Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Cladus: Boreoeutheria
Superordo: Laurasiatheria
Cladus: Scrotifera
Cladus: Ferungulata
Cladus: Euungulata
Ordo: Artiodactyla
Cladus: Artiofabula
Cladus: Cetruminantia
Subordo: Whippomorpha
Infraordo: Cetacea
Cladus: Neoceti
Parvordo: Odontoceti
Familia: Ziphiidae
Subfamilia: Hyperoodontinae
Genus: Mesoplodon
Species (15 + 3†): M. bidens – . bowdoini – M. carlhubbsi – M. densirostris – M. europaeus – M. ginkgodens – M. grayi – M. hectori – M. hotaula – M. layardii – M. mirus – M. perrini – M. peruvianus – M. stejnegeri – M. traversii – †M. posti – †M. slangkopi – †M. tumidirostris
Name
Mesoplodon Gervais, 1850
References
(2016) „A new early Pliocene species of Mesoplodon: a calibration mark for the radiation of this species-rich beaked whale genus”. Journal of Vertebrate Paleontology 36, e1055754. o. DOI:10.1080/02724634.2015.1055754.
G. Bianucci, O. Lambert, and K. Post. 2007. A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa. Geodiversitas 29(4):561-618
Miyazaki, N.; Hasegawa, Y. 1992: A new species of fossil beaked whale, Mesoplodon tumidirostris sp. nov. (Cetacea, Ziphiidae) from the Central North Pacific. Bulletin of the National Science Museum (A), 18: 167–174.
Links
Mesoplodon in Mammal Species of the World.
Wilson, Don E. & Reeder, DeeAnn M. (Editors) 2005. Mammal Species of the World – A Taxonomic and Geographic Reference. Third edition. ISBN 0-8018-8221-4.
Vernacular names
日本語: オウギハクジラ属
Türkçe: Gagalı balina
Mesoplodont whales are 16 species of toothed whale in the genus Mesoplodon, making it the largest genus in the cetacean order.[2] Two species were described as recently as 1991 (pygmy beaked whale) and 2002 (Perrin's beaked whale), and marine biologists predict the discovery of more species in the future.[3] A new species was described in 2021.[4] They are the most poorly known group of large mammals.[2] The generic name "mesoplodon" comes from the Greek meso- (middle) - hopla (arms) - odon (teeth), and may be translated as 'armed with a tooth in the centre of the jaw'.[3]
Physical description
The flank of a stranded whale, showing several round scars
Round scars from cookiecutter shark bites can be seen on the flank of this stranded Gray's beaked whale.
Mesoplodont beaked whales are small whales, 3.9 m (13 ft) (pygmy beaked whale) to 6.2 m (20 ft) (strap-toothed whale) in length,[3] even compared with closely related whales such as the bottlenose whales and giant beaked whales. The spindle-shaped body has a small dorsal fin and short and narrow flippers. The head is small and tapered and has a semicircular blow hole that is sometimes asymmetric. The beak, which vary in length between species, blends with the small melon without a crease.[3]
Sexual dimorphism is poorly known, but the females tend to be the same size or larger than males at least in some species.[3] The males typically have a bolder coloration and a unique dentition. The males of most species are covered in scars from the teeth of other males.
The lower jaw often forms a huge arch in some species, sometimes extending above the rostrum in a shape comparable to a playground slide. Every species has large (sometimes tusk-like) teeth of variable size, shape, and position.
Both sexes often have bites from cookie-cutter sharks. The dorsal fin is rather small and located between two-thirds and three-quarters down the back of the animal. Information on longevity and lactation is non-existent, and information on gestation is nearly so.
Behavior
Most species are very rarely observed, and little is known about their behavior.[2] They are typically found in groups, possibly segregated between sexes. Some species are so uncommon, they have yet to be observed alive.[2] On the surface, they are typically very slow swimmers and do not make obvious blows. They have never been observed raising their flukes above the water. They are all very deep divers, and many feed entirely on squid.
Conservation
The mesoplodonts are completely unknown as far as population estimates are concerned. They have been hunted occasionally by the Japanese, but never directly. They are also accidentally captured in drift nets. It is not known what effect this has on the population.
Species
Andrews' beaked whale (M. bowdoini) Andrews, 1908
Blainville's beaked whale (M. densirostris) Blainville, 1817
Deraniyagala's beaked whale (M. hotaula) Deraniyagala, 1963[5]
Gervais's beaked whale (M. europaeus) Gervais, 1855
Ginkgo-toothed beaked whale (M. ginkgodens) Nishiwaki and Kamiya, 1958
Gray's beaked whale (M. grayi) von Haast, 1876
Hector's beaked whale (M. hectori) Gray, 1871
Hubbs' beaked whale (M. carlhubbsi) Moore, 1963
Perrin's beaked whale (M. perrini) Dalebout, Mead, Baker, Baker & van Helden, 2002
Pygmy beaked whale (M. peruvianus) Reyes, Mead, and Van Waerebeek, 1991
Sowerby's beaked whale (M. bidens) Sowerby, 1804
Spade-toothed whale (M. traversii) Gray, 1874
Stejneger's beaked whale (M. stejnegeri) True, 1885
Strap-toothed whale (M. layardii) Gray, 1865
True's beaked whale (M. mirus) True, 1913
Ramari's beaked whale (M. eueu) Carroll et al, 2021[4]
Longman's beaked whale (Indopacetus pacificus, also known as the Indo-Pacific beaked whale or the tropical bottlenose whale) was originally assigned to Mesoplodon, but Joseph Curtis Moore placed it in its own genus, Indopacetus, a taxonomic assignment which has been followed by all researchers.[3]
Four extinct species of Mesoplodon are known, M. longirostris[6] M. posti,[7] M. slangkopi.[8] and M. tumidirostris.[9]
References
Notes
Wilson, D. E.; Reeder, D. M., eds. (2005). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. ISBN 978-0-8018-8221-0. OCLC 62265494.
Pitman 2008, p. 721
Pitman 2008, Characters and Taxonomic Relationships, pp. 721–724
Carroll et al. 2021.
Dalebout et al. 2014
Bianucci, Giovanni (1997). "The Odontoceti (Mammalia, Cetacea) from Italian Pliocene. Systematics and phylogenesis of Delphinidae". Palaeontographia Italica.
Olivier Lambert and Stephen Louwye (2016). "A new early Pliocene species of Mesoplodon: a calibration mark for the radiation of this species-rich beaked whale genus". Journal of Vertebrate Paleontology 36 (2): e1055754. doi:10.1080/02724634.2015.1055754.
G. Bianucci, O. Lambert, and K. Post. 2007. A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa. Geodiversitas 29(4):561-618
Miyazaki, N.; Hasegawa, Y. 1992: A new species of fossil beaked whale, Mesoplodon tumidirostris sp. nov. (Cetacea, Ziphiidae) from the Central North Pacific. Bulletin of the National Science Museum (A), 18: 167–174.
Sources
Dalebout, M. L.; Baker, C. S.; Steel, D.; Thompson, K.; Robertson, K. M.; Chivers, S. J.; Perrin, W. F.; Goonatilake, M.; Anderson, R. C.; Mead, J. G.; Potter, C. W.; Thompson, L.; Jupiter, D.; Yamada, T. K. (2014). "Mesoplodon hotaula: Resurrection of Mesoplodon hotaula Deraniyagala, 1963: A New Species of Beaked Whale in the tropical Indo-Pacific". Marine Mammal Science. 30 (3): 1081–1108. doi:10.1111/mms.12113. hdl:1957/51503.
Pitman, R. (2008). "Mesoplodont Whales (Mesoplodon spp.)". In Perrin, W. F.; Würsig, B.; Thewissen, J. G. M. (eds.). Encyclopedia of Marine Mammals (2nd ed.). Academic Press. pp. 721–726. ISBN 978-0-12-373553-9.
Carroll, Emma L.; McGowen, Michael R.; McCarthy, Morgan L.; Marx, Felix G.; Aguilar, Natacha; Dalebout, Merel L.; Dreyer, Sascha; Gaggiotti, Oscar E.; Hansen, Sabine S.; van Helden, Anton; Onoufriou, Aubrie B.; Baird, Robin W.; Baker, C. Scott; Berrow, Simon; Cholewiak, Danielle; Claridge, Diane; Constantine, Rochelle; Davison, Nicholas J.; Eira, Catarina; Fordyce, R. Ewan; Gatesy, John; Hofmeyr, G. J. Greg; Martín, Vidal; Mead, James G.; Mignucci-Giannoni, Antonio A.; Morin, Phillip A.; Reyes, Cristel; Rogan, Emer; Rosso, Massimiliano; Silva, Mónica A.; Springer, Mark S.; Steel, Debbie; Olsen, Morten Tange (27 October 2021). "Speciation in the deep: genomics and morphology reveal a new species of beaked whale Mesoplodon eueu". Proceedings of the Royal Society B: Biological Sciences. 288 (1961): 20211213. doi:10.1098/rspb.2021.1213. PMC 8548795. Retrieved 27 October 2021.
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