Chersobius

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Cladus: Archelosauria
Division: Pan-Testudines
Division: Testudinata
Ordo: Testudines
Subordo: Cryptodira
Superfamilia: Testudinoidea

Familia: Testudinidae
Genus: Chersobius
Species: C. boulengeri – C. signatus – C. solus
Name

Chersobius (Fitzinger 1835:112)
Type Species: Testudo (Chersobius) signatus Walbaum (= Testudo signata Gmelin, 1789), by subsequent designation by Lindholm (1929:284).

References

Fitzinger, L.J. 1835. Entwurf einer svstematischen Anordnung der Schildkröten nach den Grundsätzen der natürlichen Methode. Ann. Wiener Mus. 1: 103–128.
Hofmeyr, M.D. & Branch, W.R. 2018. The padloper’s tortuous path (Chelonia: Testudinidae): Two genera, not one. African Journal of Herpetology 2018:1-14. [1]

Chersobius is a genus of tiny tortoises in the family Testudinidae, endemic to southern Africa. The genus includes the smallest tortoises in the world. All three species were previously assigned to the genus Homopus.

Naming

As a group, these closely related species are commonly known in Europe and Africa as padlopers (originally meaning "path-walkers" in Afrikaans), due to their habit of making tiny pathways through vegetation.[1][2] In other parts of the world, such as the United States, they are known as Cape tortoises.
Distribution

The genus is indigenous and endemic to southern Africa, one within South Africa, one only in Namibia, and one possibly spanning across the border region of both countries.
Species

The genus contains these species:

Image	Common Name	Scientific name	Distribution
	Karoo padloper or Boulenger's cape tortoise	C. boulengeri	Karoo region[3]
	speckled padloper or speckled tortoise	C. signatus	South African west coast region, the smallest tortoise species in the world
	Nama padloper or Berger's cape tortoise, previously H. bergeri	C. solus	southern Namibia[4]

Conservation and captivity

They are threatened by habitat destruction, traffic on roads, overgrazing, and poaching for the pet trade.[3]

Among the Chersobius species, C. signatus adapts well to captivity, as their diets are not highly specialized.[2] The others do not generally survive well in captivity unless some effort is made to supply them with their natural food, that is, endemic plants from the Cape/Karoo regions.[2] Many are taken from their natural habitat each year, and subsequently die as a result, as they do not readily adapt to typical captive diets and environment change.[2] However, they can be very hardy in captivity, and most problems with captive care are caused by faulty nutrition, high humidity, or bad husbandry.[2]
References

"Homopus Research Foundation, reports". Home.caiway.nl. Retrieved 2017-07-17.
Corton, M., Homopus (Padloper Tortoise) Care, World Chelonian Trust (retrieved August 20, 2013).
"Homopus Research Foundation". Home.caiway.nl. Retrieved 2017-07-17.
"Nama Padloper Tortoise | Namibia". Namibian.org. Retrieved 2017-07-17.