Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohort: Theria
Cohort: Eutheria
Cohort: Placentalia
Cladus: Boreoeutheria
Superordo: Laurasiatheria
Cladus: Ferae
Ordo: Carnivora
Subordo: Feliformia
Familia: †Nimravidae
Subfamilia: †Barbourofelinae
Genus: Sansanosmilus (extinct)
Sansanosmilus is an extinct genus of carnivorous mammal of the family Barbourofelidae (false saber-tooth cats) endemic to Europe, which lived during the Miocene, 13.6—11.1 mya, existing for approximately 2.5 million years.[1]
Taxonomy
S. palmidens and Necromanis
Sansanosmilus is a member of the family Barbourofelidae, a group of feliform carnivorans related to either felids[2] or nimravids.[3] It had short legs, was very muscular and had a long tail. Sansanosmilus was 1.5 m long and probably weighed around 80 kg. In 1961, paleontologist L. Ginsburg concluded that Sansanosmilus was possessed of a plantigrade walking stance, after studying its foot bones and comparing it with those of the true felid Pseudaelurus from the same site. This is different from later barbourofelids, which are believed to have had semi-plantigrade or semi-digitigrade stances.[4]
The type species, Sansanosmilus palmidens, is known from fossils from the Orleanian and Astaracian stages in France. Although Albanosmilus was seen as a junior synonym of Sansanosmilus from the 1970s onwards, Robles et al. (2013) demonstrated that the type species of Albanosmilus, S. jourdani (which they considered to be a senior synonym of S. vallesiensis), is more closely related to Barbourofelis than to the type species of Sansanosmilus and thus generically distinct.[5] Wang et al. (2020) agreed with Robles et al. (2013) in recovering Albanosmilus as closer to Barbourofelis than to Sansanosmilus.[6]
References
PaleoBiology Database: Sansanosmilus, basic info
Michale Morlo; Stéphane Peigné & Doris Nagel (January 2004). "A new species of Prosansanosmilus: implications for the systematic relationships of the family Barbourofelidae new rank (Carnivora, Mammalia)". Zoological Journal of the Linnean Society. 140 (1): 43. doi:10.1111/j.1096-3642.2004.00087.x.
Xiaoming Wang; Stuart C. White; Jian Guan (2020). "A new genus and species of sabretooth, Oriensmilus liupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids". Journal of Systematic Palaeontology. Online edition (9): 783–803. doi:10.1080/14772019.2019.1691066. S2CID 211545222.
Anton, Mauricio (2013). Sabertooth.
Josep M. Robles; David M. Alba; Josep Fortuny; Soledad De Esteban-Trivigno; Cheyenn Rotgers; Jordi Balaguer; Raül Carmona; Jordi Galindo; Sergio Almécija; Juan V. Bertó & Salvador Moyà-Solà (2013). "New craniodental remains of the barbourofelid Albanosmilus jourdani (Filhol, 1883) from the Miocene of the Vallès-Penedès Basin (NE Iberian Peninsula) and the phylogeny of the Barbourofelini". Journal of Systematic Palaeontology. 11 (8): 993–1022. doi:10.1080/14772019.2012.724090. S2CID 85157737.
Wang, X., White, S., & Guan, J. (2020). A new genus and species of sabretooth, Oriensmilus liupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids. JOURNAL OF SYSTEMATIC PALAEONTOLOGY, 18(9), 783-803. http://dx.doi.org/10.1080/14772019.2019.1691066 Retrieved from https://escholarship.org/uc/item/0g62362j
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