Udea

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Pyraloidea

Familia: Crambidae
Subfamilia: Pyraustinae
Tribus: Spilomelini
Genus: Udea
Species: U. absolutalis – U. abstrusa – U. accolalis – U. aenigmatica – U. affinialis – U. afghanalis – U. aksualis – U. alaskalis – U. albipunctalis – U. albostriata – U. alpinalis – U. altaica – U. amitina – U. angustalis – U. annectans – U. ardekanalis – U. argoscelis – U. asychanalis – U. aurora – U. austriacalis – U. autoclesalis – U. azorensis – U. berberalis – U. beringialis – U. binoculalis – U. bipunctalis – U. bourgognealis – U. brevipalpis – U. brontias – U. bryochloris – U. cacuminicola – U. caliginosalis – U. calliastra – U. caminopis – U. capsifera – U. carniolica – U. cataphaea – U. chalcophanes – U. chloropis – U. chytropa – U. cinerea – U. confinalis – U. conisalias – U. constricta – U. conubialis – U. coranalis – U. costalis – U. costiplaga – U. crambialis – U. cretacea – U. curvata – U. cyanalis – U. decoripennis – U. decrepitalis – U. defectalis – U. delineatalis – U. derasa – U. despecta – U. detersalis – U. diopsalis – U. donzelalis – U. dracontias – U. dryadopa – U. elutalis – U. endopyra – U. endotrichialis – U. ennychioides – U. ephippias – U. epicooena – U. eucrena – U. exalbalis – U. exigualis – U. ferrealis – U. ferrugalis – U. fimbriatralis – U. flavofimbriata – U. fulcrialis – U. fumipennis – U. fusculalis – U. galapagensis – U. gigantalis – U. grisealis – U. hageni – U. hamalis – U. helioxantha – U. helviusalis – U. heterodoxa – U. hyalistis – U. ialis – U. ichinosawana – U. illineatalis – U. incertalis – U. indistincta – U. indistinctalis – U. inferioralis – U. infuscalis – U. inhospitalis – U. inquinatalis – U. institalis – U. intermedia – U. itysalis – U. juldusalis – U. karagaialis – U. khorassanalis – U. kusnezovi – U. lagunalis – U. lampadias – U. languidalis – U. latipennalis – U. lenta – U. lerautalis – U. liopis – U. liorea – U. livida – U. lugubralis – U. lutealis – U. maderensis – U. mandronalis – U. mechedalis – U. melanephra – U. melanopis – U. melanosticta – U. melanostictalis – U. metasema – U. micacea – U. montanalis – U. montensis – U. monticolans – U. murinalis – U. nea – U. nebulalis – U. nebulatalis – U. nigrescens – U. nigropunctata – U. nigrostigmalis – U. nomophiloides – U. nordeggensis – U. nordmani – U. numeralis – U. ochreocapitalis – U. ochropera – U. octosignalis – U. olivalis – U. ommatias – U. orbicentralis – U. pachygramma – U. paghmanalis – U. phaealis – U. phaethontia – U. phyllostegia – U. planalis – U. platyleuca – U. plumbalis – U. poasalis – U. poliostolalis – U. praefulvalis – U. praepetalis – U. profundalis – U. proximalis – U. prunalis – U. pseudocrocealis – U. psychropa – U. punctiferalis – U. punoalis – U. pyranthes – U. pyraustiformis – U. radiosalis – U. ragonoti – U. renalis – U. rhododendronalis – U. rubigalis – U. ruckdescheli – U. russispersalis – U. rusticalis – U. sabulosalis – U. saxifragae – U. schaeferi – U. scoparialis – U. scorialis – U. secernalis – U. secticostalis – U. sheppardi – U. sideralis – U. simplicella – U. sobrinalis – U. soratalis – U. stationalis – U. stellata – U. stigmatalis – U. subplanalis – U. suisharyonensis – U. suralis – U. swezeyi – U. tachdirtalis – U. tenoalis – U. testacea – U. tetragramma – U. thermantis – U. thermantoidis – U. thoonalis – U. thyalis – U. torvalis – U. tritalis – U. turmalis – U. uliginosalis – U. umbriferalis – U. uralica – U. vacunalis – U. vastalis – U. violae – U. viridalis – U. washingtonalis – U. zernyi
Name

Udea Guenée, 1845

Type species: Pyralis ferrugalis Jacob Hübner, 1796
References

Bolshakov, L.V. 2002. New species of pyraloid moths from the Centre of European Russia (Lepidoptera: Pyraustidae). Russian Entomological Journal 11(2): 225–228. Full article (PDF). Reference page.
Inoue, H., Yamanaka, H. & Sasaki, A. 2008: Revision of Udea orbicentralis-complex from Japan, with descriptions of four new species (Pyralidae, Pyraustinae). Tinea 20(2): 85–94. Reference page.
Kirpichnikova, V.A. & Yamanaka, H. 2014: Additional information about the genus Udea Guenee (Crambidae, Pyraustinae) from the Russian Far East. |Tinea 22(5): 313–315. Reference page. .
Landry, B. 2016. Taxonomic revision of the Spilomelinae (Lepidoptera, Pyralidae s. l.) of the Galápagos Islands, Ecuador. Revue Suisse de Zoologie 123(2): 315–399. Reference page.
Mally, R., Huemer, P. & Nuss, M. 2018. Deep intraspecific DNA barcode splits and hybridisation in the Udea alpinalis group (Insecta, Lepidoptera, Crambidae) – an integrative revision. ZooKeys 746: 51–90. DOI: 10.3897/zookeys.746.22020. Reference page.
Mally, R. & Nuss, M. 2011: Molecular and morphological phylogeny of European Udea moths (Insecta: Lepidoptera: Pyraloidea). Arthropod systematics & phylogeny, 69(1): 55–71. PDF.
Mally, R., Segerer, A.H. & Nuss, M. 2016. Udea ruckdescheli sp. n. from Crete and its phylogenetic relationships (Pyraloidea, Crambidae, Spilomelinae). Nota Lepidopterologica 39(2): 123–135. Reference page.
Munroe, E.G. 1966. Revision of North American Species of Udea Guenée (Lepidoptera: Pyralidae). The Memoirs of the Entomological Society of Canada 49: 5–57. DOI: 10.4039/entm9849fv. Reference page.
Mutuura, A. 1954: Classification of Japanese Pyrausta Group Based on the Structure of the Male and Female Genitalia (Pyr.: Lep.)(Agronomy). Bulletin of the Naniwa University Ser. B, Agriculture and biology 4: 7–33. full article Reference page.
Nuss, M., Landry, B., Mally, R., F. Vegliante, A. Tränkner, F. Bauer, J. Hayden, A. Segerer, R. Schouten, H. Li, T. Trofimova, M. A. Solis, J. De Prins & W. Speidel 2003–2019. Global Information System on Pyraloidea. [www.pyraloidea.org]
Sviridov, A.V. & Korb, S.K. 2010. An illustrated catalogue of primary type specimens of the Lepidoptera in collection of the Zoological Museum of Moscow State University described in the second half of XXth century. Eversmannia 21/22: 6–27. Reference page.
Yamanaka, H. 1985: Revision of Udea lugubralis-complex (Lepidoptera: Pyralidae) from Japan, with description of a new species. Tinea 12(12): 105-112. Reference page.
Zhang, D-D. & Li, J-W. 2016. Two new species and five newly recorded species of the genus Udea Guenée from China (Lepidoptera, Crambidae). ZooKeys 565: 123-139. DOI: 10.3897/zookeys.565.6304.Reference page.

Vernacular names
English: Celery leaftier

Udea is a genus of snout moths in the subfamily Spilomelinae of the family Crambidae. The genus was erected by Achille Guenée in 1845.[2] The currently known 214 species are present on all continents except Antarctica.[1] About 41 species are native to Hawaii.[3]

Systematics

The genus is placed in the tribe Udeini, where it is closest related to the genera Mnesictena, Deana and Udeoides.[3][4] In the past, Mnesictena was included in Udea,[5] but it is currently considered a separate genus comprising seven species.[1]

The North American and European Udea species have been treated in several studies, and a number of species groups has been proposed (see below).[3][6][7][8]
Species

Udea ferrugalis species group:[3]

Udea accolalis (Zeller, 1867)
Udea azorensis Meyer, Nuss & Speidel, 1997
Udea delineatalis (Walker in Melliss, 1875)
Udea exigualis (Wileman, 1911)
Udea ferrugalis (Hübner, 1796)
Udea hageni Viette, 1952
Udea heterodoxa (Meyrick, 1899)
Udea liopis (Meyrick, 1899)
Udea lugubralis (Leech, 1889)
Udea maderensis (Bethune-Baker, 1894)
Udea montensis Mutuura, 1954
Udea nordmani (Rebel, 1935)
Udea profundalis (Packard, 1873)
Udea pyranthes (Meyrick, 1899)
Udea rubigalis (Guenée, 1854)
Udea rusticalis (Barnes & McDunnough, 1914)
Udea stationalis Yamanaka, 1988

Udea itysalis species group:[3]

Udea abstrusa Munroe, 1966
Udea beringialis Munroe, 1966
Udea brevipalpis Munroe, 1966
Udea cacuminicola Munroe, 1966)
Udea costalis (Eversmann, 1852)
Udea derasa Munroe, 1966
Udea itysalis (Walker, 1859)
Udea livida Munroe, 1966
Udea radiosalis (Möschler, 1883)
Udea tachdirtalis (Zerny, 1935)
Udea turmalis (Grote, 1881)

Udea numeralis species group:[3][7]

Udea ardekanalis Amsel, 1961
Udea bipunctalis (Herrich-Schäffer, 1848)
Udea confinalis (Lederer, 1858)
Udea cyanalis (La Harpe, 1855)
Udea exalbalis (Caradja, 1916)
Udea fimbriatralis (Duponchel, 1833)
Udea fulvalis (Hübner, 1809)
Udea institalis (Hübner, 1819)
Udea languidalis (Eversmann, 1842)
Udea lutealis (Hübner, 1809)
Udea numeralis (Hübner, 1796)
Udea olivalis (Denis & Schiffermüller, 1775)
Udea praepetalis (Lederer, 1869)
Udea ruckdescheli Mally, Segerer & Nuss, 2016
Udea tritalis (Christoph, 1881)
Udea zernyi (Klima in Zerny, 1940)

Udea alpinalis species group:[3][8]

Udea alpinalis (Denis & Schiffermüller, 1775)
Udea altaica (Zerny, 1914)
Udea austriacalis (Herrich-Schäffer, 1851)
Udea bourgognealis Leraut, 1996
Udea carniolica Huemer & Tarmann, 1989
Udea cretacea (Filipjev, 1925)
Udea donzelalis (Guenée, 1854)
Udea juldusalis (Zerny, 1914)
Udea murinalis (Fischer von Röslerstamm, 1842)
Udea nebulalis (Hübner, 1796)
Udea plumbalis (Zerny, 1914)
Udea rhododendronalis (Duponchel, 1834)
Udea uliginosalis (Stephens, 1834)

Udea decrepitalis species group:[3]

Udea decrepitalis (Herrich-Schäffer, 1848)
Udea hamalis (Thunberg, 1792)
Udea inquinatalis (Lienig & Zeller, 1846)
Udea prunalis (Denis & Schiffermüller, 1775)

Species unplaced in any species group:

Udea absolutalis (Dyar, 1913)
Udea aenigmatica (Heinrich, 1931)
Udea affinialis (Zerny, 1914)
Udea afghanalis (Amsel, 1970)
Udea aksualis (Caradja, 1928)
Udea alaskalis (Gibson, 1920)
Udea albipunctalis Dognin, 1905
Udea albostriata Zhang & Li, 2016
Udea amitina (Butler, 1883)
Udea angustalis (Dognin, 1905)
Udea annectans Munroe, 1974
Udea argoscelis (Meyrick, 1888)
Udea asychanalis (Druce, 1899)
Udea auratalis (Warren, 1895)
Udea aurora (Butler, 1881)
Udea autoclesalis (Walker, 1859)
Udea berberalis (Barnes & McDunnough, 1918)
Udea binoculalis (Hampson, 1904)
Udea brontias (Meyrick, 1899)
Udea bryochloris (Meyrick, 1899)
Udea caliginosalis (Ragonot, 1894)
Udea calliastra (Meyrick, 1899)
Udea caminopis (Meyrick, 1899)
Udea capsifera (Meyrick, 1933)
Udea cataphaea (Meyrick, 1899)
Udea chalcophanes (Meyrick, 1899)
Udea chloropis (Meyrick, 1899)
Udea chytropa (Meyrick, 1899)
Udea cinerea (Butler, 1883)
Udea conisalias (Meyrick, 1899)
Udea constricta (Butler, 1882)
Udea conubialis Yamanaka, 1972
Udea coranialis Munroe, 1967
Udea costiplaga (Dognin, 1913)
Udea crambialis (Druce, 1899)
Udea curvata Zhang & Li, 2016
Udea decoripennis Munroe, 1967
Udea defectalis (Sauber, 1899)
Udea despecta (Butler, 1877)
Udea detersalis (Walker, 1866)
Udea diopsalis (Hampson, 1913)
Udea dracontias (Meyrick, 1899)
Udea dryadopa (Meyrick, 1899)
Udea elutalis (Denis & Schiffermüller, 1775)
Udea endopyra (Meyrick, 1899)
Udea endotrichialis (Hampson, 1918)
Udea ennychioides (Butler, 1881)
Udea ephippias (Meyrick, 1899)
Udea epicoena (Meyrick, 1937)
Udea eucrena (Meyrick, 1888)
Udea ferrealis (Hampson, 1900)
Udea flavofimbriata (Moore, 1888)
Udea fulcrialis (Sauber, 1899)
Udea fumipennis (Warren, 1892)
Udea fusculalis (Hampson, 1899)
Udea gigantalis Dognin, 1912
Udea grisealis Inoue, Yamanaka & Sasaki, 2008
Udea helioxantha (Meyrick, 1899)
Udea helviusalis (Walker, 1859)
Udea hyalistis (Lower, 1902)
Udea ialis (Walker, 1859)
Udea ichinosawana (Matsumura, 1925)
Udea illineatalis (Dognin, 1904)
Udea incertalis (Caradja in Caradja & Meyrick, 1937)
Udea indistincta (Butler, 1883)
Udea indistinctalis Warren, 1892
Udea inferioralis (Walker, 1866)
Udea infuscalis Zeller, 1852
Udea inhospitalis Warren, 1892
Udea intermedia Inoue, Yamanaka & Sasaki, 2008
Udea karagaialis (Caradja, 1916)
Udea khorassanalis (Amsel, 1950)
Udea kusnezovi Sinev, 2008
Udea lagunalis (Schaus, 1913)
Udea lampadias (Meyrick, 1904)
Udea latipennalis (Caradja, 1928)
Udea lenta (Meyrick, 1936)
Udea lerautalis Tautel, 2014
Udea litorea (Butler, 1883)
Udea mandronalis (Walker, 1859)
Udea mechedalis (Amsel, 1950)
Udea melanephra (Hampson, 1913)
Udea melanopis (Meyrick, 1899)
Udea melanosticta (Butler, 1883)
Udea melanostictalis (Hampson in Poulton, 1916)
Udea metasema (Meyrick, 1899)
Udea micacea (Butler, 1881)
Udea minnehaha (Pryer, 1877)
Udea montanalis (Schaus, 1912)
Udea monticolens (Butler, 1882)
Udea nea (Strand, 1918)
Udea nebulatalis Inoue, Yamanaka & Sasaki, 2008
Udea nigrescens (Butler, 1881)
Udea nigripunctata Warren, 1892
Udea nigrostigmalis Warren, 1896
Udea nomophilodes (Hampson, 1913)
Udea nordeggensis (McDunnough, 1929)
Udea ochreocapitalis (Ragonot, 1894)
Udea ochropera (Hampson, 1913)
Udea octosignalis (Hulst, 1886)
Udea ommatias (Meyrick, 1899)
Udea orbicentralis (Christoph, 1881)
Udea pachygramma (Meyrick, 1899)
Udea paghmanalis (Amsel, 1970)
Udea phaealis (Hampson, 1899)
Udea phaethontia (Meyrick, 1899)
Udea phyllostegia (Swezey, 1946)
Udea planalis (South in Leech & South, 1901)
Udea platyleuca (Meyrick, 1899)
Udea poasalis (Schaus, 1912)
Udea poliostolalis (Hampson, 1918)
Udea praefulvalis (Amsel, 1970)
Udea proximalis Inoue, Yamanaka & Sasaki, 2008
Udea pseudocrocealis (South in Leech & South, 1901)
Udea psychropa (Meyrick, 1899)
Udea punctiferalis (South in Leech & South, 1901)
Udea punoalis Munroe, 1967
Udea pyraustiformis (Toll, 1948)
Udea ragonotii (Butler, 1883)
Udea renalis Moore, 1888
Udea russispersalis (Zerny, 1914)
Udea sabulosalis Warren, 1892
Udea saxifragae (McDunnough, 1935)
Udea schaeferi (Caradja in Caradja & Meyrick, 1937)
Udea scoparialis (Hampson, 1899)
Udea scorialis (Zeller, 1847)
Udea secernalis (Möschler, 1890)
Udea secticostalis (Hampson, 1913)
Udea sheppardi (McDunnough, 1929)
Udea simplicella (La Harpe, 1861)
Udea sobrinalis (Guenée, 1854)
Udea soratalis Munroe, 1967
Udea stellata (Butler, 1883)
Udea stigmatalis (Wileman, 1911)
Udea subplanalis (Caradja in Caradja & Meyrick, 1937)
Udea suisharyonensis (Strand, 1918)
Udea suralis (Schaus, 1933)
Udea swezeyi (Zimmerman, 1951)
Udea tenoalis Munroe, 1974
Udea testacea (Butler, 1879)
Udea tetragramma (J. F. G. Clarke, 1965)
Udea thermantis (Meyrick, 1899)
Udea thermantoidis (Swezey, 1913)
Udea thoonalis (Walker, 1859)
Udea thyalis (Walker, 1859)
Udea torvalis (Möschler, 1864)
Udea umbriferalis (Hampson, 1918)
Udea uralica Slamka, 2013
Udea vacunalis (Grote, 1881)
Udea vastalis (Christoph in Romanoff, 1887)
Udea violae (Swezey, 1933)
Udea viridalis (Dognin, 1904)
Udea washingtonalis (Grote, 1882)

Former species

Udea catilualis (Hampson, 1900), synonymized with Udea numeralis[9]
Udea conquisitalis (Guenee, 1849), transferred to the Glaphyriinae genus Evergestis[1]
Udea illutalis (Guenée, 1854), a synonym of Udea numeralis[9]
Udea pauperalis (Staudinger, 1879), transferred to the Pyraustinae genus Pyrausta[9]
Udea perfervidalis (Hampson, 1900), transferred to the Spilomelinae genus Metasia[9]
Udea sviridovi Bolshakov, 2002, synonymized with Udea exalbalis[9]

References

Nuss, Matthias; Landry, Bernard; Mally, Richard; Vegliante, Francesca; Tränkner, Andreas; Bauer, Franziska; Hayden, James; Segerer, Andreas; Schouten, Rob; Li, Houhun; Trofimova, Tatiana; Solis, M. Alma; De Prins, Jurate; Speidel, Wolfgang (2003–2020). "Global Information System on Pyraloidea (GlobIZ)". www.pyraloidea.org. Retrieved 2020-02-01.
Pitkin, Brian; Jenkins, Paul (November 5, 2004). "Udea Guenée, 1845". Butterflies and Moths of the World. Natural History Museum, London. Retrieved June 26, 2018.
Mally, Richard; Nuss, Matthias (2011). "Molecular and morphological phylogeny of European Udea moths (Insecta: Lepidoptera: Pyraloidea)" (PDF). Arthropod Systematics & Phylogeny. 69 (1): 55–71.
Mally, Richard; Hayden, James E.; Neinhuis, Christoph; Jordal, Bjarte H.; Nuss, Matthias (2019). "The phylogenetic systematics of Spilomelinae and Pyraustinae (Lepidoptera: Pyraloidea: Crambidae) inferred from DNA and morphology" (PDF). Arthropod Systematics & Phylogeny. 77 (1): 141–204. doi:10.26049/ASP77-1-2019-07. ISSN 1863-7221.
Munroe, Eugene Gordon (1983). "Pyralidae (except Crambinae)". In Hodges, R. W.; Dominick, T.; Franclemont, J. G.; Munroe, Eugene Gordon; Powell, J. A. (eds.). Check List of the Lepidoptera of America North of Mexico Including Greenland. London: E. W. Classey Ltd. and the Wedge Entomological Research Foundation. pp. 67–76, 78–85.
Munroe, Eugene G. (1966). "Revision of the North American species of Udea Guenée (Lepidoptera: Pyralidae)". Memoirs of the Entomological Society of Canada. 49: 1–57. doi:10.4039/entm9849fv.
Mally, Richard; Segerer, Andreas H.; Nuss, Matthias (2016). "Udea ruckdescheli sp. n. from Crete and its phylogenetic relationships (Pyraloidea, Crambidae, Spilomelinae)". Nota Lepidopterologica. 39 (2): 123–135. doi:10.3897/nl.39.9090.
Mally, Richard; Huemer, Peter; Nuss, Matthias (2018). "Deep intraspecific DNA barcode splits and hybridisation in the Udea alpinalis group (Insecta, Lepidoptera, Crambidae) – an integrative revision". ZooKeys (746): 51–90. doi:10.3897/zookeys.746.22020. PMID 29674896.
Slamka, František (2013). Pyraloidea (Lepidoptera) of Europe, Volume 3 – Pyraustinae & Spilomelinae. František Slamka. pp. 1–360. ISBN 978-80-969052-8-7.