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Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Cladus: Boreoeutheria
Superordo: Laurasiatheria
Cladus: Scrotifera
Cladus: Ferungulata
Cladus: Ferae
Cladus: Pancarnivora
Cladus: Carnivoramorpha
Cladus: Carnivoraformes
Ordo: Carnivora
Subordo: Feliformia

Familia: †Nimravidae
Subfamilia: †Barbourofelinae
Genus: †Prosansanosmilus
Species: †P. eggeri – †P. peregrinus
Name

Prosansanosmilus Heizmann, Ginsburg & Bulot, 1980: 9

Type species: Prosansanosmilus peregrinus [Heizmann et al., 1980: 9]
References
Primary references

Heizmann, E.P.J., Ginsburg, L. & Bulot, C. 1980. Prosansanosmilus peregrinus, ein neuer machairodontider Felide aus dem Miocän Deutschlands und Frankreichs. Stuttgarter Beiträge zur Naturkunde B 58: 1–27. BHL Reference page.

Additional references

Morlo, M., Peigné, S. & Nagel, D. 2004. A new species of Prosansanosmilus: implications for the systematic relationships of the family Barbourofelidae new rank (Carnivora, Mammalia). Zoological Journal of the Linnean Society 140(1): 43–61. DOI: 10.1111/j.1096-3642.2004.00087.x ResearchGate Reference page.

Prosansanosmilus is an extinct genus of barbourofelid that lived in Europe during the Late Miocene epoch (16.9—7.246 mya), existing for approximately 9.654 million years.[1] It contains Prosansanosmilus peregrinus, which died out in the Miocene epoch.
Taxonomy

Prosansanosmilus was named in 1980 by Heizmann et al. with the type species Prosansanosmilus peregrinus.[2] It was assigned to Nimravinae by Heizmann et al. (1980); to Felidae by Carroll (1988); to Barbourofelinae by Bryant (1991); and to Barbourofelidae by Morlo et al. (2004) and Morlo (2006).

A second species, P. eggeri from the Middle Miocene locality of Sandelzhausen, Germany, was described in 2004. It differed from other European barbourofelids in having a more plesiomorphic ("ancestral") morphology, with less developed sabretooth adaptations and being smaller. However, the species is stratigraphically younger than P. peregrinus; and probably part of the African faunal immigration into Europe during the Middle Eocene.[3]
Distribution

P. peregrinus lived in MN4 of France and Germany.[3] Two fossils of P. peregrinus have been found in France and another two have been found in Germany. P. peregrinus was a ground-dwelling creature.[2]
Morphology

Like all barbourofelids, Prosansanosmilus was very muscular, short legged and probably walked plantigrade (flat-footed). There are only two species of Prosansanosmilus, which lived in Spain, France and Germany during the Late Miocene epoch.
Time range

P. peregrinus is thought to have lived between 16.9 and 16 Mya. New evidence suggests that P. peregrinus lived 20–16 Mya. German scientists excavated a fossil of P. peregrinus dating 20 Mya. Other scientists think that P. peregrinus lived between 16.9 and 15.7 Mya.
References

Prosansanosmilus in the Paleobiology Database
Heizmann, E. P. J.; Ginsburg, L.; Bulot, C. (1980). "Prosansanosmilus peregrinus, ein neuer machairodontider Felide aus dem Miocän Deutschlands und Frankreichs" [Prosansanosmilus peregrinus, a new machairodontid felid from the Miocene of Germany and France]. Stuttgarter Beiträge zur Naturkunde Ser. B. (in German). 58: 1–27.
Morlo, Michael; Peigné, Stéphane; Nagel, Doris (2004). "A new species of Prosansanosmilus: implications for the systematic relationships of the family Barbourofelidae new rank (Carnivora, Mammalia)". Zoological Journal of the Linnean Society. 140 (1): 43–61. doi:10.1111/j.1096-3642.2004.00087.x.

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