ochi, Kon & Kawahara, 2008
Copris camerunus Felsche, 1904
Copris caobangensis Bui, Dumack & Bonkowski, 2018 [3]
Copris capensis Waterhouse, 1891
Copris cariniceps Felsche, 1910
Copris carinicus Gillet, 1910
Copris carmelita Fabricius, 1801
Copris cassius Péringuey, 1901
Copris celebensis Lansberge, 1886
Copris cheni Ochi, Kon & Bai, 2007
Copris chimalapensis Mora-Aguilar & Delgado, 2015
Copris ciliatus
Copris colmanti Gillet, 1908
Copris complexus Nguyen-phung, 1989
Copris compressipennis Gillet, 1910
Copris confucius Harold, 1877
Copris coriarius Gillet, 1907
Copris cornifrons Boheman, 1860
Copris corniger Sahlberg, 1823
Copris corpulentus Gillet, 1910
Copris costaricensis Gahan, 1894
Copris costariensis Matthews
Copris crassus Deschodt & Davis, 2015
Copris cribratus Gillet, 1927
Copris cribricollis Gillet, 1910
Copris curlettii Moretto, 2011
Copris davidis (Deyrolle, 1878)
Copris davisi Nguyen-phung & Cambefort, 1986
Copris davisoni Waterhouse, 1891
Copris decellei Cambefort, 1992
Copris delicatus Arrow, 1931
Copris denticulatus Nguyen-phung, 1988
Copris diversus Waterhouse, 1891
Copris draco Arrow, 1906
Copris dracunculus Fereira, 1959
Copris druidum Heer, 1862
Copris dudleyi Cambefort, 1992
Copris eburneus Cambefort, 1992
Copris elpheneroides Felsche, 1910
Copris elphenor Klug, 1855
Copris erratus Lansberge, 1886
Copris evanidus Klug, 1855
Copris excisus Waterhouse, 1891
Copris fallaciosus Gillet, 1907
Copris fallax Felsche, 1910
Copris felschei Reitter, 1892
Copris fidius (Olivier, 1789)
Copris frankenbergerianus Balthasar, 1958
Copris freyi Ferreira, 1961
Copris fricator (Fabricius, 1787)
Copris fukiensis Balthasar, 1952
Copris furciceps Felsche, 1910
Copris garambae Cambefort, 1992
Copris gazellarum Gillet, 1918
Copris gibbulus Lansberge, 1886
Copris gilleti Kolbe, 1907
Copris gladiator Arrow, 1933
Copris gopheri Hubbard, 1894
Copris gracilis Waterhouse, 1891
Copris graueri Kolbe, 1914
Copris halffteri Matthews, 1959
Copris harrisi Waterhouse, 1891
Copris hartli Frey, 1975
Copris hatayamai Ochi & Araya, 1992
Copris hispanus (Linnaeus, 1764)
Copris howdeni Matthews & Halffter, 1959
Copris humilis Gillet, 1908
Copris hybridus Gillet, 1910
Copris igualensis Warner, 1990
Copris illotus Balthasar, 1942
Copris imitans Felsche, 1910
Copris inaequabilis Zhang, 1997
Copris incertus Say, 1835
Copris inemarginatus Blatchley, 1918
Copris inhalatus Quedenfeldt, 1884
Copris insidiosus Péringuey, 1901
Copris integer Reiche, 1847
Copris interioris Kolbe, 1897
Copris iris Sharp, 1875
Copris jacchoides Nguyen-phung & Cambefort, 1987
Copris jacchus (Fabricius, 1775)
Copris jahi Nguyen-phung & Cambefort, 1986
Copris johkii Ochi & Kon, 2014
Copris juanmorai Mora-Aguilar & Delgado, 2015
Copris jucundus Gillet, 1932
Copris kachinensis Ochi, Kon & Kawahara, 2008
Copris kartlinus Kabakov, 1988
Copris kasagii Ochi & Kon, 1996
Copris kasanka Josso & Prévost, 2016
Copris katangae Cambefort & Nguyen-phung, 1996
Copris keralensis Gill, 1986
Copris kiuchii Masumoto, 1989
Copris klapperichi Balthasar, 1942
Copris klugi Harold, 1869
Copris kusakabei Ochi, Kon & Kawahara, 2005
Copris laeviceps Harold, 1862
Copris laevigatus Gillet, 1927
Copris laiiformis Nguyen-phung & Cambefort, 1987
Copris laioides Boucomont, 1932
Copris laius Harold, 1868
Copris lannathai Hanboonsong & Masumoto & Ochi, 2003
Copris leakeyorum Paulian, 1976
Copris lecontei Matthews, 1962
Copris leruei Josso & Prévost, 2017
Copris lugubris Boheman, 1858
Copris lunaris [fr] (Linnaeus, 1758)
Copris macacus Lansberge, 1886
Copris macclevei Warner, 1990
Copris macer Péringuey, 1901
Copris maesi Ratcliffe, 1998
Copris magicus Harold, 1881
Copris manni Ochi, Kon & Bai, 2009
Copris marcus Gillet, 1921
Copris martinae Nguyen-phung, 1988
Copris martinezi Matthews and Halffter, 1968
Copris matthewsi Delgado & Kohlmann, 2001
Copris medogensis Zhang, 1988
Copris megaceratoides Waterhouse, 1891
Copris megasoma Matthews & Halffter, 1959
Copris mesacanthus Harold, 1878
Copris mexicanus Matthews & Halffter, 1959
Copris minutus (Drury, 1773)
Copris misellus Péringuey, 1901
Copris moechus LeConte, 1854
Copris moffartsi Gillet, 1907
Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003
Copris montivagus Gillet, 1908
Copris morphaeus Gillet, 1932
Copris mourgliai Zelenka, 1993
Copris neglectus Moxey, 1963
Copris neotridens Génier & Prévost, 2017
Copris nepos Gillet, 1908
Copris nevinsoni Waterhouse, 1891
Copris novaki Zelenka, 1992
Copris nubilosus Kohlmann & Cano & Delgado, 2003
Copris numa Lansberge, 1886
Copris obenbergeri Balthasar, 1933
Copris obesus Boheman, 1857
Copris ochus (Motschulsky, 1860)
Copris orion Klug, 1835
Copris orphanus Guérin-Méneville, 1847
Copris parapecuarius Ochi, Kon & Kawahara, 2008
Copris pauliani Nguyen-phung & Cambefort, 1986
Copris pecuarius Lewis, 1884
Copris pedarioides Lansberge, 1886
Copris phungae Cambefort, 1992
Copris phylax Gillet, 1908
Copris poggii Ochi & Kon, 2005
Copris potanini Semenov, 1891
Copris prevosti Josso, 2011
Copris pristinus Pierce, 1946
Copris pseudobootes Ferreira, 1962
Copris pseudochus Ochi & Kon, 2004
Copris pseudomoffartsi Moretto, 2011
Copris pseudosinicus Balthasar, 1958
Copris pueli Mollandin De Boissy, 1905
Copris punctatus Gillet, 1910
Copris puncticollis Boheman, 1857
Copris punctipennis Boucomont, 1914
Copris punctulatus Wiedemann, 1823
Copris punjabensis Gillet, 1921
Copris quasilaevigatus Ochi, Kon & Bai, 2007
Copris ramosiceps Gillet, 1921
Copris rebouchei Harold, 1869
Copris remotus Leconte, 1866
Copris renwarti Nguyen-phung & Cambefort, 1986
Copris repertus Walker, 1858
Copris ribbei Lansberge, 1886
Copris riekoae Ochi & Kon, 2003
Copris ritsemae Harold, 1875
Copris rugosus Gillet, 1908
Copris ruricola Balthasar, 1933
Copris sabinus Gillet, 1910
Copris sacontala Redtenbacher, 1848
Copris sallei Harold, 1869
Copris sarpedon Harold, 1868
Copris saundersi Harold, 1869
Copris schoolmeestersi Ochi & Kon, 2010
Copris scorpio Zelenka, 1993
Copris serius Nguyen-phung, 1987
Copris sexdentatus Thunberg, 1818
Copris siamensis Gillet, 1910
Copris siangensis Biswas, 1980
Copris sierrensis Matthews, 1961
Copris signatus Walker, 1858
Copris simonettai Cambefort, 1992
Copris simulator Nguyen-phung & Cambefort, 1986
Copris singularis Nguyen-phung & Cambefort, 1986
Copris sinicus Hope, 1842
Copris sinon (Olivier, 1789)
Copris sodalis Walker, 1858
Copris sonensis Bui, Dumack & Bonkowski, 2018 [3]
Copris sorex Balthasar, 1942
Copris sphaeropterus Harold, 1877
Copris spinator Harold, 1881
Copris subdolus Balthasar, 1958
Copris subpunctatus Gillet, 1910
Copris subsidens Péringuey, 1901
Copris subterraneus Heer, 1862
Copris sumatrensis Gillet, 1921
Copris surdus Arrow, 1931
Copris szechouanicus Balthasar, 1958
Copris tetraodon Gillet, 1910
Copris tridentatus Solis & Kohlmann, 2003
Copris tripartitus Waterhouse, 1875
Copris truncatus Felsche, 1901
Copris tsukamotoi Ochi, Kon & Kawahara, 2008
Copris typhoeus Gerstaecker, 1884
Copris uenoi Hanboonsong & Masumoto & Ochi, 2003
Copris umbilicatus Abeille De Perrin, 1901
Copris urus Boheman, 1857
Copris usambaricus Gillet, 1908
Copris vankhaii Nguyen-phung, 1988
Copris victorini Boheman, 1857
Copris vietnamicus Kabakov, 1994
Copris vilhenai Ferreira, 1962
Copris vrydaghi Ferreira, 1962
Copris warneri Mccleve & Kohlmann, 2005
Copris wiesei Kolbe, 1914
Copris yangi Ochi, Kon & Bai, 2007
Copris youngai Balthasar, 1967
Copris zhangi Ochi, Kon & Bai, 2009
Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Scarabaeinae
Tribus: Coprini
Genus: Copris
Subgenera (3): C. (Copris) – C. (Paracopris) – C. (Sinocopris)

Invalid Subgenera: C. (Microcopris)

Species (248): C. acutidens – C. afgoi – C. agnus – C. algol – C. amabilis – C. amyntor – C. anceus – C. andrewesi – C. angusticornis – C. angustus – C. anomiopseoides – C. antares – C. arcturus – C. ares – C. arimotoi – C. arizonensis – C. armatus – C. armeniacus – C. armiger – C. arrowi – C. aspericollis – C. atropolitus – C. barclayi – C. basilewskyi – C. basipunctatus – C. bellator – C. bengalensis – C. bidens – C. bootes – C. boucardi – C. bovinus – C. brachypterus – C. brittoni – C. caelatus – C. caliginosus – C. cambeforti – C. cambodiensis – C. camerunus – C. capensis – C. cariniceps – C. carinicus – C. carmelita – C. cassius – C. celebensis – C. cheni – C. chimalapensis – C. colmanti – C. complexus – C. compressipennis – C. confucius – C. coriarius – C. cornifrons – C. corniger – C. corpulentus – C. costaricensis – C. crassus – C. cribratus – C. cribricollis – C. curlettii – C. davidis – C. davisi – C. davisoni – C. decellei – C. delicatus – C. denticulatus – C. diversus – C. draco – C. dracunculus – C. dudleyi – C. eburneus – C. elpheneroides – C. elphenor – C. erratus – C. evanidus – C. excisus – C. fallaciosus – C. fallax – C. felschei – C. fidius – C. frankenbergerianus – C. freyi – C. fricator – C. fukiensis – C. furciceps – C. garambae – C. gazellarum – C. gibbulus – C. gilleti – C. gladiator – C. gopheri – C. gracilis – C. graueri – C. halffteri – C. harrisi – C. hartli – C. hatayamai – C. hispanus – C. howdeni – C. humilis – C. hybridus – C. igualensis – C. illotus – C. imitans – C. inaequabilis – C. incertus – C. inemarginatus – C. inhalatus – C. insidiosus – C. integer – C. interioris – C. iris – C. jacchoides – C. jacchus – C. jahi – C. johkii – C. juanmorai – C. jucundus – C. kachinensis – C. kasagii – C. kasanka – C. katangae – C. keralensis – C. kiuchii – C. klapperichi – C. klugi – C. kusakabei – C. laeviceps – C. laevigatus – C. laiiformis – C. laioides – C. laius – C. lannathai – C. lecontei – C. lugubris – C. lunaris – C. macacus – C. macclevei – C. macer – C. maesi – C. magicus – C. manni – C. marcus – C. martinae – C. martinezi – C. matthewsi – C. medogensis – C. megaceratoides – C. megasoma – C. mesacanthus – C. mexicanus – C. minutus – C. misellus – C. moechus – C. moffartsi – C. mongkhoni – C. montivagus – C. morphaeus – C. mourgliai – C. neglectus – C. nepos – C. nevinsoni – C. novaki – C. nubilosus – C. numa – C. obenbergeri – C. obesus – C. ochus – C. orion – C. orphanus – C. parapecuarius – C. pauliani – C. pecuarius – C. pedarioides – C. phungae – C. phylax – C. poggii – C. potanini – C. prevosti – C. pseudobootes – C. pseudochus – C. pseudomoffartsi – C. pseudosinicus – C. pueli – C. punctatus – C. puncticollis – C. punctipennis – C. punctulatus – C. punjabensis – C. quasilaevigatus – C. ramosiceps – C. rebouchei – C. remotus – C. renwarti – C. repertus – C. ribbei – C. riekoae – C. ritsemae – C. rugosus – C. ruricola – C. sabinus – C. sacontala – C. sallei – C. sarpedon – C. saundersi – C. schoolmeestersi – C. scorpio – C. serius – C. sexdentatus – C. siamensis – C. siangensis – C. sierrensis – C. signatus – C. simonettai – C. simulator – C. singularis – C. sinicus – C. sinon – C. sodalis – C. sorex – C. sphaeropterus – C. spinator – C. subdolus – C. subpunctatus – C. subsidens – C. sumatrensis – C. surdus – C. szechouanicus – C. tetraodon – C. tridens – C. tridentatus – C. tripartitus – C. truncatus – C. tsukamotoi – C. typhoeus – C. uenoi – C. umbilicatus – C. urus – C. usambaricus – C. vankhaii – C. victorini – C. vietnamicus – C. vilhenai – C. vrydaghi – C. warneri – C. wiesei – C. yangi – C. youngai – C. zhangi
Name

Copris Geoffroy, 1762 BHL

Type species: Scarabaeus lunaris Linnaeus, 1758 by subsequent designation by Latreille 1810: 428 BHL.

Synonym:

Copris obliquatus Voet, 1806 = Canthon quadriguttatus (Olivier, 1789)

References
Primary references

Geoffroy, E.L. 1762. Histoire abrégée des insectes qui se trouvent aux environs de Paris; dans laquelle ces animaux sont rangés suivant un ordre méthodique. Tome seconde. Imprimerie de J. Durand, Paris, 690 pp., 22 pls. BHL [original description]

Additional references

Deschodt, C.M., Davis. L.V. & Scholtz, C.H. 2015. A new synonymy in the fidius group of Copris Müller 1764 (Coleoptera: Scarabaeidae: Scarabaeinae) and a new species from the highland grasslands of South Africa. Zootaxa 3949(3): 431–438. DOI: 10.11646/zootaxa.3949.3.8 Preview (PDF). Reference page.
Falahee, S.L.; Angus, R.B. 2010: Chromosomal separation of difficult species of Copris Geoffroy, 1762 and Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae), with discussion of O. massai Baraud as a British Pleistocene fossil. ZooKeys, 34: 17–32. DOI: 10.3897/zookeys.34.256
Latreille, P.A. 1810. Considérations générales sur l'ordre naturel des animaux composant les classes des crustacés, des arachnides, et des insectes : avec un tableau méthodique de leurs genres, disposés en familles. Paris: Schoell, 444 pp. BHL Reference page.
Marchisio, R.; Zunino, M. 2012: Il genere Copris Müller. Tassonomia, filogenesi e note di zoogeografia. World Biodiversity Association monographs, (2) ISBN 978-88-906379-1-9 preview only seen
McCleve, S.; Kohlmann, B. 2005: New species and new records of Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and the United States. Zootaxa, 1096: 17–28. Abstract & excerpt PDF
Ochi, T.; Kon, M. 2010: Two new taxa of Copris (Coleoptera: Scarabaeidae) from Asia. Entomological review of Japan, 65(1): 127–130. [Publication date: 'June 30, 2010', sourced from first page of article]
Ochi, T. & Kon, M. 2021. Notes on the Coprophagous Scarab Beetles (Coleoptera, Scarabaeidae) from Southeast Asia. XXXIV Four new species of Copris Müller from Myanmar, Laos and Cambodia. Giornale Italiano di Entomologia 16(66): 149-158. Reference page.
Ochi, T., Kon, M. & Bai, M. 2009. Three new species of Copris (Coleoptera: Scarabaeidae) from China, with description of a new subgenus. Entomological review of Japan 64(2): 207-216. Reference page.
Ochi, T.; Kon, M.; Kawahara, M. 2011: Four new taxa of Scarabaeoidea (Coleoptera) from Southeast Asia. Special publication of the Japanese Society of Scarabaeoidology, (1): 153–162. [Publication date: 'April 25, 2011', sourced from first page of article]
Ziani, S. 2017. Morphological revision of the western Palaearctic species of the genus Copris Geoffroy, 1762 with three foretibial external teeth (Coleoptera: Scarabaeoidea: Scarabaeidae). Insecta mundi 0528: 1–26. Full text (PDF) Reference page. [Key to Western Palaearctic Copris: p. 14]

Links

Schoolmeesters, P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on March 27, 2017]

Copris is a genus of dung beetles in the tribe Coprini (subfamily Scarabaeinae) of the scarab family.[1] It comprises more than 250 tunnelling species and has an almost worldwide distribution.[1]
Contents

1 Species
2 Gallery
3 References
4 External links

Species
Main article: List of Copris species

References

Scholtz, Clarke H.; Davis, Adrian L. V.; Kryger, Ute (2009). Evolutionary biology and conservation of dung beetles. Sofia-Moscow: Pensoft Pub. ISBN 978-954-642-517-1.

Copris acutidens Motschulsky, 1860
Copris afgoi Cambefort, 1992
Copris agnus Sharp, 1875
Copris algol Nguyen-phung, 1989
Copris amabilis Kolbe, 1914
Copris amyntor Klug, 1855
Copris anceus (Olivier, 1789)
Copris andrewesi Waterhouse, 1891
Copris angusticornis Arrow, 1933
Copris angustus Nguyen-phung & Cambefort, 1986
Copris anomiopseoides Boucomont, 1924
Copris antares Ferreira, 1958
Copris arcturus Gillet, 1907
Copris ares Zelenka, 1993
Copris arimotoi Ochi & Araya, 1992
Copris arizonensis Schaeffer, 1906
Copris armatus Harold, 1869
Copris armeniacus Faldermann, 1835
Copris armiger Gillet, 1910
Copris arrowi Felsche, 1910
Copris aspericollis Gillet, 1910
Copris atropolitus Gillet, 1910
Copris bacchus Thunberg, 1787
Copris barclayi Ochi, Kon & Bai, 2009
Copris basilewskyi Ferreira, 1962
Copris basipunctatus Balthasar, 1942
Copris bellator Chevrolat, 1844
Copris bengalensis Gillet, 1911
Copris bidens Kolbe, 1893
Copris binodis
Copris bootes Klug, 1855
Copris boucardi Harold, 1869
Copris bovinus Gillet, 1908
Copris brachypterus Nomura, 1964
Copris brittoni Ferreira, 1962
Copris caelatus (Fabricius, 1794)
Copris caliginosus Kohlmann, Cano & Delgado, 2003
Copris cambeforti Nguyen-phung, 1988
Copris cambodiensis Ochi, Kon & Kawahara, 2008
Copris camerunus Felsche, 1904
Copris caobangensis Bui, Dumack & Bonkowski, 2018 [3]
Copris capensis Waterhouse, 1891
Copris cariniceps Felsche, 1910
Copris carinicus Gillet, 1910
Copris carmelita Fabricius, 1801
Copris cassius Péringuey, 1901
Copris celebensis Lansberge, 1886
Copris cheni Ochi, Kon & Bai, 2007
Copris chimalapensis Mora-Aguilar & Delgado, 2015
Copris ciliatus
Copris colmanti Gillet, 1908
Copris complexus Nguyen-phung, 1989
Copris compressipennis Gillet, 1910
Copris confucius Harold, 1877
Copris coriarius Gillet, 1907
Copris cornifrons Boheman, 1860
Copris corniger Sahlberg, 1823
Copris corpulentus Gillet, 1910
Copris costaricensis Gahan, 1894
Copris costariensis Matthews
Copris crassus Deschodt & Davis, 2015
Copris cribratus Gillet, 1927
Copris cribricollis Gillet, 1910
Copris curlettii Moretto, 2011
Copris davidis (Deyrolle, 1878)
Copris davisi Nguyen-phung & Cambefort, 1986
Copris davisoni Waterhouse, 1891
Copris decellei Cambefort, 1992
Copris delicatus Arrow, 1931
Copris denticulatus Nguyen-phung, 1988
Copris diversus Waterhouse, 1891
Copris draco Arrow, 1906
Copris dracunculus Fereira, 1959
Copris druidum Heer, 1862
Copris dudleyi Cambefort, 1992
Copris eburneus Cambefort, 1992
Copris elpheneroides Felsche, 1910
Copris elphenor Klug, 1855
Copris erratus Lansberge, 1886
Copris evanidus Klug, 1855
Copris excisus Waterhouse, 1891
Copris fallaciosus Gillet, 1907
Copris fallax Felsche, 1910
Copris felschei Reitter, 1892
Copris fidius (Olivier, 1789)
Copris frankenbergerianus Balthasar, 1958
Copris freyi Ferreira, 1961
Copris fricator (Fabricius, 1787)
Copris fukiensis Balthasar, 1952
Copris furciceps Felsche, 1910
Copris garambae Cambefort, 1992
Copris gazellarum Gillet, 1918
Copris gibbulus Lansberge, 1886
Copris gilleti Kolbe, 1907
Copris gladiator Arrow, 1933
Copris gopheri Hubbard, 1894
Copris gracilis Waterhouse, 1891
Copris graueri Kolbe, 1914
Copris halffteri Matthews, 1959
Copris harrisi Waterhouse, 1891
Copris hartli Frey, 1975
Copris hatayamai Ochi & Araya, 1992
Copris hispanus (Linnaeus, 1764)
Copris howdeni Matthews & Halffter, 1959
Copris humilis Gillet, 1908
Copris hybridus Gillet, 1910
Copris igualensis Warner, 1990
Copris illotus Balthasar, 1942
Copris imitans Felsche, 1910
Copris inaequabilis Zhang, 1997
Copris incertus Say, 1835
Copris inemarginatus Blatchley, 1918
Copris inhalatus Quedenfeldt, 1884
Copris insidiosus Péringuey, 1901
Copris integer Reiche, 1847
Copris interioris Kolbe, 1897
Copris iris Sharp, 1875
Copris jacchoides Nguyen-phung & Cambefort, 1987
Copris jacchus (Fabricius, 1775)
Copris jahi Nguyen-phung & Cambefort, 1986
Copris johkii Ochi & Kon, 2014
Copris juanmorai Mora-Aguilar & Delgado, 2015
Copris jucundus Gillet, 1932
Copris kachinensis Ochi, Kon & Kawahara, 2008
Copris kartlinus Kabakov, 1988
Copris kasagii Ochi & Kon, 1996
Copris kasanka Josso & Prévost, 2016
Copris katangae Cambefort & Nguyen-phung, 1996
Copris keralensis Gill, 1986
Copris kiuchii Masumoto, 1989
Copris klapperichi Balthasar, 1942
Copris klugi Harold, 1869
Copris kusakabei Ochi, Kon & Kawahara, 2005
Copris laeviceps Harold, 1862
Copris laevigatus Gillet, 1927
Copris laiiformis Nguyen-phung & Cambefort, 1987
Copris laioides Boucomont, 1932
Copris laius Harold, 1868
Copris lannathai Hanboonsong & Masumoto & Ochi, 2003
Copris leakeyorum Paulian, 1976
Copris lecontei Matthews, 1962
Copris leruei Josso & Prévost, 2017
Copris lugubris Boheman, 1858
Copris lunaris [fr] (Linnaeus, 1758)
Copris macacus Lansberge, 1886
Copris macclevei Warner, 1990
Copris macer Péringuey, 1901
Copris maesi Ratcliffe, 1998
Copris magicus Harold, 1881
Copris manni Ochi, Kon & Bai, 2009
Copris marcus Gillet, 1921
Copris martinae Nguyen-phung, 1988
Copris martinezi Matthews and Halffter, 1968
Copris matthewsi Delgado & Kohlmann, 2001
Copris medogensis Zhang, 1988
Copris megaceratoides Waterhouse, 1891
Copris megasoma Matthews & Halffter, 1959
Copris mesacanthus Harold, 1878
Copris mexicanus Matthews & Halffter, 1959
Copris minutus (Drury, 1773)
Copris misellus Péringuey, 1901
Copris moechus LeConte, 1854
Copris moffartsi Gillet, 1907
Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003
Copris montivagus Gillet, 1908
Copris morphaeus Gillet, 1932
Copris mourgliai Zelenka, 1993
Copris neglectus Moxey, 1963
Copris neotridens Génier & Prévost, 2017
Copris nepos Gillet, 1908
Copris nevinsoni Waterhouse, 1891
Copris novaki Zelenka, 1992
Copris nubilosus Kohlmann & Cano & Delgado, 2003
Copris numa Lansberge, 1886
Copris obenbergeri Balthasar, 1933
Copris obesus Boheman, 1857
Copris ochus (Motschulsky, 1860)
Copris orion Klug, 1835
Copris orphanus Guérin-Méneville, 1847
Copris parapecuarius Ochi, Kon & Kawahara, 2008
Copris pauliani Nguyen-phung & Cambefort, 1986
Copris pecuarius Lewis, 1884
Copris pedarioides Lansberge, 1886
Copris phungae Cambefort, 1992
Copris phylax Gillet, 1908
Copris poggii Ochi & Kon, 2005
Copris potanini Semenov, 1891
Copris prevosti Josso, 2011
Copris pristinus Pierce, 1946
Copris pseudobootes Ferreira, 1962
Copris pseudochus Ochi & Kon, 2004
Copris pseudomoffartsi Moretto, 2011
Copris pseudosinicus Balthasar, 1958
Copris pueli Mollandin De Boissy, 1905
Copris punctatus Gillet, 1910
Copris puncticollis Boheman, 1857
Copris punctipennis Boucomont, 1914
Copris punctulatus Wiedemann, 1823
Copris punjabensis Gillet, 1921
Copris quasilaevigatus Ochi, Kon & Bai, 2007
Copris ramosiceps Gillet, 1921
Copris rebouchei Harold, 1869
Copris remotus Leconte, 1866
Copris renwarti Nguyen-phung & Cambefort, 1986
Copris repertus Walker, 1858
Copris ribbei Lansberge, 1886
Copris riekoae Ochi & Kon, 2003
Copris ritsemae Harold, 1875
Copris rugosus Gillet, 1908
Copris ruricola Balthasar, 1933
Copris sabinus Gillet, 1910
Copris sacontala Redtenbacher, 1848
Copris sallei Harold, 1869
Copris sarpedon Harold, 1868
Copris saundersi Harold, 1869
Copris schoolmeestersi Ochi & Kon, 2010
Copris scorpio Zelenka, 1993
Copris serius Nguyen-phung, 1987
Copris sexdentatus Thunberg, 1818
Copris siamensis Gillet, 1910
Copris siangensis Biswas, 1980
Copris sierrensis Matthews, 1961
Copris signatus Walker, 1858
Copris simonettai Cambefort, 1992
Copris simulator Nguyen-phung & Cambefort, 1986
Copris singularis Nguyen-phung & Cambefort, 1986
Copris sinicus Hope, 1842
Copris sinon (Olivier, 1789)
Copris sodalis Walker, 1858
Copris sonensis Bui, Dumack & Bonkowski, 2018 [3]
Copris sorex Balthasar, 1942
Copris sphaeropterus Harold, 1877
Copris spinator Harold, 1881
Copris subdolus Balthasar, 1958
Copris subpunctatus Gillet, 1910
Copris subsidens Péringuey, 1901
Copris subterraneus Heer, 1862
Copris sumatrensis Gillet, 1921
Copris surdus Arrow, 1931
Copris szechouanicus Balthasar, 1958
Copris tetraodon Gillet, 1910
Copris tridentatus Solis & Kohlmann, 2003
Copris tripartitus Waterhouse, 1875
Copris truncatus Felsche, 1901
Copris tsukamotoi Ochi, Kon & Kawahara, 2008
Copris typhoeus Gerstaecker, 1884
Copris uenoi Hanboonsong & Masumoto & Ochi, 2003
Copris umbilicatus Abeille De Perrin, 1901
Copris urus Boheman, 1857
Copris usambaricus Gillet, 1908
Copris vankhaii Nguyen-phung, 1988
Copris victorini Boheman, 1857
Copris vietnamicus Kabakov, 1994
Copris vilhenai Ferreira, 1962
Copris vrydaghi Ferreira, 1962
Copris warneri Mccleve & Kohlmann, 2005
Copris wiesei Kolbe, 1914
Copris yangi Ochi, Kon & Bai, 2007
Copris youngai Balthasar, 1967
Copris zhangi Ochi, Kon & Bai, 2009

References
Bui, Van Bac; Dumack, Kenneth; Bonkowski, Michael (2018). "Two new species and one new record for the genus Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Vietnam with a key to Vietnamese species". European Journal of Entomology. 15. doi:10.14411/eje.2018.016.
References

Bui, Van Bac; Dumack, Kenneth; Bonkowski, Michael (2018). "Two new species and one new record for the genus Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Vietnam with a key to Vietnamese species". European Journal of Entomology. 15. doi:10.14411/eje.2018.016.