Menoceras

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Cladus: Boreoeutheria
Superordo: Laurasiatheria
Cladus: Scrotifera
Cladus: Ferungulata
Cladus: Euungulata
Ordo: Perissodactyla
Subordo: Ceratomorpha
Superfamilia: Rhinocerotoidea

Familia: Rhinocerotidae
Genus: †Menoceras

Menoceras ("Crescent Horns"[4]) is a genus of extinct, small rhinocerotids endemic to most of southern North America and ranged as far south as Panama during the early Miocene epoch. It lived from around 30.7—19.7 Ma, existing for approximately 11 million years.
Description

Male Menoceras sported two horns side by side at the tip of the nose, whereas the females were hornless. All other rhino genera, save the related genus Diceratherium, have their horns arranged one behind the other. Both sexes of Menoceros grew to a length of 5 feet (1.5 meters) long.
Paleobiology
1913 illustration of M. arikarense.

Menoceras roamed across a tropical, savanna-like grassland and plains environment that covered much of North America.[5] Because of the massive accumulations of fossil bones of this animal, particularly at Agate Springs Nebraska, Menoceras may have lived in large herds. Other sites include Martin-Anthony site Martin County, Florida, and Cady Mountains Horse Quarry, San Bernardino County, California.[6]
Taxonomy
Female (top) and male (bottom) M. arikarense skulls

Menoceras was named by Troxell and assigned to Rhinocerotidae in 1921. It was synonymized subjectively with Diceratherium by Matthew in 1931 and Wood in 1964. Again assigned to Rhinocerotidae by Prothero, Guerrin, Manning in 1989.[7] Tanner (1969), Wilson and Schiebout (1981), Prothero and Manning (1987), Carroll (1988) and Prothero et al. (1989); and to Menoceratinae by Prothero (1998).[8]
Fossil distribution
Life reconstruction of M. barbouri

Fossil distribution is as far north as New Jersey, south to Florida (3 collections) and Texas (6 collections), as far west as Nebraska (7 collections) and California (2 collections).

The Panamanian find was determined to be 19.7 Ma (AEO). It was found in the Gaillard Cut in Panama in "a 45 m thick section (narrow stratigraphic interval)"[9] It was reposited in the Smithsonian Institution, Washington, D.C. Other sites:

Lay Ranch Beds, Goshen County, Wyoming, a deposit of several carnivores and herbivores.
Agate Springs Quarries, Sioux County, Nebraska, a deposit of Miocene herbivores and carnivores such as Moropus elatus, Cynelos, Cephalogale, and a number of Artiodactyla.
Martin Canyon Quarry A, Logan County, Colorado, a very substantial number of carnivores and herbivores.

Notes

Prothero, 2005, p. 189.
Prothero, 2005, p. 59.
Prothero, 2005, pp. 65-67.
"Glossary. American Museum of Natural History". Archived from the original on 20 November 2021.
Prothero, 2005
PaleoDB collection 19233, authorized by Dr. John Alroy, May 15, 2002.
D. R. Prothero, C. Guerin, and E. Manning. 1989. The history of the Rhinocerotoidea. In D. R. Prothero and R. M. Schoch (eds.), The Evolution of Perissodactyls 321-340.
D. R. Prothero. 1998. Rhinocerotidae. In C. M. Janis, K. M. Scott, and L. L. Jacobs (eds.), Evolution of Tertiary mammals of North America 595-605.

Cucaracha Formation, Gaillard Cut, Panama. J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals.

References

Prothero, Donald R. 2005. The Evolution of North American Rhinoceroses. Cambridge University Press, Cambridge, 218 pp. ISBN 0-521-83240-3