Eudaminae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Hesperiidae
Subfamilia: Eudaminae
Tribus (4): Entheini – Eudamini – Oileidini – Phocidini
Name

Eudaminae Mabille, 1877: 162.

Included Genera: Astraptes – Aguna – Augiades – Autochton –Cecropterus – Cephise – Chioides –Codatractus –Drephalys –Ectomis –Entheus –Epargyreus –Hyalothyrus –Lobocla –Lobotractus –Narcosius –Phanus –Polygonus –Proteides –Ridens –Spathilepia –Spicauda –Tarsoctenus –Telegonus –Telemiades –Udranomia –Urbanus –Venada –Zestusa –Zeutus

References

Bell, E.L. 1947. A New Genus and some New Species and Subspecies of Neotropical Hesperiidae (Lepidoptera, Rhopalocera). American Museum novitates 1354: 1–12. PDF. Reference page.
Cong, Q., Zhang, J., Shen, J.H. & Grishin, N.V. 2019. Fifty new genera of Hesperiidae (Lepidoptera). Insecta Mundi 0731: 1–56. Zoobank. Reference page.
Lamas, G. 2004. (ed.) Checklist: Part 4A. Hesperioidea - Papilionoidea. In Heppner, J.B. (ed.) Atlas of Neotropical Lepidoptera. Vol.5A, Pt.4A. Assn. for Tropical Lepidoptera/Scientific Publishers, Gainesville. 439pp. Reference page.
Lee, C.L. 1962. Some new species of Rhopalocera from China. II. Acta entomologica sinica 11(2): 139–148. PDF Reference page.
Li, W., Cong. Q., Shen, J.H., Zhang, J., Hallwachs, W., Janzen, D.H. & Grishin, N.V. 2019. Genomes of skipper butterflies reveal extensive convergence of wing patterns. PNAS 116(13): 6232–6237. DOI: 10.1073/pnas.1821304116. Text. SI Appendix. Reference page.
Mabille, P. 1877. Descriptions de Lépidoptères nouveaux du groupe des Hespérides. Petites Nouvelles Entomologiques 2(167): 114; 2(179): 161–162. Reference page.
Sahoo, R.K., Warren, A.D., Wahlberg, N., Brower, A.V.Z., Lukhtanov, V.A. & Kodandaramaiah, U. 2016. Ten genes and two topologies: an exploration of higher relationships in skipper butterflies (Hesperiidae). PeerJ 4: e2653. DOI: 10.7717/peerj.2653 Reference page.
Steinhauser, S.R., 1989. Taxonomic notes and descriptions of new taxa in the neotropical Hesperiidae. Part I. Pyrginae. Bulletin of the Allyn Museum 127: 1–70. Reference page.
Toussaint, E.F.A., Breinholt, J.W., Earl, C., Warren, A.D., Brower, A.V.Z., Yago, M., Dexter, K.M., Espeland, M., Pierce, N.E., Lohman, D.J. & Kawahara, A.Y. 2018. Anchored phylogenomics illuminates the skipper butterfly tree of life. BMC Evolutionary Biology 18: 101. DOI: 10.1186/s12862-018-1216-z Reference page.
Warren, A.D., Ogawa, J.R. & Brower, A.V.Z. 2009. Revised classification of the family Hesperiidae (Lepidoptera: Hesperioidea) based on combined molecular and morphological data. Systematic entomology 34(3): 467–523. Reference page.
Zhang, J., Cong. Q., Shen, J.H., Brockmann, E. & Grishin, N.V. 2019. Three new subfamilies of skipper butterflies (Lepidoptera, Hesperiidae). ZooKeys 861: 91–105. DOI: 10.3897/zookeys.861.34686 Reference page.

The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus Eudamus is today a junior synonym of Urbanus. They are largely found in the Neotropics, with some extending into temperate North America and the puzzling Lobocla (which seems closest to Zestusa) occurring in East Asia.[1]
Typical resting positions

They were formerly included in subfamily Pyrginae (spread-winged skippers) as a tribe Eudamini. This was mainly due to a perceived similarity with the pyrgine tribes Celaenorrhini and Pyrgini, where certain eudamine genera were indeed sometimes placed. But in reality, these genera seem to be quite closely related to Dyscophellus, and the two lineages of moderately advanced skippers are quite distinct.[1]
Genera

The following 55 genera are placed in the Eudaminae in their delimitation as distinct subfamily. In phylogenetic sequence, they are:[1]

Chioides
Proteides
Spathilepia
Thorybes
Achalarus
Autochton
Urbanus (paraphyletic?)
Astraptes
Narcosius
Cabares
Aguna
Codatractus
Typhedanus
Lobocla (formerly in Celaenorrhini)
Zestusa
Calliades
Drephalys
Udranomia
Hyalothyrus
Augiades
Cabirus
Entheus
Phanus
Phareas
Tarsoctenus
Bungalotis
Nicephellus
Salatis
Porphyrogenes
Oileides
Aurina
Sarmientola Berg, 1897
Polygonus
Telemiades (formerly in Pyrgini)
Phocides
Hypocryptothrix
Nascus
Pseudonascus
Dyscophellus
Euriphellus
Cogia (formerly in Pyrgini)
Paracogia (formerly in Pyrgini)
Marela (formerly in Pyrgini)
Oechydrus (formerly in Pyrgini)
Nerula (formerly in Pyrgini)
Venada

Some genera are of unresolved relationships. Presumably, at least most of them are indeed Eudamini, but it cannot be ruled out that some belong elsewhere in the Hesperiidae:[1]

Cephise
Chrysoplectrum
Ectomis (formerly in Pyrgini)
Epargyreus
Heronia
Polythrix
Ridens
Thessia

Footnotes

Brower & Warren (2009)

References
Wikispecies has information related to Eudaminae.
Wikimedia Commons has media related to Eudaminae.

Brower, Andrew V.Z. & Warren, Andrew (2009): Tree of Life Web Project – Eudaminae. Version of 2009-JUN-11. Retrieved 2009-DEC-24.