Geomitridae

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Heterobranchia
Infraclassis: Euthyneura
Cohors: Tectipleura
Subcohors: Panpulmonata
SuperOrdo: Eupulmonata
Ordo: Stylommatophora
SubOrdo: Helicina
InfraOrdo: Helicoidei
Superfamilia: Helicoidea

Familia: Geomitridae
Subfamiliae (2): Geomitrinae – Helicellinae
Overview of genera (42)

Actinella – Alteniella – Backeljaia – Callina – Candidula – Caseolus – Cernuella – Cochlicella – Craspedaria – Discula – Disculella – Geomitra – Helicella – Helicopsis – Heterostoma – Hystricella – Keraea – Lemniscia – Microxeromagna – Monilearia – Moreletina – Obelus – Orexana – Plentuisa – Ponentina – Pseudocampylaea – Pseudoxerophila – Ripkeniella – Serratorotula – Spirorbula – Trochoidea – Wollastonaria – Xerocrassa – Xerogyra – Xerolenta – Xeroleuca – Xeromunda – Xeroplexa – Xerosecta – Xerotricha – Zarateana – Xeropicta
Name

Geomitridae Boettger, 1909
References
Links

Geomitridae in the World Register of Marine Species

Geomitridae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.[1][2]

Anatomy

The family is characterized by the presence of a free right ommatophoral retractor (passing outside the peni-oviducal angle) and a double stimulatory apparatus.[1] A free right ommatophoral retractor has been linked to adaptation to xeric habitats[3]
Taxonomy

The family Geomitridae is subdivided in the following subfamilies (according to Razkin et al., 2015[1])

• Subfamily Geomitrinae
  • tribe Cochlicellini Schileyko, 1972
  • tribe Geomitrini C. Boettger, 1909 - synonym: Ochthephilinae Zilch, 1960 (n.a.)
  • tribe Ponentinini Razkin et al., 2015
• Subfamily Helicellinae
  • tribe Cernuellini Schileyko, 1991
  • tribe Helicellini Ihering, 1909 - synonym: Jacostidae Pilsbry, 1948 (inv.)
  • tribe Helicopsini H. Nordsieck, 1987
  • tribe Plentuisini Razkin et al., 2015
  • tribe Trochoideini H. Nordsieck, 1987

Genera

Genera with the family Geomitridae include:[1][4]

subfamily Geomitrinae

• tribe Cochlicellini
  • Cochlicella Férussac, 1821
• tribe Geomitrini
  • Actinella R. T. Lowe, 1852^[5]
  • Caseolus R. T. Lowe, 1852^[5]
  • Discula R. T. Lowe, 1852^[5]
  • Disculella Pilsbry, 1895^[5]
  • Hystricella R. T. Lowe 1855^[5]
  • Geomitra Swainson, 1840^[5]
  • Heterostoma W. Hartmann, 1843^[5]
  • Hystricella R. T. Lowe, 1855^[5]
  • Lemniscia R. T. Lowe, 1855^[5]
  • Moreletina de Frias Martins, 2002^[5]
  • Pseudocampylaea L. Pfeiffer, 1877^[5]
  • Serratorotula Groh & Hemmen, 1986^[5]
  • Spirorbula R. T. Lowe, 1852^[5]
• tribe Ponentinini
  • Ponentina P. Hesse, 1921

subfamily Helicellinae

• tribe Cernuellini
  • Candidula Kobelt, 1871
  • Cernuella Schlüter, 1838
  • Cernuellopsis Manganelli & Giusti, 1988
  • Xerogyra Monterosato, 1892
  • Xerosecta Monterosato, 1892
• tribe Helicellini
  • Backeljaia Chueca et al., 2018
  • Helicella Férussac, 1821
  • Helicopsis Fitzinger, 1833
  • Microxeromagna Ortiz De Zárate López, 1950
  • Orexana Chueca et al., 2018
  • Xeroleuca Kobelt, 1877
  • Xeroplexa Monterosato, 1892
  • Xerotricha Monterosato, 1892
  • Zarateana Chueca et al., 2018
  • tribe Helicopsini H. Nordsieck, 1987
  • Helicopsis Fitzinger, 1833
  • Pseudoxerophila Westerlund, 1879
  • Xerolenta Monterosato, 1892
  • Xeromunda Monterosato, 1892
  • Xeropicta Monterosato, 1892
• tribe Plentuisini
  • Plentuisa Puente & Prieto, 1992
• tribe Trochoideini
  • Trochoidea T. Brown, 182
  • Xerocrassa Monterosato, 1892

References

Razkin, Oihana; Gómez-Moliner, Benjamín Juán; Prieto, Carlos Enrique; Martínez-Ortí, Alberto; Arrébola, José Ramón; Muñoz, Benito; Chueca, Luis Javier; Madeira, María José (February 2015). "Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution. 83: 99–117. doi:10.1016/j.ympev.2014.11.014. PMID 25485783.
MolluscaBase eds. (2021). MolluscaBase. Geomitridae C.R. Boettger, 1909. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=994707 on 2021-02-01
Giusti, F; Manganelli, G (1987). "Notulae malacologicae, XXXVI. On some Hygromiidae (Gastropoda: Helicoidea) living in Sardinia and in Corsica.(Studies on the Sardinian and Corsican malacofauna VI)". Bollettino Malacologico. 23: 123–206.
Chueca, Luis J.; Gómez-Moliner, Benjamín J.; Madeira, María José; Pfenninger, Markus (January 2018). "Molecular phylogeny of Candidula (Geomitridae) land snails inferred from mitochondrial and nuclear markers reveals the polyphyly of the genus". Molecular Phylogenetics and Evolution. 118: 357–368. doi:10.1016/j.ympev.2017.10.022. ISSN 1055-7903. PMID 29107619.
"Geomitrini". Fauna Europaea, last update 27 January 2011, accessed 4 August 2011.

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