Turdus merula ( w:User:SonNy cZ, Blackbird 2, Size, CC BY-SA 3.0 )
Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis: Sarcopterygii
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Cladus: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Euornithes
Cladus: Ornithuromorpha
Cladus: Ornithurae
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Cladus: Telluraves
Cladus: Australaves
Ordo: Passeriformes
Subordo: Passeri
Infraordo: Passerida
Superfamilia: Muscicapoidea
Familia: Turdidae
Genus: Turdus
Species: T. merula
Subspecies: T. m. aterrimus – T. m. azorensis – T. m. cabrerae – T. m. intermedius – T. m. mauretanicus – T. m. merula – T. m. sowerbyi – T. m. syriacus
Name
Turdus merula Linnaeus, 1758
References
Primary references
Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Tomus I. Holmiæ (Stockholm): impensis direct. Laurentii Salvii. 824 pp. DOI: 10.5962/bhl.title.542 [first availability: page 170] BHL Reference page.
Additional references
Clement, P., Hathway, R., & Wilczur, J. 2000. Thrushes (Helm Identification Guides). Christopher Helm Publishers Ltd. ISBN 0-7136-3940-7.
Collar, N.J. 2005. Family Turdidae (thrushes). In: del Hoyo, J. et al., eds. Handbook of the Birds of the World 10: 514–807. Lynx Edicions, Barcelona.
Nylander, J.A.A; Olsson, U.; Alström, P.; & Sanmartín, I. 2008. Accounting for Phylogenetic Uncertainty in Biogeography: A Bayesian Approach to Dispersal-Vicariance Analysis of the Thrushes (Aves: Turdus) Systematic Biology 57(2): 257–268 DOI: 10.1080/10635150802044003 Open access Reference page.
Vernacular names
Alemannisch: Amsle
aragonés: Tordos
العربية: شحرور
asturianu: Nerbatu
azərbaycanca: Qara qaratoyuq
башҡортса: Ҡара барҡылдаҡ
беларуская (тарашкевіца): Дрозд чорны
беларуская: Дрозд чорны
български: Кос
brezhoneg: Moualc'h zu
bosanski: Kos
català: Merla
нохчийн: Járźa h́oza
کوردی: ڕەشیشە
corsu: Merula
kaszëbsczi: Kòs
čeština: Kos černý
Cymraeg: Mwyalchen
dansk: Solsort
Deutsch: Amsel
dolnoserbski: Kós
Ελληνικά: Κότσυφας
emiliàn e rumagnòl: Mêrel
English: Common Blackbird
Esperanto: Merlo
español: Mirlo
eesti: Musträstas
euskara: Zozo
estremeñu: Mielra
فارسی: توکای سیاه
suomi: Mustarastas
føroyskt: Kvørkveggja
Nordfriisk: Kramper
français: Merle noir
furlan: Mierli
Frysk: Klyster
Gaeilge: Lon dobh
Gàidhlig: Lon dubh
galego: Merlo
ગુજરાતી: કસ્તુરી
Gaelg: Lhondoo
עברית: שחרור
hrvatski: Kos
hornjoserbsce: Kós
magyar: Fekete rigó
հայերեն: Սև կեռնեխ
Bahasa Indonesia: Burung sikatan hitam
Ido: Merlo
íslenska: Svartþröstur
italiano: Merlo, Merlo nero
日本語: クロウタドリ
қазақша: Қара сайрақ
한국어: 대륙검은지빠귀
kurdî: Boqijeya reş
kernowek: Molgh dhu
Latina: Merula
Lëtzebuergesch: Märel
Limburgs: Merel
lietuvių: Juodasis strazdas
latviešu: Melnais mežastrazds
македонски: Обичен кос
മലയാളം: കരിംകിളി
Malti: Malvizz iswed
Mirandés: Mielro, Mierlo, Mierlho
Nedersaksies: Gietelink
Plattdüütsch: Swattdrossel
Nederlands: Merel
norsk nynorsk: Svarttrast
norsk: Svarttrost
Nouormand: Mêle
occitan: Mèrle
polski: Kos zwyczajny
Piemontèis: Merlo
português: Melro-preto
Runa Simi: Kastilla suksu
rumantsch: Merlotscha
română: Mierlă
русский: Чёрный дрозд
sicilianu: Merlu
Scots: Merl
sardu: Merula
davvisámegiella: Čáhppesrásttis
srpskohrvatski / српскохрватски: Kos
slovenčina: Drozd čierny
slovenščina: Kos
shqip: Cërre
српски / srpski: Кос
svenska: Koltrast
Kiswahili: Mkesha mweusi
ไทย: นกแบล็กเบิร์ด
Türkçe: Karatavuk
татарча/tatarça: Кара баркылдык
українська: Дрізд чорний
Tiếng Việt: Hoét đen
West-Vlams: Meirel
walon: Måvi
Zeêuws: Maerelaer
中文: 烏鶇
The common blackbird (Turdus merula) is a species of true thrush. It is also called the Eurasian blackbird (especially in North America, to distinguish it from the unrelated New World blackbirds),[2] or simply the blackbird where this does not lead to confusion with a similar-looking local species. It breeds in Europe, Asiatic Russia, and North Africa, and has been introduced to Australia and New Zealand.[3] It has a number of subspecies across its large range; a few of the Asian subspecies are sometimes considered to be full species. Depending on latitude, the common blackbird may be resident, partially migratory, or fully migratory.
The adult male of the common blackbird (Turdus merula merula, the nominate subspecies), which is found throughout most of Europe, is all black except for a yellow eye-ring and bill and has a rich, melodious song; the adult female and juvenile have mainly dark brown plumage. This species breeds in woods and gardens, building a neat, cup-shaped nest, bound together with mud. It is omnivorous, eating a wide range of insects, earthworms, berries, and fruits.
Both sexes are territorial on the breeding grounds, with distinctive threat displays, but are more gregarious during migration and in wintering areas. Pairs stay in their territory throughout the year where the climate is sufficiently temperate. This common and conspicuous species has given rise to a number of literary and cultural references, frequently related to its song.
Taxonomy and systematics
The common blackbird was described by Carl Linnaeus in his landmark 1758 10th edition of Systema Naturae as Turdus merula (characterised as T. ater, rostro palpebrisque fulvis).[4] The binomial name derives from two Latin words, turdus, "thrush", and merula, "blackbird", the latter giving rise to its French name, merle,[5] and its Scots name, merl.[6]
About 65 species of medium to large thrushes are in the genus Turdus, characterised by rounded heads, longish, pointed wings, and usually melodious songs. Although two European thrushes, the song thrush and mistle thrush, are early offshoots from the Eurasian lineage of Turdus thrushes after they spread north from Africa, the blackbird is descended from ancestors that had colonised the Canary Islands from Africa and subsequently reached Europe from there.[7] It is close in evolutionary terms to the island thrush (T. poliocephalus) of Southeast Asia and islands in the southwest Pacific, which probably diverged from T. merula stock fairly recently.[8]
It may not immediately be clear why the name "blackbird", first recorded in 1486, was applied to this species, but not to one of the various other common black English birds, such as the carrion crow, raven, rook, or jackdaw. However, in Old English, and in modern English up to about the 18th century, "bird" was used only for smaller or young birds, and larger ones such as crows were called "fowl". At that time, the blackbird was therefore the only widespread and conspicuous "black bird" in the British Isles.[9] Until about the 17th century, another name for the species was ouzel, ousel or wosel (from Old English osle, cf. German Amsel). Another variant occurs in Act 3 of Shakespeare's A Midsummer Night's Dream, where Bottom refers to "The Woosell cocke, so blacke of hew, With Orenge-tawny bill". The ouzel usage survived later in poetry, and still occurs as the name of the closely related ring ouzel (Turdus torquatus), and in water ouzel, an alternative name for the unrelated but superficially similar white-throated dipper (Cinclus cinclus) and American dipper (‘’C. mexicanus’’).[10]
Two related Asian Turdus thrushes, the white-collared blackbird (T. albocinctus) and the grey-winged blackbird (T. boulboul), are also named blackbirds,[8] and the Somali thrush (T. (olivaceus) ludoviciae) is alternatively known as the Somali blackbird.[11]
The icterid family of the New World is sometimes called the blackbird family because of some species' superficial resemblance to the common blackbird and other Old World thrushes, but they are not evolutionarily close, being related to the New World warblers and tanagers.[page needed][12] The term is often limited to smaller species with mostly or entirely black plumage, at least in the breeding male, notably the cowbirds,[13] the grackles,[14] and for around 20 species with "blackbird" in the name, such as the red-winged blackbird and the melodious blackbird.[12]
Subspecies
As would be expected for a widespread passerine bird species, several geographical subspecies are recognised. The treatment of subspecies in this article follows Clement et al. (2000).[8]
Female of subspecies merula
T. m. merula, the nominate subspecies, breeds commonly throughout much of Europe from Iceland, the Faroes and the British Isles east to the Ural Mountains and north to about 70 N, where it is fairly scarce. A small population breeds in the Nile Valley. Birds from the north of the range winter throughout Europe and around the Mediterranean including Cyprus and North Africa. The introduced birds in Australia and New Zealand are of the nominate race.[8]
T. m. azorensis is a small race which breeds in the Azores. The male is darker and glossier than merula.[15]
T. m. cabrerae, named for Ángel Cabrera, Spanish zoologist, resembles azorensis and breeds in Madeira and the western Canary Islands.[15]
T. m. mauretanicus, another small dark species with a glossy black male plumage, breeds in central and northern Morocco, coastal Algeria and northern Tunisia.[15]
First-summer male, probably subspecies aterrimus
T m. aterrimus breeds in Hungary, south and east to southern Greece, Crete northern Turkey and northern Iran. It winters in southern Turkey, northern Egypt, Iraq and southern Iran. It is smaller than merula with a duller male and paler female plumage.[15]
T. m. syriacus breeds on the Mediterranean coast of southern Turkey south to Jordan, Israel and the northern Sinai. It is mostly resident, but part of the population moves south west or west to winter in the Jordan Valley and in the Nile Delta of northern Egypt south to about Cairo. Both sexes of this subspecies are darker and greyer than the equivalent merula plumages.[8]
T. m. intermedius is an Asiatic race breeding from Central Russia to Tajikistan, western and north east Afghanistan, and eastern China. Many birds are resident but some are altitudinal migrants and occur in southern Afghanistan and southern Iraq in winter.[8] This is a large subspecies, with a sooty-black male and a blackish-brown female.[16]
The Asian subspecies, the relatively large intermedius also differs in structure and voice, and may represent a distinct species.[16] Alternatively, it has been suggested that they should be considered subspecies of T. maximus,[8] but they differ in structure, voice and the appearance of the eye-ring.[16][17]
Similar species
In Europe, the common blackbird can be confused with the paler-winged first-winter ring ouzel (Turdus torquatus) or the superficially similar common starling (Sturnus vulgaris).[18] A number of similar Turdus thrushes exist far outside the range of the common blackbird, for example the South American Chiguanco thrush (Turdus chiguanco).[19] The Indian blackbird, the Tibetan blackbird, and the Chinese blackbird were formerly considered subspecies of the common blackbird.[20]
Description
Historic image of blackbird in Nederlandsche Vogelen (1770)
The common blackbird of the nominate subspecies T. m. merula is 23.5–29 cm (9+1⁄4–11+1⁄2 in) in length, has a long tail, and weighs 80–125 g (2+7⁄8–4+3⁄8 oz). The adult male has glossy black plumage, blackish-brown legs, a yellow eye-ring and an orange-yellow bill. The bill darkens somewhat in winter.[18] The adult female is sooty-brown with a dull yellowish-brownish bill, a brownish-white throat and some weak mottling on the breast. The juvenile is similar to the female, but has pale spots on the upperparts, and the very young juvenile also has a speckled breast. Young birds vary in the shade of brown, with darker birds presumably males.[18] The first year male resembles the adult male, but has a dark bill and weaker eye ring, and its folded wing is brown, rather than black like the body plumage.[8]
Distribution and habitat
The common blackbird breeds in temperate Eurasia, North Africa, the Canary Islands, and South Asia. It has been introduced to Australia and New Zealand.[8] Populations are sedentary in the south and west of the range, although northern birds migrate south as far as northern Africa and tropical Asia in winter.[page needed][8] Urban males are more likely to overwinter in cooler climes than rural males, an adaptation made feasible by the warmer microclimate and relatively abundant food that allow the birds to establish territories and start reproducing earlier in the year.[21] Recoveries of blackbirds ringed on the Isle of May show that these birds commonly migrate from southern Norway (or from as far north as Trondheim) to Scotland, and some onwards to Ireland. Scottish-ringed birds have also been recovered in England, Belgium, Holland, Denmark, and Sweden.[22] Female blackbirds in Scotland and the north of England migrate more (to Ireland) in winter than do the males[23]
Common over most of its range in woodland, the common blackbird has a preference for deciduous trees with dense undergrowth. However, gardens provide the best breeding habitat with up to 7.3 pairs per hectare (nearly three pairs per acre), with woodland typically holding about a tenth of that density, and open and very built-up habitats even less.[page needed][24] They are often replaced by the related ring ouzel in areas of higher altitude.[25] The common blackbird also lives in parks, gardens and hedgerows.[26]
The common blackbird occurs at elevations up to 1,000 m (3,300 ft) in Europe, 2,300 m (7,500 ft) in North Africa, and at 900–1,800 m (3,000–5,900 ft) in peninsular India and Sri Lanka, but the large Himalayan subspecies range much higher, with T. m. maximus breeding at 3,200–4,800 m (10,500–15,700 ft) and remaining above 2,100 m (6,900 ft) even in winter.[8]
This widespread species has occurred as a vagrant in many locations in Eurasia outside its normal range, but records from North America are normally considered to involve escapees, including, for example, the 1971 bird in Quebec.[27] However, a 1994 record from Bonavista, Newfoundland, has been accepted as a genuine wild bird,[8] and the species is therefore on the North American list.[28]
Behaviour and ecology
The male common blackbird defends its breeding territory, chasing away other males or utilising a "bow and run" threat display. This consists of a short run, the head first being raised and then bowed with the tail dipped simultaneously. If a fight between male blackbirds does occur, it is usually short and the intruder is soon chased away. The female blackbird is also aggressive in the spring when it competes with other females for a good nesting territory, and although fights are less frequent, they tend to be more violent.[24]
The bill's appearance is important in the interactions of the common blackbird. The territory-holding male responds more aggressively towards models with orange bills than to those with yellow bills, and reacts least to the brown bill colour typical of the first-year male. The female is, however, relatively indifferent to bill colour, but responds instead to shinier bills.[29]
As long as winter food is available, both the male and female will remain in the territory throughout the year, although occupying different areas. Migrants are more gregarious, travelling in small flocks and feeding in loose groups in the wintering grounds. The flight of migrating birds comprises bursts of rapid wing beats interspersed with level or diving movement, and differs from both the normal fast agile flight of this species and the more dipping action of larger thrushes.[15]
Breeding
The male common blackbird attracts the female with a courtship display which consists of oblique runs combined with head-bowing movements, an open beak, and a "strangled" low song. The female remains motionless until she raises her head and tail to permit copulation.[24] This species is monogamous, and the established pair will usually stay together as long as they both survive.[15] Pair separation rates of up to 20% have been noted following poor breeding.[30] Although the species is socially monogamous, there have been studies showing as much as 17% extra-pair paternity.[31]
The nominate T. merula may commence breeding in March, but eastern and Indian races are a month or more later, and the introduced New Zealand birds start nesting in August (late winter).[8][25] The breeding pair prospects for a suitable nest site in a creeper or bush, favouring evergreen or thorny species such as ivy, holly, hawthorn, honeysuckle or pyracantha.[32] Sometimes the birds will nest in sheds or outbuildings where a ledge or cavity is used. The cup-shaped nest is made with grasses, leaves and other vegetation, bound together with mud. It is built by the female alone. She lays three to five (usually four) bluish-green eggs marked with reddish-brown blotches,[24] heaviest at the larger end;[25] the eggs of nominate T. merula are on average 29 mm × 21 mm (1+1⁄8 in × 13⁄16 in) in size and weigh 7.2 g (1⁄4 oz), of which 6% is shell.[33] Eggs of birds of the southern Indian races are paler than those from the northern subcontinent and Europe.[8]
The female incubates for 12–14 days before the altricial chicks are hatched naked and blind. Fledging takes another 10–19 (average 13.6) days, with both parents feeding the young and removing faecal sacs.[15] The nest is often ill-concealed compared with those of other species, and many breeding attempts fail due to predation.[34] The young are fed by the parents for up to three weeks after leaving the nest, and will follow the adults begging for food. If the female starts another nest, the male alone will feed the fledged young.[24] Second broods are common, with the female reusing the same nest if the brood was successful, and three broods may be raised in the south of the common blackbird's range.[8]
A common blackbird has an average life expectancy of 2.4 years,[35] and, based on data from bird ringing, the oldest recorded age is 21 years and 10 months.[36]
Breeding
Songs and calls
Alarm calls
In its native Northern Hemisphere range, the first-year male common blackbird of the nominate race may start singing as early as late January in fine weather in order to establish a territory, followed in late March by the adult male. The male's song is a varied and melodious low-pitched fluted warble, given from trees, rooftops or other elevated perches mainly in the period from March to June, sometimes into the beginning of July. It has a number of other calls, including an aggressive seee, a pook-pook-pook alarm for terrestrial predators like cats, and various chink and chook, chook vocalisations. The territorial male invariably gives chink-chink calls in the evening in an attempt (usually unsuccessful) to deter other blackbirds from roosting in its territory overnight.[24] During the northern winter, blackbirds can be heard quietly singing to themselves.[37] Like other passerine birds, it has a thin high seeet alarm call for threats from birds of prey since the sound is rapidly attenuated in vegetation, making the source difficult to locate.[38]
Feeding
Adult male feeding on berries in Lausanne, Switzerland
The common blackbird is omnivorous, eating a wide range of insects, earthworms, seeds and berries. It feeds mainly on the ground, running and hopping with a start-stop-start progress. It pulls earthworms from the soil, usually finding them by sight, but sometimes by hearing, and roots through leaf litter for other invertebrates. Small amphibians, lizards and (on rare occasions) small mammals are occasionally hunted.[39][40] This species will also perch in bushes to take berries and collect caterpillars and other active insects.[24] Animal prey predominates, and is particularly important during the breeding season, with windfall apples and berries taken more in the autumn and winter. The nature of the fruit taken depends on what is locally available, and frequently includes exotics in gardens.
Natural threats
Near human habitation the main predator of the common blackbird is the domestic cat, with newly fledged young especially vulnerable. Foxes and predatory birds, such as the sparrowhawk and other accipiters, also take this species when the opportunity arises.[41][42] However, there is little direct evidence to show that either predation of the adult blackbirds or loss of the eggs and chicks to corvids, such as the European magpie or Eurasian jay, decrease population numbers.[32]
A male attempting to distract a kestrel close to its nest
This species is occasionally a host of parasitic cuckoos, such as the common cuckoo (Cuculus canorus), but this is minimal because the common blackbird recognizes the adult of the parasitic species and its non-mimetic eggs.[43] In the UK, only three nests of 59,770 examined (0.005%) contained cuckoo eggs.[44] The introduced merula blackbird in New Zealand, where the cuckoo does not occur, has, over the past 130 years, lost the ability to recognize the adult common cuckoo but still rejects non-mimetic eggs.[45]
As with other passerine birds, parasites are common. Intestinal parasites were found in 88% of common blackbirds, most frequently Isospora and Capillaria species.[46] and more than 80% had haematozoan parasites (Leucocytozoon, Plasmodium, Haemoproteus and Trypanosoma species).[47]
Common blackbirds spend much of their time looking for food on the ground where they can become infested with ticks, which are external parasites that most commonly attach to the head of a blackbird.[48] In France,74% of rural blackbirds were found to be infested with Ixodes ticks, whereas, only 2% of blackbirds living in urban habitats were infested.[48] This is partly because it is more difficult for ticks to find another host on lawns and gardens in urban areas than in uncultivated rural areas, and partly because ticks are likely to be commoner in rural areas, where a variety of tick hosts, such as foxes, deer and boar, are more numerous.[48] Although ixodid ticks can transmit pathogenic viruses and bacteria, and are known to transmit Borrelia bacteria to birds,[49] there is no evidence that this affects the fitness of blackbirds except when they are exhausted and run down after migration.[48]
The common blackbird is one of a number of species which has unihemispheric slow-wave sleep. One hemisphere of the brain is effectively asleep, while a low-voltage EEG, characteristic of wakefulness, is present in the other. The benefit of this is that the bird can rest in areas of high predation or during long migratory flights, but still retain a degree of alertness.[50]
Status and conservation
The common blackbird has an extensive range, estimated at 32.4 million square kilometres (12.5 million square miles), and a large population, including an estimated 110 to 174 million individuals in Europe alone. The species is not believed to approach the thresholds for the population decline criterion of the IUCN Red List (i.e., declining more than 30% in ten years or three generations), and is therefore evaluated as least concern[1] In the western Palearctic, populations are generally stable or increasing,[1][15] but there have been local declines, especially on farmland, which may be due to agricultural policies that encouraged farmers to remove hedgerows (which provide nesting places), and to drain damp grassland and increase the use of pesticides, both of which could have reduced the availability of invertebrate food.[41]
The common blackbird was introduced to Australia by a bird dealer visiting Melbourne in early 1857,[51] and its range has expanded from its initial foothold in Melbourne and Adelaide to include all of southeastern Australia, including Tasmania and the Bass Strait islands.[52] The introduced population in Australia is considered a pest because it damages a variety of soft fruits in orchards, parks and gardens, including berries, cherries, stone fruit and grapes.[51] It is thought to spread weeds, such as blackberry, and may compete with native birds for food and nesting sites.[51][53]
The introduced common blackbird is, together with the native silvereye (Zosterops lateralis), the most widely distributed avian seed disperser in New Zealand. Introduced there along with the song thrush (Turdus philomelos) in 1862, it has spread throughout the country up to an elevation of 1,500 metres (4,921 ft), as well as outlying islands such as the Campbell and Kermadecs.[54] It eats a wide range of native and exotic fruit, and makes a major contribution to the development of communities of naturalised woody weeds. These communities provide fruit more suited to non-endemic native birds and naturalised birds than to endemic birds.[55]
In popular culture
"Sing a Song for Sixpence" cover illustration
The common blackbird was seen as a sacred though destructive bird in Classical Greek folklore, and was said to die if it consumed pomegranates.[56] Like many other small birds, it has in the past been trapped in rural areas at its night roosts as an easily available addition to the diet,[57] and in medieval times the practice of placing live birds under a pie crust just before serving may have been the origin of the familiar nursery rhyme:[page needed][57]
Sing a song of sixpence,
A pocket full of rye;
Four and twenty blackbirds baked in a pie!
When the pie was opened the birds began to sing,
Oh, wasn't that a dainty dish to set before the king?[58]
The common blackbird's melodious, distinctive song is mentioned in the poem Adlestrop by Edward Thomas;
And for that minute a blackbird sang
Close by, and round him, mistier,
Farther and farther, all the birds
Of Oxfordshire and Gloucestershire.[59]
In the English Christmas carol "The Twelve Days of Christmas", the line commonly sung today as "four calling birds" is believed to have originally been written in the 18th century as "four colly birds", an archaism meaning "black as coal" that was a popular English nickname for the common blackbird.[60]
The common blackbird, unlike many black creatures, is not normally seen as a symbol of bad luck,[57] but R. S. Thomas wrote that there is "a suggestion of dark Places about it",[61] and it symbolised resignation in the 17th century tragic play The Duchess of Malfi;[62] an alternate connotation is vigilance, the bird's clear cry warning of danger.[62]
The common blackbird is the national bird of Sweden,[63] which has a breeding population of 1–2 million pairs,[15] and was featured on a 30 öre Christmas postage stamp in 1970;[64] it has also featured on a number of other stamps issued by European and Asian countries, including a 1966 4d British stamp and a 1998 Irish 30p stamp.[65] This bird—arguably—also gives rise to the Serbian name for Kosovo, which is the possessive adjectival form of Serbian kos ("blackbird"), as in Kosovo polje ("Blackbird's Field").[66]
A common blackbird can be heard singing on the Beatles song "Blackbird".[67]
References
BirdLife International (2016). "Turdus merula". IUCN Red List of Threatened Species. 2016: e.T103888106A87871094. doi:10.2305/IUCN.UK.2016-3.RLTS.T103888106A87871094.en. Retrieved 19 November 2021.
Alderfer, Jonathan, ed. (2006). Complete Birds of North America. Washington, D.C.: National Geographic Society. p. 489. ISBN 0-7922-4175-4.
Long, John L. (1981). Introduced Birds of the World. Agricultural Protection Board of Western Australia. pp. 21–493. ISBN 9780876633182.
Linnaeus, Carolus (1758). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata (in Latin). Holmiae (Laurentii Salvii). p. 170.
Le Robert, Paul (2001). Le Grand Robert de la langue française (in French). Dictionnaires Le Robert. ISBN 2-85036-673-0.
Dictionary of the Scots Language. University of Edinburgh: Scottish Language Dictionaries.
Reilly, John (2018). The Ascent of Birds. Pelagic Monographs. Exeter: Pelagic. pp. 221–225. ISBN 978-1-78427-169-5.
Clement, Peter; Hathway, Ren; Wilczur, Jan (2000). Thrushes (Helm Identification Guides). Christopher Helm Publishers Ltd. ISBN 0-7136-3940-7.
Oxford English Dictionary, Oxford University Press, 1933: Bird (sense 2), Blackbird
Lockwood, W. B. (1984). Oxford Book of British Bird Names. Oxford: Oxford University Press. ISBN 0-19-214155-4.
Sinclair, I., & P. Ryan (2003). Birds of Africa south of the Sahara. Struik Publishers, Cape Town. ISBN 1-86872-857-9
Jaramillo, Alvaro; Burke, Peter (1997). New World Blackbirds: The Icterids. Helm Identification Guides. Christopher Helm Publishers Ltd. ISBN 0-7136-4333-1.
"All About Birds: Bronzed Cowbird". Cornell Lab of Ornithology. 2003. Retrieved 18 February 2008.
"All About Birds: Great-tailed Grackle". Cornell Lab of Ornithology. 2003. Retrieved 18 February 2008.
Snow, David; Perrins, Christopher M., eds. (1998). The Birds of the Western Palearctic concise edition (2 volumes). Oxford: Oxford University Press. ISBN 0-19-854099-X. p. 1215–1218
Collar, N. J. (2005). Common Blackbird (Turdus merula). p. 645 in: del Hoyo, J., Elliott, A., & Christie, D. A. eds. (2005) Handbook of the Birds of the World. Vol. 10. Cuckoo-shrikes to Thrushes. Lynx Edicions, Barcelona. ISBN 84-87334-72-5
Collar, N. J. (2005). Tibetan Blackbird (Turdus maximus). p. 646 in: del Hoyo, J., Elliott, A., & Christie, D. A. eds. (2005) Handbook of the Birds of the World. Vol. 10. Cuckoo-shrikes to Thrushes. Lynx Edicions, Barcelona. ISBN 84-87334-72-5
Mullarney, Killian; Svensson, Lars, Zetterstrom, Dan; Grant, Peter (2001). Birds of Europe. Princeton University Press. pp. 304–306. ISBN 0-691-05054-6
Fjeldså, J., & N. Krabbe (1990). The Birds of the High Andes. Zoological Museum, University of Copenhagen, Copenhagen. ISBN 87-88757-16-1
Gill, F.; Donsker, D. (20 July 2015). "Thrushes". IOC World Bird List, version 5.3. Retrieved 29 September 2015.
Partecke, J.; Gwinner, E. (2007). "Increased sedentariness in European blackbirds following urbanization: a consequence of local adaptation?". Ecology. 88 (4): 882–90. doi:10.1890/06-1105. PMID 17536705.
Eggeling W. J. (1960) The Isle of May: a Scottish nature reserve. Oliver and Boyd. p. 108.
Snow D. (1958) A Study of Blackbirds. George Allen & Unwin. p. 173.
Snow, David (1988). A Study of Blackbirds. British Museum (Natural History). ISBN 0-565-01021-2.
Evans G (1972). The Observer's Book of Birds' Eggs. London: Warne. p. 78. ISBN 0-7232-0060-2.
Holden, Peter (2012). RSPB Handbook Of British Birds. p. 225. ISBN 978-1-4081-2735-3.
McNeil, Raymond; Cyr, André (October 1971). "General Notes: European Blackbird (Turdus merula) in Quebec" (PDF). The Auk. 88 (4): 919–920. doi:10.2307/4083850. JSTOR 4083850.
"The A.O.U. Check-list of North American Birds". Check-list of North American Birds (Seventh ed.). AOU. Archived from the original on December 11, 2007. Retrieved 14 December 2007.
Bright, Ashleigh; Waas, Joseph R. (August 2002). "Effects of bill pigmentation and UV reflectance during territory establishment in blackbirds" (PDF). Animal Behaviour. 64 (2): 207–213. doi:10.1006/anbe.2002.3042. S2CID 51833485. Archived from the original (PDF) on 2011-09-29. Retrieved 2007-12-16.
Streif, Michael; Rasa O. Anne E. (2001). "Divorce and its consequences in the Common blackbird Turdus merula". Ibis. 143 (4): 554–560. doi:10.1111/j.1474-919X.2001.tb04882.x.
Garamszegia, László Zsolt; Anders Pape Møller (2004). "Extrapair paternity and the evolution of bird song". Behavioral Ecology. 15 (3): 508–519. doi:10.1093/beheco/arh041.
"Blackbird – Turdus merula". The Royal Horticultural Society/The Wildlife Trusts. Archived from the original on 29 October 2013. Retrieved 9 April 2012.
"Blackbird Turdus merula [Linnaeus, 1758]". BTOWeb BirdFacts. British Trust for Ornithology. Retrieved 30 December 2007.
"89% of Blackbird nest failures are attributed to predators". Game and Wildfowl Conservation Trust. Retrieved 23 January 2012.
"British garden birds – lifespan". garden-birds.co.uk. Archived from the original on 2007-04-24. Retrieved 7 April 2007.
"European Longevity Records". euring.org. Retrieved 15 December 2007.
"Blackbird". British Garden Birds. Archived from the original on 2016-01-09. Retrieved 4 January 2016.
Burton, Robert (1985). Bird behaviour. London: Granada. p. 125. ISBN 0-246-12440-7.
"We watched a blackbird kill and then fly away with a shrew. Is this unusual?". The RSPB. Retrieved 2021-04-16.
Elliott, Valerie. "Birds get taste for tadpoles as summer heat keeps worms under ground". The Times. ISSN 0140-0460. Retrieved 2021-04-16.
"Threats". Blackbird. Royal Society for the Protection of Birds. Retrieved 19 December 2007.
"Blackbird Action Plan" (PDF). Lambeth Council's Parks and Greenspaces Business Unit. Archived from the original (PDF) on 5 October 2007. Retrieved 11 December 2007.
Davies, N. B.; Brooke, M. de L. (1989). "An experimental study of co-evolution between the Cuckoo, Cuculus canorus, and its hosts. I. Host egg discrimination". Journal of Animal Ecology. 58 (1): 207–224. doi:10.2307/4995. JSTOR 4995.
Glue, David; Morgan, Robert (1972). "Cuckoo hosts in British habitats". Bird Study. 19 (4): 187–192. doi:10.1080/00063657209476342.
Hale, Katrina; Briskie, James V. (March 2007). "Response of introduced European birds in New Zealand to experimental brood parasitism" (PDF). Journal of Avian Biology. 38 (2): 198–204. doi:10.1111/j.2007.0908-8857.03734.x. ISSN 0908-8857. Archived from the original (PDF) on 2012-01-13. Retrieved 2012-01-16.
Misof, Katharina (2005). Eurasian Blackbirds (Turdus merula) and their gastrointestinal parasites: A role for parasites in life-history decisions? (PDF). Doctoral dissertation, Bonn. Archived from the original (PDF) on 2011-07-19.
Hatchwell, B. J.; Wood; Anwar, M. J. M.; Perrins C. M. (2000). "The prevalence and ecology of the haematozoan parasites of European blackbirds, Turdus merula". Canadian Journal of Zoology. 78 (4): 684–687. doi:10.1139/cjz-78-4-684.
Gregoire, Arnaud; Faivre, Bruno; Heeb, Philipp; Cezilly, Frank (2002). "A comparison of infestation patterns by Ixodes ticks in urban and rural populations of the Common Blackbird Turdus merula" (PDF). Ibis. 144 (4): 640–645. doi:10.1046/j.1474-919X.2002.00102.x. Archived from the original (PDF) on 2008-12-17.
Kipp, Susanne; Goedecke, Andreas; Dorn, Wolfram; Wilske, Bettina; Fingerle, Volke (May 2006). "Role of birds in Thuringia, Germany, in the natural cycle of Borrelia burgdorferi sensu lato, the Lyme disease spirochaete". International Journal of Medical Microbiology. 296: 125–128. doi:10.1016/j.ijmm.2006.01.001. PMID 16530003.
Rattenbourg, Neils C.; Amlaner, C. J.; Lima, S.L. (2000). "Behavioral, neurophysiological and evolutionary perspectives on unihemispheric sleep". Neuroscience and Biobehavioral Reviews. 24 (8): 817–842. doi:10.1016/S0149-7634(00)00039-7. PMID 11118608. S2CID 7592942.
Clarke, G. M.; Gross, S., Matthews, M.; Catling, P. C.; Baker, B.; Hewitt, C. L.; Crowther, D.; Saddler, S. R. (2000), Environmental Pest Species in Australia, Australia: State of the Environment, Second Technical Paper Series (Biodiversity), Department of the Environment and Heritage, Canberra. hdl:102.100.100/203987
"Common Blackbird". Birds in Backyards. Australian Museum. Retrieved 30 December 2007.
"Blackbird". Department of Agriculture, Western Australia. 16 December 2014. Retrieved 17 August 2017.
Falla, R. A., R. B. Sibson, and E. G. Turbott (1979). The new guide to the birds of New Zealand and outlying islands. Collins, Auckland. ISBN 0-00-216928-2
Williams, Peter A (2006). "The role of blackbirds (Turdus merula) in weed invasion in New Zealand" (PDF). New Zealand Journal of Ecology. 30 (2): 285–291.
Cooper, J.C. (1992). Symbolic and Mythological Animals. London: Aquarian Press. p. 38. ISBN 1-85538-118-4.
Cocker, Mark; Mabey, Richard (2005). Birds Britannica. London: Chatto & Windus. pp. 349–353. ISBN 0-7011-6907-9.
Opie, I. and P. (1997). The Oxford Dictionary of Nursery Rhymes (2nd ed.). Oxford: Oxford University Press. pp. 394–5. ISBN 0-19-869111-4.
"Adlestrop". Poets' Graves. Retrieved 7 December 2007.
"Birds of the Twelve Days of Christmas". 10,000 Birds. Retrieved 19 December 2013.
"A Blackbird Singing". Poemhunter. 13 January 2003. Retrieved 7 December 2007.
de Vries, Ad (1976). Dictionary of Symbols and Imagery. Amsterdam: North-Holland Publishing Company. p. 51. ISBN 0-7204-8021-3.
"Background – Sweden". Nationmaster. Retrieved 12 December 2007.
"Bird stamps from Sweden". Theme Birds on Stamps. Kjell Scharning. Retrieved 13 December 2007.
"218 Thrushes Turdidae". Theme Birds on Stamps. Kjell Scharning. Retrieved 8 June 2015.
Trbovich, Ana S. (2008). A Legal Geography of Yugoslavia's Disintegration. Oxford: Oxford University Press. p. 76. ISBN 978-0-19-533343-5.
"Blackbird". Rolling Stone. 10 April 2020.
Further reading
Snow, David W. (1987). The Blackbird. Shire Natural History ISBN 0-85263-854-X
Snow, David W. (1958). "The breeding of the Blackbird Turdus merula at Oxford". Ibis. 100 (1): 1–30. doi:10.1111/j.1474-919X.1958.tb00362.x.
Retrieved from "http://en.wikipedia.org/"
All text is available under the terms of the GNU Free Documentation License