Pammene

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Superfamilia: Tortricoidea

Familia: Tortricidae
Subfamilia: Olethreutinae
Tribus: Grapholitini
Genus: Pammene
Species: P. aceris – P. adusta – P. agnotana – P. ainorum – P. albuginana – P. amygdalana – P. argyrana – P. aurana – P. aurita – P. avetianae – P. bathysema – P. blanda – P. blockian – P. bowmanana – P. caeruleata – P. caliginosa – P. castanicola – P. casticola – P. christophana – P. clanculana – P. cocciferana – P. crataegicola – P. crataegophila – P. cyanatra – P. cytisana – P. engadinensis – P. epanthista – P. exscribana – P. fasciana – P. felicitana – P. festiva – P. flavicellula – P. fulminea – P. gallicana – P. gallicolana – P. germmana – P. giganteana – P. ginkgoicola – P. griseana – P. griseomaculana – P. grunini – P. hilarocrossa – P. ignorata – P. insolentana – P. instructana – P. insulana – P. japonica – P. juniperana – P. laserpitiana – P. leucitis – P. luculentana – P. luedersiana – P. macrolepis – P. mariana – P. medioalbana – P. megalocephala – P. monotincta – P. nannodes – P. nemorosa – P. nequior – P. nescia – P. nigritana – P. obscurana – P. ochsenheimeriana – P. ocliferia – P. oreina – P. orientana – P. ornata – P. oxycedrana – P. paula – P. percognata – P. perstructana – P. phthoneris – P. piceae – P. populana – P. pulchella – P. pullana – P. purpureana – P. querceti – P. regiana – P. reisseri – P. rhediella – P. salvana – P. shicotanica – P. snellenana – P. soyoensis – P. spiniana – P. splendidulana – P. subsalvana – P. suspectana – P. tauriana – P. trauniana – P. tricuneana – P. tsugae – P. vernana
Name

Pammene Hübner, 1816

Type species: Tortrix trauniana Denis & Schiffermüller, 1775

Pammene aurana , Belgian High Ardennes .

References

Bae, Y.-S. & Park, K.-T. 1998. The Genus: Pammene Hubner (Lepidoptera: Tortricidae) from Korea, with Descriptions of Two New Species. Zoological Studies 37(4): 291–301. Full article (pdf). Reference page.
Brown, J.W. 2005. World Catalogue of Insects. Volume 5: Tortricidae (Lepidoptera) 1–741. Apollo Books. Reference page.
Clausi, M., Leone, D., Oreste, M. & Tarasco, E. 2016. Comments on the morphology and biology of Pammene castanicola Trematerra & Clausi, 2009 (Lepidoptera: Tortricidae: Grapholitini). Zootaxa 4168(3): 570–572. DOI: 10.11646/zootaxa.4168.3.8. Reference page.
Diakonoff, A., 1976: New and rare Mediterranean Laspeyresiini collected by Dr. J. Klimesch (Lepidoptera, Tortricidae). Zoologische Mededelingen, 50 (6): 91–105. Full article: [1].
Hübner, 1816, Verz. bekannter Schmett. 378
Huemer, P. & S. Erlebach, 1999: Pammene laserpitiana sp.n., ein neuer Endemit der Südalpen Lepidoptera: Tortricidae). Qudrifina 2: 231–240. Full article: [2].
Liu, Y.-q., 1992: Two new species of Laspeyresiini damaged cones and seeds of forest.(Lepidoptera: Tortricidae). Sinozoologia 9: 249–252. Full article: [3].
Liu, Y.-q. & F. Komai, 1993: A new species of the genus Pammene (Lepidoptera: Tortricidae) injurious to fruits of hawthorn in China. Tinea 13 (23): 241–244. Full article: [4].
Trematerra, P. & M. Clausi, 2009: Pammene casticola sp. n. a new Tortricidae from Sicily. Boll. Zool. Agric. Bachic (II) 41 (2): 123–128.

The tortrix moth, genus Pammene, belongs to the tribe Grapholitini of the subfamily Olethreutinae.[1]
Species

89 species are currently recognized in Pammene:[2]

Pammene aceris Kuznetzov in Danilevsky & Kuznetsov, 1968
Pammene adusta Kuznetzov 1972
Pammene agnotana Rebel, 1914
Pammene ainorum Kuznetzov in Danilevsky & Kuznetsov, 1968
Pammene albuginana (Guenée, 1845)
Pammene amygdalana (Duponchel in Godart, 1842)
Pammene argyrana (Hübner, [1799])
Pammene aurana (Fabricius, 1775)
Pammene aurita Razowski, 1992
Pammene avetianae Kuznetzov, 1964
Pammene bathysema Diakonoff, 1976
Pammene blanda Bae & Park, 1998
Pammene blockiana (Herrich-Schäffer, 1851)
Pammene bowmanana (McDunnough, 1927)
Pammene caeruleata Kuznetzov, 1970
Pammene caliginosa Kuznetzov, 1972
Pammene christophana (Moschler, 1862)
Pammene clanculana (Tengström, 1869)
Pammene cocciferana Walsingham, 1903
Pammene crataegicola Liu & Komai, 1993
Pammene crataegophila Amsel, 1935
Pammene cyanatra (Diakonoff, 1976)
Pammene cytisana (Zeller, 1847)
Pammene engadinensis Müller-Rutz, 1920
Pammene epanthista (Meyrick, 1922)
Pammene exscribana Kuznetzov, 1986
Pammene fasciana (Linnaeus, 1761) – chestnut leafroller
Pammene felicitana Heinrich, 1923
Pammene festiva Kuznetzov, 1972
Pammene flavicellula Kuznetzov, 1971
Pammene fulminea (Komai, 1999)
Pammene gallicana (Guenée, 1845)
Pammene gallicolana (Lienig & Zeller, 1846)
Pammene germmana (Hübner, [1799])
Pammene giganteana (Peyerimhoff, 1863)
Pammene ginkgoicola Liu, 1992
Pammene griseana Walsingham, 1900
Pammene griseomaculana Kuznetzov, 1960
Pammene grunini (Kuznetzov, 1960)
Pammene ignorata Kuznetzov, in Danilevsky & Kuznetsov, 1968
Pammene insolentana Kuznetzov, 1964
Pammene instructana Kuznetzov, 1964
Pammene insulana (Guenée, 1845)
Pammene japonica Kuznetzov in Danilevsky & Kuznetsov, 1968
Pammene juniperana (Millière, 1858)
Pammene laserpitiana Huemer & Erlebach, 1999
Pammene leucitis (Meyrick, 1907)
Pammene luculentana Kuznetzov in Danilevsky, Kuznetsov & Falkovitsh, 1962
Pammene luedersiana (Sorhagen, 1885)
Pammene macrolepis Diakonoff, 1976
Pammene mariana (Zerny, 1920)
Pammene medioalbana Knudson, 1986
Pammene megalocephala Diakonoff, 1983
Pammene monotincta Kuznetzov, 1976
Pammene nannodes Walsingham, 1900
Pammene nemorosa Kuznetzov in Danilevsky & Kuznetsov, 1968
Pammene nequior Kuznetzov, 1964
Pammene nescia Kuznetzov, 1972
Pammene nigritana (Mann, 1862)
Pammene obscurana (Stephens, 1834)
Pammene ochsenheimeriana (Lienig & Zeller, 1846)
Pammene ocliferia (Heinrich, 1926)
Pammene oreina Kuznetzov, 1973
Pammene orientana Kuznetzov, 1960
Pammene ornata Walsingham, 1903
Pammene oxycedrana (Millière, 1876)
Pammene paula (Heinrich, 1926)
Pammene percognata Diakonoff, 1976
Pammene perstructana (Walker, 1863)
Pammene phthoneris Diakonoff, 1976
Pammene piceae Komai, 1999
Pammene populana (Fabricius, 1787)
Pammene pulchella Amsel, 1935
Pammene pullana Kuznetzov, 1986
Pammene purpureana (Constant, 1888)
Pammene querceti (Gozmány, 1957)
Pammene regiana (Zeller, 1849)
Pammene rhediella (Clerck, 1759)
Pammene salvana (Staudinger, 1859)
Pammene shicotanica Kuznetzov in Danilevsky & Kuznetsov, 1968
Pammene soyoensis Bae & Park, 1998
Pammene spiniana (Duponchel in Godart, 1842)
Pammene splendidulana (Guenée, 1845)
Pammene subsalvana Kuznetzov, 1960
Pammene suspectana (Lienig & Zeller, 1846)
Pammene tauriana Kuznetzov, 1960
Pammene trauniana (Denis & Schiffermüller, 1775)
Pammene tricuneana (Kennel, 1900)
Pammene tsugae Issiki in Issiki & Mutuura, 1961

Synonyms

Obsolete scientific names (junior synonyms and others) of Pammene are:[3]

Encelis (lapsus)
Eucelis Hübner, [1825]
Encells (lapsus)
Halonota Stephens, 1852
Hemene (lapsus)
Hemerosia Stephens, 1852
Hemimene Hübner, 1825
Heusimene Stephens, 1834
Metasphaeroeca Fernald, 1908

Orchemia Guenée, 1845
Palla Billberg, 1820 (non Hübner, 1819: preoccupied)
Pamene (lapsus)
Pammena (lapsus)
Phthoroblastis Lederer, 1859
Psudotomia Stephens, 1829
Pyrodes Guenée, 1845 (non Audinet-Serville, 1832: preoccupied)
Sphaeroeca Meyrick, 1895 (non Lauterborn, 1894: preoccupied)
Trycheris Guenée, 1845

The synonymy of Pammene has been subject to some confusion with its close relatives Cydia and Epinotia: Eucelis, Orchemia and Trycheris are sometimes[4] listed as junior synonyms of the former, and Halonota has been listed[5] as synonym of the latter. But the type species of Eucelis and Trycheris is Tortrix mediana (a junior synonym of P. aurana), that of Orchemia is O. gallicana (a junior synonym of P. gallicana), and that of Halonota is Pyralis populana (a junior synonym of P. populana). Thus, Eucelis, Halonota, Orchemia and Trycheris are all junior subjective synonym of the present genus.[1]
Footnotes

Baixeras et al. (2009a)
Baixeras et al. (2009b)
Baixeras et al. (2009a), and see references in Savela (2005)
See e.g. references in Savela (2005)

See e.g. references in Savela (2009)

References
Wikimedia Commons has media related to Pammene.

Baixeras, J.; Brown, J.W. & Gilligan, T.M. (20091): Online World Catalogue of the Tortricidae – Genus: Pammene account. Version 1.3.1. Retrieved 2009-APR-20.
Baixeras, J.; Brown, J.W. & Gilligan, T.M. (2009b): Online World Catalogue of the Tortricidae – Pammene species list. Version 1.3.1. Retrieved 2009-JAN-20.
Savela, Markku (2005): Markku Savela's Lepidoptera and some other life forms – Pammene. Version of 2005-SEP-13. Retrieved 2010-APR-20.
Savela, Markku (2009): Markku Savela's Lepidoptera and some other life forms – Epinotia. Version of 2009-JUL-07. Retrieved 2010-APR-19.