Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Superordo: Xenarthra
Ordo: Cingulata
Familia: †Peltephilidae
Genus: †Epipeltephilus
Species: E. bifasciatus – E. forficulatus – E. kanti
Name
†Epipeltephilus Ameghino, 1904
References
Ameghino, F. 1904: Nuevas especies de mamiferos Cretaceos y Terciarios de la Republica Argentina. Anales de la Sociedad Científica Argentina, 58: 56–71.
Epipeltephilus is an extinct genus of armadillo, belonging to the family Peltephilidae, the "horned armadillos", whose most famous relative was Peltephilus. Epipeltephilus is the last known member of its family, becoming extinct during the Chasicoan period. It was found in the Rio Mayo Formation and the Arroyo Chasicó Formation of Argentina, and in northern Chile.
History and Etymology
Epipeltephilus was originally described in 1904 by Florentino Ameghino, its holotype being MACN A 11641, which included remains of the occipital and frontal areas of the skull, and fragments of the right mandible preserving five molariformes, coming from the Mayoan Rio Mayo Formation. [1] In 1915, remains from the Arroyo Chasicó Formation are recovered, and they were first mentioned in literature in 1931. In 2012, Gonzalez-Ruiz, Scillato-Yané, Krmpotic and Carlini described the second species, E. kanti, with MLP 92-XI-19-7, three osteoderms, as holotype, with other osteoderms and a single molariform as among the remaining material associated to the species, all coming from the Arroyo Chasicó Formation.[2] Finally, in 2016, a new species, E. caraguensis, is described from remains coming from Northern Chile by Montoya-Sanhuez, Moreno, Bobe, Carrano, García and Corgne, with, as holotype, SGO-PV 21102, composed of four osteoderms and fragmentary cranial and postcranial remains.[3]
Epipeltephilus means, in Latin, "above Peltephilus".
Description
Epipeltephilus's skull was larger than Peltephilus, and lower and wider in its posterior area, and its temporals were more extended laterally. The sagittal and occipital crests were less visible on Epipeltephilus, and its mandibles were larger, with vertically implanted, sub-elliptical molariforms, the last one being of similar size with the others, in opposition to several of its relatives. Like its relatives, it had the typical "horn" osteoderms.[1][2]
Species
Epipeltephilus recurvus
E. recurvus is the type species, described in 1904 by Ameghino. It is only known from remains coming from the Mayo Formation in Mayoan-aged rocks. While its holotype is only composed of the remains of a skull, several peltephilid osteoderms from the locality are commonly associated with the genus.
Epipeltephilus kanti
E. kanti is the last known member of the genus Epipeltephilus, and of the peltephilids as a whole. It is known from Chasicoan remains found in the Arroyo Chasicó Formation. The longitudinal crests of the quadrangular osteoderms were higher and more developed than in E. recurvus, and their exposed surface was rougher and more similar to those of Peltephilus and Peltecoelus. The teeth were subtriangular.[2]
The species was named in honour of the Prussian philosopher Immanuel Kant.[2]
Epipeltephilus caraguensis
E. caraguensis was found in Mayoan period rocks from the Chilean Precordillera. Its osteoderms were larger and very different from all other horned armadillos, with a tubular, rough and raised anterior edge.
The name "caraguensis" means "from Caragua", the locality near the town of Putre where the remains of the species were extracted.[3]
Palaeoecology
While originally recovered as carnivorous armadillos, peltephilids are now considered to be more generalist, able to dig to find tough tubers, but also eating potentially carrion.[4] Finally, in 2016, a new species, E. caraguensis, is described from remains coming from Northern Chile by Montoya-Sanhuez, Moreno, Bobe, Carrano, García and Corgne, with, as holotype, SGO-PV 21102, composed of four osteoderms and fragmentary cranial and postcranial remains.[3] They were probably burrowers. While on the edge of extinction during the late Miocene, they were still widely distributed, known from Patagonia to the Central Andes.[3]
References
Gonzalez-Ruiz, Laureano (October 2010). 1. Los Cingulata (Mammalia, Xenarthra) del Mioceno temprano y medio de Patagonia (edades Santacrucense y "Friasense"). Revisión sistemática y consideraciones bioestratigráficas (Doctor). Facultad de Ciencias Naturales y Museo Universidad Nacional de La Plata.
Gonzalez-Ruiz, L.; Scillato-Yané, G.; Krmpotic, C.; Carlini, A. (2012). "A new species of Peltephilidae (Mammalia: Xenarthra: Cingulata) from the late Miocene (Chasicoan SALMA) of Argentina". Zootaxa. 3359: 55–64. doi:10.11646/zootaxa.3359.1.5. hdl:11336/196599.
Montoya-Sanhueza, G.; Moreno, K.; Bobe, R.; Carrano, M.T.; García, M.; Corgne, A. (2017). "Peltephilidae and Mesotheriidae (Mammalia) from late Miocene strata of Northern Chilean Andes, Caragua". Journal of South American Earth Sciences. 75: 51–65. Bibcode:2017JSAES..75...51M. doi:10.1016/j.jsames.2017.01.009.
Vizcaino, S.F.; FARIÑA, R.A. (2007). "Diet and locomotion of the armadillo Peltephilus: a new view". Lethaia. 30: 79–86. doi:10.1111/j.1502-3931.1997.tb00446.x.
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