Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Heterobranchia
Infraclassis: Euthyneura
Cohors: Tectipleura
Subcohors: Panpulmonata
SuperOrdo: Eupulmonata
Ordo: Stylommatophora
SubOrdo: Helicina
InfraOrdo: Limacoidei
Superfamilia: Helicarionoidea
Familia: Urocyclidae
Subfamiliae (3): Rhysotininae – Sheldoniinae – Urocyclinae
Genera Incertae sedis (1 + 1†): Selatodryas – †Koruella
Overview of genera (71 + 1†)
Extant groups (71): Acantharion – Africarion – Amatarion – Amphiblema – Angustivestis – Anisotoxon – Aspidelus – Aspidotomium – Atoxon – Atoxonoides – Atrichotoxon – Belonarion – Buettneria – Bukobia – Camerunarion – Carinazingis – Centrafricarion – Chlamydarion – Colparion – Degneria – Dendrolimax – Elisolimax – Emphysetes – Entagaricus – Estria – Falloonella – Fuellebornia – Granularion – Guillainia – Gymnarion – Kerkophorus – Lacrimarion – Leptichnoides – Leptichnus – Malagarion – Mesafricarion – Microcyclus – Microkerkus – Montanobloyetia – Morrumbalia – Nupnus – Paragranularion – Pembatoxon – Percivalia – Phaneroporus – Plicatonanina – Polytoxon – Pseudatoxon – Ptilototheca – Rhopalogonium – Rhysotina – Selatodryas – Senegalarion – Sheldonia – Sjostedtina – Sylvarion – Tanzalimax – Thapsia – Thielarion – Tresia – Trichotoxon – Trochonanina – Trochozonites – Tropidocochlion – Udzungwalimax – Upembella – Urocyclus – Verdcourtia – Verrucarion – Zingis – Zonitarion
Fossil groups (1): †Koruella
Name
Urocyclidae Simroth, 1888
References
Template:Simroth, 1888
Rowson, B., Paustian, M. & van Goethem, J. 2017. New species and records of terrestrial slugs from East Africa (Gastropoda, Urocyclidae, Veronicellidae, Agriolimacidae). ZooKeys 723: 11–42. DOI: 10.3897/zookeys.723.21817. Reference page.
Links
Urocyclidae in the World Register of Marine Species
Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea, (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[2]
Some species within this family make and use love darts before mating.[citation needed]
Contents
1 Distribution
2 Taxonomy
3 Genera
4 Cladogram
5 References
6 External links
Distribution
The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.[3][4]
Taxonomy
The family Urocyclidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005),[2] that follows classification of Urocyclinae by Van Goethem (1977):[5]
subfamily Rhysotininae Schileyko, 2002
subfamily Sheldoniinae Connolly, 1925 (1912)[6] - synonyms: Peltatinae Godwin-Austen, 1912;[7] Trochonanininae Connolly, 1912;[8] Trochozonitinae Iredale, 1914;[9] Ledoulxiinae Pilsbry, 1919;[10] Gymnarionidae Von Mol, 1970;[11] Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002
subfamily Urocyclinae Simroth, 1889
tribe Urocyclini Simroth, 1889 - synonyms: Atoxonini Schileyko, 2002; Buettneriini Schileyko, 2002
tribe Dendrolimacini Van Goethem, 1977[5]
tribe Leptichnini Van Goethem, 1977[5]
tribe Upembellini Van Goethem, 1977[5]
Genera
Genera within the family Urocyclidae include:
unsorted
† Koruella Pickford, 2009
Subfamily Rhysotininae Schileyko, 2002
Rhysotina Ancey, 1887[2]
Thomeonanina Germain, 1909 : synonym of Rhysotina Ancey, 1887 (objective synonym)
Subfamily Sheldoniinae
Acantharion Binder & Tillier, 1985[2]
Africarion Godwin-Austen, 1883
Amatarion Van Mol, 1970
Angustivestis Pilsbry, 1919
Aspidelus Morelet, 1884
Aspidotomium Degner, 1932
Belonarion Pilsbry, 1919
Bloyetia Bourguignat, 1890
Camerunarion Van Mol, 1970
Carinazingis van Bruggen & de Winter, 1990
Centrafricarion Van Mol, 1970 [11]
Chlamydarion Van Mol, 1968
Colparion Laidlaw, 1938
Degneria Verdcourt, 1956
Entagaricus Pilsbry, 1919
Estria Poirier, 1887
Falloonella Preston, 1914
Fuellebornia Verdcourt, 1998
Granularion Germain, 1912
Gymnarion Pilsbry, 1919[2]
Hamya Bourguignat, 1885 †
Kerkophorus Godwin-Austen, 1912
Lacrimarion Connolly, 1939
Malagarion Tillier, 1979
Mesafricarion Pilsbry, 1919
Microkerkus Godwin-Austen, 1912
Montanobloyetia Verdcourt, 1961
Morrumbalia Verdcourt, 1998
Paragranularion Van Mol, 1970
Percivalia Preston, 1914
Plicatonanina Verdcourt, 1961
Principicochlea D. Holyoak & G. Holyoak, 2020
Principitrochoidea D. Holyoak & G. Holyoak, 2020
Ptilototheca Herbert, 2016
Rhopalogonium Degner, 1932
Selatodryas Herbert, 2017
Senegalarion Van Mol, 1970
Sheldonia Ancey, 1887 - the type genus of the subfamily Sheldoniinae[2][12]
Sjostedtina Verdcourt, 1961
Sylvarion Van Mol, 1970
Thapsia Albers, 1860
Thielarion Van Mol, 1970
Thomithapsia D. Holyoak & G. Holyoak, 2020
Thomitrochoidea D. Holyoak & G. Holyoak, 2020
Tresia Van Goethem, 1975
Trochonanina Mousson, 1869[2]
Trochozonites Pfeffer, 1883[2]
Tropidocochlion Verdcourt, 1998
Verdcourtia Van Mol, 1970
Verrucarion Van Mol, 1970 [13]
Zingis E. von Martens, 1878
Zonitarion Pfeffer, 1883[2]
Ledoulxia Bourguignat, 1885:[2] synonym of Trochonanina Mousson, 1869
Peltatus Godwin-Austen, 1908:[2] synonym of Sheldonia Ancey, 1887
Subfamily Urocyclinae
Tribe Dendrolimacini
Dendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini[2]
Tribe Leptichnini
Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini[2]
Tribe Upembellini
Leptichnoides Van Goethem, 1975
Upembella Van Goethem, 1969 - the type genus of the tribe Upembellini[2]
tribe Urocyclini
Anisotoxon Van Goethem, 1975
Atoxon Simroth, 1888[2]
Atoxonoides Van Goethem, 1973
Atrichotoxon Simroth, 1910
Buettneria Simroth, 1910[14]
Bukobia Simroth, 1896
Elisolimax Cockerell, 1893
Emphysetes Verdcourt, 2003
Microcyclus Simroth, 1896
Nupnus Van Goethem, 1975
Pembatoxon Van Goethem, 1975
Phaneroporus Simroth, 1888
Polytoxon Simroth, 1897
Pseudatoxon Van Goethem, 1975
Tanzalimax Rowson, Paustian & Van Goethem, 2017
Trichotoxon Simroth, 1888
Udzungwalimax Rowson, Paustian & Van Goethem, 2017
Urocyclus Gray, 1864 - the type genus of the family Urocyclidae[2]
Cladogram
The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade:[3]
| limacoid clade |
|
References
Simroth H. (1899). "Beiträge zur Kenntniss der Nacktschnecken". Nova Acta, Kaiserlich Leopoldinish-Carolinische Deutsche Akademie der Naturforscher (= Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum) 54(1): 1-91, plates 1-4. p. 62.
Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.
Herbert, Dai; Kilburn, Dick (2004). Field guide to the land snails and slugs of eastern south Africa (1st ed.). Pietermaritzburg :Natal Museum: Natal Museum. ISBN 0-620-32415-5.
Van Goethem J. (1977). "Révision systématique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)". Musée Royal de l'Afrique Centrale , Annales, Sciences zoologiques 218: 355 pp., 4 plates.
Connolly (1925). Annals and Magazine of Natural History (9)15: 467.
Godwin-Austen H. H. (1912). "A review of South-African Land-Mollusca belonging to the Family Zonitidae". Annals and Magazine of Natural History (8)9: 122-139. Plate 1-7. p. 124.
Connolly (1912). Annals of the South African Museum 11(3): 101.
Iredale (1914). Proceedings of the Malacological Society of London 11(2): 122.
Pilsbry H. A. (1919). Bulletin of the American Museum of Natural History 40: 245.
Van Mol J. J. (1970). "Révision des Urocyclidae (Mollusca, Gastropoda, Pulmonata). Anatomie - systématique - zoogéographie". Annales Musée Royal de l’Afrique Centrale, Sciences Zoologiques 180: 1-231.
Herbert D. G. (2004). Sheldonia puzeyi. 2006 IUCN Red List of Threatened Species. Downloaded on 7 August 2007.
de Winter A. J. (1987). "Verrucarion demeryi, a new semi-slug from Sierra Leone (Mollusca: Pulmonata: Urocyclidae)". Basteria 51(4-6): 87-93. abstract.
Simroth H. (1910). In: Voeltzkow Reise O.-Afr., Wiss. Ergebn. 2(5): 611.
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