Rhaphium

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Antliophora
Ordo: Diptera
Subordo: Brachycera
Infraordo: Asilomorpha
Superfamilia: Empidoidea

Familia: Dolichopodidae
Subfamilia: Rhaphiinae
Genus: Rhaphium
Species (197): R. aequale – R. albibarbum – R. albifrons – R. albomaculatum – R. aldrichi – R. anale – R. angusticorne – R. antennatum – R. apicinigrum – R. apophysatum – R. arboreum – R. armatum – R. atkinsoni – R. auctum – R. baihuashanum – R. banksi – R. barbipes – R. basale – R. beringiense – R. bidilatatum – R. bilobum – R. bisectum – R. boreale – R. brachyceras – R. brevicorne – R. brevilamellatum – R. browni – R. bukzeevae – R. bulyginskayae – R. calcaratum – R. caliginosum – R. campestre – R. canadense – R. caudatum – R. ciliatum – R. coloradense – R. colute – R. commune – R. confine – R. congonsis – R. crassipes – R. crinitum – R. currani – R. daqinggouense – R. dialatatum – R. dichromum – R. dilatatum – R. discigerum – R. discolor – R. dispar – R. dorsiseta – R. doroninae – R. doroteum – R. dubium – R. eburnea – R. effilatum – R. elegantulum – R. elongatum – R. ensicorne – R. essoensis – R. exile – R. fasciatum – R. fascipes – R. femineum – R. femoratum – R. firsovi – R. fissum – R. flavicoxa – R. flavilabre – R. fluviatile – R. foliatum – R. furcatum – R. furcifer – R. gangchanum – R. gansuanum – R. gibsoni – R. glaciale – R. gracile – R. grande – R. grandicercum – R. gravipes – R. gruniniani – R. gussakovskii – R. heilongjiangense – R. hirtimanum – R. holmgreni – R. hungaricum – R. igorjani – R. insolitum – R. intermedium – R. jamalensis – R. johnsoni – R. laticorne – R. latifacies – R. lehri – R. longibara – R. longicorne – R. longipalpe – R. longipes – R. lugubre – R. macalpini – R. macrocerum – R. mcveighi – R. medicoris – R. mediocre – R. mediocris – R. melampus – R. micans – R. monotrichum – R. montanum – R. montium – R. nasutum – R. neimengense – R. neolatifacies – R. nigribarbatum – R. nigricoxa – R. nigrociliatum – R. nigrovittatum – R. nigrum – R. nubilum – R. nudiusculum – R. nudum – R. nuortevai – R. obscuripes – R. obtusum – R. occipitale – R. orientale – R. ornatum – R. ovsyannikovae – R. palliaristatum – R. palpale – R. parentianum – R. patellitarse – R. patulum – R. paulseni – R. pectinatum – R. pectiniger – R. penicillatum – R. picketti – R. pitkini – R. pollex – R. popularis – R. praerosum – R. psilopoda – R. punctitarse – R. qinghaiense – R. quadrispinosum – R. reaveyi – R. relatus – R. richterae – R. riparium – R. rivale – R. robustum – R. rotundiceps – R. sachalinense – R. sagittiferum – R. septentrionale – R. sexsetosum – R. shaliuhense – R. shamshevi – R. shannoni – R. sichotense – R. sichuanense – R. signiferum – R. simplicipes – R. sinense – R. slossonae – R. speciosum – R. spinitarse – R. stackelbergi – R. steyskali – R. suave – R. subarmatum – R. subfurcatum – R. temerarium – R. terminale – R. tianshuiense – R. triangulatum – R. tricaudatum – R. tridactylum – R. tripartitum – R. triste – R. tuberculatum – R. turanicola – R. umbripenne – R. vanduzeei – R. venustum – R. viklundi – R. vockerothi – R. wheeleri – R. wuduanum – R. xinjiangense – R. xipheres – R. xiphias – R. zakonnikovae – R. zetterstedti – R. zhongdianum

[Source: Catalogue of Life: 2013 Annual Checklist]

Add (7): R. borisovi – R. brooksi – R. huzhuense – R. minhense – R. sibiricus – R. srilankensis – R. subtridactylum
Name

Rhaphium Meigen, 1803: 272

Type species: Rhaphium macrocerum Meigen, 1824, by subsequent designation by Curtis, 1835: pl. 568.
Synonyms

Hydrochus Fallén, 1823 [junior homonym of Hydrochus Leach, 1817]
Type species: Hydrochus longicornis Fallen, 1823, by subsequent designation by Coquillet, 1910: 553.
Raphium Meigen, 1824: pl. 34 [incorrect spelling]
Raphium Latreille, 1829: 472 [unjustified emendation] [syn.: O’Hara, Cerretti, Pape & Evenhuis, 2011: 158]
Raphium Rondani, 1856: 146, 221 [incorrect spelling, or subsequent usage of Raphium Latreille, 1829]

References
Primary references

Coquillett, D.W. 1910. The type-species of the North American genera of Diptera. Proceedings of the United States National Museum 37: 499–647. BHL.
Latreille, P.A. 1829. Tome V. Suite et fin des insectes. In: Cuvier G.: Le règne animal distribué d'après son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction à l'anatomie comparée, Nouvelle édition, revue et augmentée. Déterville, Paris, xxii + 556 pp. BHL Reference page.
Meigen, J. 1803. Versuch einer neuen Gattungseintheilung der europäischen zweiflügeligen Insekten. Magazin für Insektenkunde 2: 259–281. BHL Reference page.
Meigen, J. 1824. Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten 4, xi i+ 428 pp., 9 pls. Hamm, Schulz-Wundermann'sche Buchhandlung. BHL. Reference page.
O’Hara, J.E., Cerretti, P., Pape, T. & Evenhuis, N.L. 2011. Nomenclatural studies toward a world list of Diptera genus-group names. Part II: Camillo Rondani. Zootaxa 3141: 1–268. Preview PDF Reference page.

Additional references

Evenhuis, N.L. & Pape, T. 2019. Nomenclatural Studies Toward a World List of Diptera Genus-Group Names Part VII: Johann Wilhelm Meigen. Zootaxa 4703(1): 1–193. DOI: 10.11646/zootaxa.4703.1.1 Paywall. Reference page.
Qilemoge, Lin, C., Qi, P., Li, D. & Yang, D. 2020. Two new species of Rhaphium from Qinghai Province, China (Diptera, Dolichopodidae, Rhaphiinae). ZooKeys 931: 73–94. DOI: 10.3897/zookeys.931.49671 Open access. Reference page.
Qilemoge, Wang, M-Q. & Yang, D. 2019. Three new species of Rhaphium from China, with an updated key to Chinese Rhaphium (Diptera, Dolichopodidae, Rhaphiinae). Zookeys 840: 87–99. DOI: 10.3897/zookeys.840.31602 Open access Reference page.
Tang, C-F., Wang, N. & Yang, D. 2016. Rhaphium (Diptera: Dolichopodidae: Rhaphiinae) from China with six new species. Zootaxa 4162(3): 581–593. DOI: 10.11646/zootaxa.4162.3.11. Reference page.

Rhaphium is a genus of flies in the family Dolichopodidae. It is the largest genus within the subfamily Rhaphiinae, with over 200 species currently known.[2]
Gallery

Rhaphium sp. in copula

Species

Rhaphium aequale Van Duzee, 1927
Rhaphium albibarbum (Van Duzee, 1924)
Rhaphium albifrons Zetterstedt, 1843
Rhaphium albomaculatum (Becker, 1891)
Rhaphium aldrichi (Van Duzee, 1922)
Rhaphium antennatum (Carlier, 1835)
Rhaphium apicinigrum Yang & Saigusa, 1999[3]
Rhaphium apophysatum Tang, Wang & Yang, 2016[4]
Rhaphium appendiculatum Zetterstedt, 1849
Rhaphium arboreum Curran, 1924
Rhaphium armatum Curran, 1924
Rhaphium atkinsoni Curran, 1926
Rhaphium auctum Loew, 1857
Rhaphium baihuashanum Yang, 1998[5]
Rhaphium banksi Van Duzee, 1926
Rhaphium barbipes (Van Duzee, 1923)
Rhaphium basale Loew, 1850
Rhaphium beringiense Negrobov & Vockeroth, 1979
Rhaphium bidilatatum (Parent, 1954)
Rhaphium bilobum Tang, Wang & Yang, 2016[4]
Rhaphium bisectum Tang, Wang & Yang, 2016[4]
Rhaphium boreale (Van Duzee, 1923)
Rhaphium borisovi Negrobov, Barkalov & Selivanova, 2012[6]
Rhaphium brachycerus Meuffels & Grootaert, 1999[7]
Rhaphium brevicorne Curtis, 1835
Rhaphium brevilamellatum Van Duzee in Curran, 1926
Rhaphium brooksi Negrobov, Barkalov & Selivanova, 2011
Rhaphium browni Curran, 1931
Rhaphium bukzeevae Grichanov, 1995
Rhaphium bulyginskayae Grichanov, 1995
Rhaphium calcaratum Van Duzee, 1928
Rhaphium caliginosum Meigen, 1824
Rhaphium campestre Curran, 1924
Rhaphium canadense Curran, 1924
Rhaphium canniccii Grootaert, Taylor & Guenard, 2019[8]
Rhaphium caucasicum Negrobov, Grichanov & Selivanova, 2013[9]
Rhaphium caudatum Van Duzee in Curran, 1926
Rhaphium ciliatum Curran, 1929
Rhaphium coloradense Curran, 1926
Rhaphium colute Harmston & James, 1942
Rhaphium commune (Meigen, 1824)
Rhaphium confine Zetterstedt, 1843
Rhaphium crassipes (Meigen, 1824)
Rhaphium crinitum Negrobov & Onishchenko, 1991
Rhaphium cupreum Macquart, 1827
Rhaphium currani (Parent, 1939)
Rhaphium curvitarsus Negrobov, Maslova & Selivanova, 2020[10]
Rhaphium daqinggouense Tang, Wang & Yang, 2016[4]
Rhaphium dichromum Negrobov, 1976
Rhaphium dilatatum Wiedemann, 1830
Rhaphium discigerum Stenhammar, 1850
Rhaphium discolor Zetterstedt, 1838
Rhaphium dispar Coquillett, 1898
Rhaphium doroninae Grichanov, 1995
Rhaphium doroteum Negrobov, 1979
Rhaphium dorsiseta Tang, Wang & Yang, 2016[4]
Rhaphium dubium (Van Duzee, 1922)
Rhaphium eburnea (Parent, 1926)[8]
Rhaphium effilatum (Wheeler, 1899)
Rhaphium elegantulum (Meigen, 1824)
Rhaphium elongatum Van Duzee, 1933
Rhaphium ensicorne Meigen, 1824
Rhaphium essoense Negrobov, 1979
Rhaphium exile Curran, 1926
Rhaphium fasciatum Meigen, 1824
Rhaphium fascipes (Meigen, 1824)
Rhaphium femineum (Van Duzee, 1922)
Rhaphium femoratum (Van Duzee, 1922)
Rhaphium firsovi Stackelberg & Negrobov in Negrobov, 1976
Rhaphium fissum Loew, 1850
Rhaphium flavicoxa (Van Duzee, 1922)
Rhaphium flavilabre Negrobov, 1979
Rhaphium foliatum Curran, 1926
Rhaphium fulvipes (Macquart, 1827)
Rhaphium furcatum Yang & Saigusa, 2000[11]
Rhaphium furciferum Curran, 1931
Rhaphium gangchanum Qilemoge, Wang & Yang, 2019[2]
Rhaphium gansuanum Yang, 1998[5]
Rhaphium gibsoni Curran, 1926
Rhaphium glaciale (Ringdahl, 1920)
Rhaphium gracile Curran, 1924
Rhaphium grandicercum Negrobov, 1979
Rhaphium grande (Curran, 1923)
Rhaphium gravipes Haliday in Walker, Stainton & Wilkinson, 1851
Rhaphium gruniniani Negrobov, 1979
Rhaphium gussakovskii Stackelberg & Negrobov in Negrobov, 1976
Rhaphium heilongjiangense Wang, Yang & Masunaga, 2005
Rhaphium hirtimanum Van Duzee, 1933
Rhaphium holmgreni (Mik, 1878)
Rhaphium hongkongense Grootaert, Taylor & Guenard, 2019[8]
Rhaphium hungaricum (Becker, 1918)
Rhaphium huzhuense Qilemoge, Lin, Qi, Li & Yang, 2020[12]
Rhaphium impetuum Curran, 1926
Rhaphium insolitum Curran, 1926
Rhaphium intermedium (Becker, 1918)
Rhaphium jamalense Negrobov, 1986
Rhaphium johnrichardi Negrobov & Grichanov, 2010[13]
Rhaphium johnsoni (Van Duzee, 1923)
Rhaphium lanceolatum Loew, 1850
Rhaphium latifacies Van Duzee, 1930
Rhaphium latimanum Kahanpaa, 2007
Rhaphium lehri Negrobov, 1977
Rhaphium longibara Van Duzee, 1930
Rhaphium longicorne (Fallen, 1823)
Rhaphium longipalpe Curran, 1926
Rhaphium longipes (Loew, 1864)
Rhaphium lugubre Loew, 1861
Rhaphium lumbricum Wei, 2006
Rhaphium macalpini Negrobov, 1986
Rhaphium macrocerum Meigen, 1824
Rhaphium maculipes (Meigen, 1824)
Rhaphium magnicorne (Zetterstedt, 1843)
Rhaphium mcveighi Grichanov, 1995
Rhaphium mediocre (Becker, 1922)
Rhaphium melampus (Loew, 1861)
Rhaphium mesasiaticum Negrobov, Selivanova & Maslova, 2013[14]
Rhaphium micans (Meigen, 1824)
Rhaphium minhense Qilemoge, Lin, Qi, Li & Yang, 2020[12]
Rhaphium monotrichum Loew, 1850
Rhaphium montanum (Van Duzee, 1920)
Rhaphium nasutum (Fallen, 1823)
Rhaphium neimengense Tang, Wang & Yang, 2016[4]
Rhaphium neolatifacies Yang & Wang, 2006
Rhaphium nigrum (Van Duzee, 1923)
Rhaphium nigribarbatum (Becker, 1900)
Rhaphium nigricoxa (Loew, 1861)
Rhaphium nigrociliatum Curran, 1926
Rhaphium nigrovittatum Curran, 1926
Rhaphium nubilum Van Duzee, 1930
Rhaphium nudiusculum Negrobov, 1976
Rhaphium nuortevai Negrobov, 1977
Rhaphium obscuripes Zetterstedt, 1849
Rhaphium obtusum Van Duzee, 1928
Rhaphium occipitale Curran, 1927
Rhaphium orientale Curran, 1926
Rhaphium ovsyannikovae Grichanov, 1995
Rhaphium palliaristatum Yang & Saigusa, 2001[15]
Rhaphium palpale Curran, 1926
Rhaphium parentianum Negrobov, 1979
Rhaphium patellitarse (Becker, 1900)
Rhaphium patulum (Raddatz, 1873)
Rhaphium paulseni Philippi, 1865
Rhaphium pectinatum (Loew, 1859)
Rhaphium pectinigerum (Parent, 1938)[16]
Rhaphium penicillatum Loew, 1850
Rhaphium petchi Curran, 1924
Rhaphium picketti Grichanov, 1995
Rhaphium pitkini Grichanov, 1995
Rhaphium pollex (Van Duzee, 1922)
Rhaphium psilopodum (Becker, 1918)
Rhaphium punctitarse Curran, 1924
Rhaphium qinghaiense Yang, 1998[17]
Rhaphium quadrispinosum (Strobl, 1898)
Rhaphium reaveyi Grichanov, 1995
Rhaphium relatum (Becker, 1922)
Rhaphium rhaphioides Zetterstedt, 1838
Rhaphium richterae Negrobov, 1977
Rhaphium riparium (Meigen, 1824)
Rhaphium rivale (Loew, 1869)
Rhaphium robustum Curran, 1926
Rhaphium rossi Harmston & Knowlton, 1940
Rhaphium rotundiceps (Loew, 1861)
Rhaphium sachalinense Negrobov, 1979
Rhaphium septentrionale Curran, 1931
Rhaphium sexsetosum (Vanschuytbroeck, 1951)
Rhaphium shaliuhense Qilemoge, Wang & Yang, 2019[2]
Rhaphium shamshevi Grichanov, 1995
Rhaphium shannoni Curran, 1926
Rhaphium sibiricum Negrobov, Barkalov & Selivanova, 2011[18]
Rhaphium sichotense Negrobov, 1979
Rhaphium sichuanense Yang & Saigusa, 1999[3]
Rhaphium signifer (Osten Sacken, 1878)
Rhaphium simplicipes Curran, 1926
Rhaphium sinense Negrobov, 1979
Rhaphium speciosum (S. Abreu, 1929)
Rhaphium spinitarse Curran, 1924
Rhaphium spinulatum Grootaert, Taylor & Guenard, 2019[8]
Rhaphium srilankense Naglis & Grootaert, 2011
Rhaphium stackelbergi Negrobov, 1976
Rhaphium steyskali Robinson, 1967[19]
Rhaphium suave (Loew, 1859)
Rhaphium subarmatum Curran, 1924
Rhaphium subfurcatum Van Duzee, 1932
Rhaphium subtridactylum Negrobov, Barkalov & Selivanova, 2011
Rhaphium temerarium (Becker, 1922)
Rhaphium terminale (Van Duzee, 1924)
Rhaphium thoracicum (Meigen, 1824)
Rhaphium tianshanicum Negrobov, Grichanov & Selivanova, 2013[9]
Rhaphium tianshuiense Qilemoge, Wang & Yang, 2019[2]
Rhaphium tibiale (von Roser, 1840)
Rhaphium triangulatum (Van Duzee, 1922)
Rhaphium tricaudatum (Van Duzee, 1923)
Rhaphium tridactylum (Frey, 1915)
Rhaphium tripartitum (Frey, 1913)
Rhaphium tuberculatum (Negrobov, 1973)
Rhaphium turanicola (Stackelberg, 1927)
Rhaphium umbripenne (Frey, 1915)
Rhaphium vanduzeei Curran, 1926
Rhaphium venustum Negrobov, 1977
Rhaphium vockerothi Robinson, 1964
Rhaphium wheeleri Van Duzee, 1932
Rhaphium wuduanum Wang, Yang & Masunaga, 2005
Rhaphium xinjiangense Yang, 1998[5]
Rhaphium xipheres (Wheeler, 1899)
Rhaphium xiphias Meigen, 1824
Rhaphium zairense Negrobov, Grichanov & Bakary, 1982
Rhaphium zakonnikovae Grichanov, 1995
Rhaphium zhongdianum Yang & Saigusa, 2001[20]

References

O'Hara, James E.; Cerretti, Pierfilippo; Pape, Thomas; Evenhuis, Neal L. (2011). "Nomenclatural Studies Toward a World List of Diptera Genus-Group Names. Part II: Camillo Rondani" (PDF). Zootaxa. 3141: 1–268. doi:10.11646/zootaxa.3141.1.1.
Qilemoge, M.; Wang; Yang, D. (2019). "Three new species of Rhaphium from China, with an updated key to Chinese Rhaphium (Diptera, Dolichopodidae, Rhaphiinae)". ZooKeys (840): 87–99. doi:10.3897/zookeys.840.31602. PMC 6482115. PMID 31065228.
Yang, D.; Saigusa, T. (1999). "New and little known species of Dolichopodidae from China (VI): Diptera from Emei Mountain (1)" (PDF). Bulletin de l'Institut royal des sciences naturelles de Belgique. Entomologie. 69: 233–250.
Tang, C.; Wang, N.; Yang, D. (2016). "Rhaphium (Diptera: Dolichopodidae: Rhaphiinae) from China with six new species". Zootaxa. 4162 (3): 581–593. doi:10.11646/zootaxa.4162.3.11. PMID 27615993.
Yang, D. (1998). "New and little known species of Dolichopodidae from China (I)" (PDF). Bulletin de l'Institut royal des sciences naturelles de Belgique. Entomologie. 68: 151–164.
Negrobov, O. P.; Barkalov, A. B.; Selivanova, O. B. (2012). "Rhaphium Meigen (Diptera, Dolichopodidae) from the Taimyr Peninsula (Russia), with description of a new species" (PDF). Zootaxa. 3548: 75–87. doi:10.11646/zootaxa.3548.1.6.
Meuffels, H. J. G.; Grootaert, Patrick (1999). "New names in the family Dolichopodidae (Diptera)" (PDF). Bulletin de l'Institut royal des Sciences naturelles de Belgique. Entomologie. 69: 289–292. Retrieved 30 March 2018.
Grootaert, P.; Taylor, C.; Guénard, B. (2019). "Three new species of Rhaphium Meigen, 1803 from mangroves in Hong Kong (Diptera: Dolichopodidae: Rhaphiinae)". European Journal of Taxonomy (540): 1–21. doi:10.5852/ejt.2019.540.
Negrobov, O.P.; Grichanov, I.Ya.; Selivanova, O.V. (2013). "Palearctic species of the Rhaphium albifrons group (Diptera, Dolichopodidae)" (PDF). Euroasian Entomological Journal. 12 (6): 601–606.
Negrobov, O. P.; Maslova, O. O.; Selivanova, O. V. (2020). "A new species of the genus Rhaphium (Diptera: Dolichopodidae) from Sakhalin Island". Far Eastern Entomologist. 409: 21–24. doi:10.25221/fee.409.3.
Yang, D.; Saigusa, T. (2000). "New and little known species of Dolichopodidae from China (VII): Diptera from Emei Mountain (2)" (PDF). Bulletin de l'Institut royal des Sciences naturelles de Belgique. Entomologie. 70: 219–242.
Qilemoge; Lin, Chen; Qi, Peifeng; Li, Deming; Yang, Ding (2020). "Two new species of Rhaphium from Qinghai Province, China (Diptera, Dolichopodidae, Rhaphiinae)". ZooKeys (931): 73–94. doi:10.3897/zookeys.931.49671. PMC 7205839. PMID 32405242.
Negrobov, O.P.; Grichanov, I.Ya. (2010). "The Rhaphium crassipes species group in the Palearctic Region with the description of a new species from Uzbekistan (Diptera: Dolichopodidae)" (PDF). Caucasian Entomological Bulletin. 6 (1): 117–122. doi:10.23885/1814-3326-2010-6-1-117-122.
Negrobov, O.P.; Selivanova, O.V.; Maslova, O.O. (2013). "Новый вид рода Rhaphium Meigen, 1803 (Dolichopodidae, Diptera) из Средней Азии" [New Species Rhaphium Meigen, 1803 (Dolichopodidae, Diptera) from Central Asia] (PDF). Byulleten Moskovskogo Obschestva Ispytatelei Prirody. Otdel Biologicheskii [Bulletin of Moscow Society of Naturalists, Biological Series] (in Russian). 118 (6): 22–26.
Yang, D.; Saigusa, T. (2001). "New and little known species of Dolichopodidae (Diptera) from China (XI)" (PDF). Bulletin de l'Institut royal des Sciences naturelles de Belgique. Entomologie. 71: 237–256.
Grichanov, I. Ya. (2001). "Afrotropical Syntormon Loew and new synonyms in the genus Rhaphium Loew (Diptera: Dolichopodidae)". International Journal of Dipterological Research. 12 (4): 181–194.
Yang, D. (1998). "New and little known species of Dolichopodidae from China (III)" (PDF). Bulletin de l'Institut royal des Sciences naturelles de Belgique. Entomologie. 68: 177–183.
Negrobov, O. P.; Barkalov, A. V.; Selivanova, O. V. (2011). "A New Species of the Genus Rhaphium (Dolichopodidae, Diptera) from Siberia" (PDF). Vestnik Zoologii. 45 (3): e-42–e-44. doi:10.2478/v10058-011-0018-1.
Robinson, H. (1967). "New species of Dolichopodidae from the United States and Mexico (Diptera)". Proceedings of the Entomological Society of Washington. 69: 114–127.

Yang, D.; Saigusa, T. (2001). "New and little known species of Dolichopodidae (Diptera) from China (IX)" (PDF). Bulletin de l'Institut royal des Sciences naturelles de Belgique. Entomologie. 71: 165–188.

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