Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Subsectio: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Euornithes
Cladus: Ornithuromorpha
Cladus: Ornithurae
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Ordo: Bucerotiformes
Familia: Bucerotidae
Genus: Rhyticeros
Species: R. cassidix – R. everetti – R. narcondami – R. plicatus – R. subruficollis – R. undulatus
Name
Rhyticeros Reichenbach, 1849
References
Rhyticeros is a genus of medium to large hornbills (family Bucerotidae) found in forests from Southeast Asia to the Solomons. They are sometimes included in the genus Aceros. On the other hand, most species generally placed in Aceros are sometimes moved to Rhyticeros, leaving Aceros as a monotypic genus only containing the rufous-necked hornbill.
All species generally placed in Rhyticeros have relatively low, conspicuously wreathed casques and a mainly dull whitish horn-colored bill. Both sexes have mainly black plumage, but the head and neck of the males are white or rufous. The tail is white except in the black-tailed Sumba hornbill. They have conspicuous inflatable skin on the throat, which is blue in all except the males of the plain-pouched and wreathed hornbills, where it is yellow.
Species
As generally recognized, the following species belong in the genus Rhyticeros:
| Image | Scientific name | Common Name | Distribution |
|---|---|---|---|
 |
Rhyticeros plicatus | Papuan (or Blyth's) hornbill | Wallacea and Melanesia. |
 |
Rhyticeros narcondami | Narcondam hornbill | Narcondam |
| Rhyticeros subruficollis | Plain-pouched hornbill | southern Myanmar, adjacent parts of western Thailand and northern Peninsular Malaysia | |
 |
Rhyticeros undulatus | Wreathed hornbill | north-eastern India and Bhutan, east and south through mainland Southeast Asia and the Greater Sundas in Indonesia |
 |
Rhyticeros everetti | Sumba hornbill | Sumba in the Lesser Sunda Islands |
 |
Rhyticeros cassidix | Knobbed hornbill | Sulawesi, Buton, Lembeh, Togian and Muna Island. |
An undescribed extinct hornbill species from Lifou in the Loyalty Islands, living until at least some 30,000 years ago, was initially placed in Aceros, but its biogeography places it with the species now in Rhyticeros (Steadman, 2006).
References
Kemp, A. C. (2001). Family Bucerotidae (Hornbills). pp. 436–523 in: del Hoyo, J., Elliott, A., & Sargatal, J. eds. (2001). Handbook of the Birds of the World. Vol. 6. Mousebirds to Hornbills. Lynx Edicions, Barcelona. ISBN 84-87334-30-X
Steadman, David William (2006): Extinction and Biogeography of Tropical Pacific Birds. University of Chicago Press. ISBN 0-226-77142-3
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