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Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Histeroidea

Familia: Histeridae
Subfamilia: Histerinae
Tribus: Histerini
Genus: Atholus
Subgenera: A. (Atholus) – A. (Euatholus)
Name

Atholus Thomson, 1859
Synonymy

Peranus Lewis, 1906

References

Bousquet, Y.; Laplante, S. 1999: Les Coléoptères Histéridés du Québec Association des Entomologiste Amateurs du Québec

Atholus is a genus of clown beetles in the family Histeridae. There are more than 70 described species in Atholus.[1][2][3]
Atholus scutellaris

Species

These 78 species belong to the genus Atholus:

Atholus aequistrius (Marseul, 1854)
Atholus americanus (Paykull, 1811)
Atholus amplificipes Mazur, 2013
Atholus arcatus (Lewis, 1908)
Atholus arrowi (Desbordes, 1923)
Atholus astragali Olexa, 1987
Atholus atricolor Lewis, 1907
Atholus baberii (Lewis, 1901)
Atholus bakeri (Bickhardt, 1914)
Atholus bifrons (Marseul, 1854)
Atholus bimaculatus (Linnaeus, 1758)
Atholus bolteri Wenzel, 1944
Atholus brancuccii Mazur, 2013
Atholus cinctipygus (Lewis, 1900)
Atholus cochinchinae (Schmidt, 1889)
Atholus coelestis (Marseul, 1857)
Atholus concordans (Marseul, 1870)
Atholus confinis (Erichson, 1834)
Atholus conformis (Erichson, 1834)
Atholus corvinus (Germar, 1817)
Atholus crenatifrons (Lewis, 1899)
Atholus cycloides (Burgeon, 1939)
Atholus daldorffi (Bedel, 1906)
Atholus debeauxi (Moro, 1942)
Atholus dentipes (Lewis, 1892)
Atholus depistor (Marseul, 1873)
Atholus duodecimstriatus (Schrank, 1781)
Atholus erichsoni Lundgren in Johnson et al., 1991
Atholus euphorbiae (Peyerimhoff, 1925)
Atholus falli (Bickhardt, 1912)
Atholus famulus (Lewis, 1892)
Atholus gestroi (Schmidt, 1897)
Atholus goudotii (Marseul, 1854)
Atholus graueri (G. Müller, 1944)
Atholus helferi (Reichardt, 1932)
Atholus holzschuhi Olexa, 1988
Atholus hucheti Gomy, 2004
Atholus infirmus (Schmidt, 1889)
Atholus khnzoriani Olexa, 1982
Atholus kuijteni Kanaar, 1983
Atholus laequatus (Lewis, 1905)
Atholus lao Mazur, 2013
Atholus levis Mazur, 2013
Atholus maindronii (Lewis, 1901)
Atholus malaysi Lewis, 1908
Atholus minutus Ross, 1940
Atholus myrmidon (Marseul, 1861)
Atholus nitidissimus Desbordes, 1925
Atholus nubilus (J. L. LeConte, 1859)
Atholus omar (Lewis, 1913)
Atholus paganettii (Bickhardt, 1911)
Atholus peloponnesus Kapler, 1992
Atholus perplexus (J. L. LeConte, 1863)
Atholus philippinensis (Marseul, 1854)
Atholus pinnulae (Lewis, 1900)
Atholus pirithous (Marseul, 1873)
Atholus praetermissus (Peyron, 1856)
Atholus relictus (Marseul, 1870)
Atholus rubricatus (Lewis, 1897)
Atholus rudesculptus (Reichardt, 1922)
Atholus scutellaris (Erichson, 1834)
Atholus sedecimstriatus (Say, 1825)
Atholus sessilis (Lewis, 1899)
Atholus siculus (Tournier, 1869)
Atholus silvicola (Lewis, 1901)
Atholus singalanus (Marseul, 1880)
Atholus somali (Lewis, 1885)
Atholus staudingeri (Schmidt, 1889)
Atholus striatipennis (Lewis, 1892)
Atholus striatithorax Desbordes, 1930
Atholus tenuistriatus (Lewis, 1889)
Atholus terraemotus (Lewis, 1900)
Atholus tetricus (Lewis, 1902)
Atholus tornatus (J. L. LeConte, 1880)
Atholus torquatus (Marseul, 1854)
Atholus vacillans (Lewis, 1900)
Atholus viennai Secq, 2000
Atholus yelamosi Gomy, 2011

References

"Atholus Report". Integrated Taxonomic Information System. Retrieved 2019-09-23.
"Atholus". GBIF. Retrieved 2019-09-23.

"Atholus genus Information". BugGuide.net. Retrieved 2019-09-23.

Further reading

Bousquet, Yves; Laplante, Serge (1999). "Les Coléoptères Histéridés du Québec". Fabreries Supplément. 8: 1–190. ISSN 0318-6725.
Mazur, Slawomir (2011). A Concise Catalogue of the Histeridae (Insecta: Coleoptera). WULS - SGGW Press. ISBN 978-83-7583-3171.
Lobl, I.; Smetana, A., eds. (2015). Catalogue of Palaearctic Coleoptera, Volume 2: Hydrophiloidea - Staphylinoidea. Apollo Books. ISBN 978-90-04-29685-5.

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