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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Subsectio: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Infraclassis: Aves
Ordo: Passeriformes
Subordo: Passeri
Infraordo: Passerida
Superfamilia: Sylvioidea

Familia: Pycnonotidae
Genus: Ixos
Species: I. everetti - I. malaccensis - I. mcclellandii - I. nicobariensis - I. philippinus - I. rufigularis - I. siquijorensis - I. virescens
Name

Ixos Temminck, 1825
References
Nouveau recueil de planches coloriées d'oiseaux livr.64 pl.382 fig.1

Ixos is a genus of passerine birds in the bulbul family, Pycnonotidae.

Taxonomy and systematics

The genus Ixos was introduced in 1825 by the Dutch zoologist Coenraad Jacob Temminck to accommodate the Javan bulbul.[1] The genus name is the Ancient Greek for "mistletoe".[2]

Some authorities have advocated a complete merger of the genus Ixos with Hypsipetes – and even the entire "Hypsipetes group" of bulbuls, which also includes Hemixos, Iole and Tricholestes –, being the oldest genus name Ixos would apply to all of them, rather than Hypsipetes as is often believed.[3] This re-classification seems hardly appropriate however, since Alophoixus and Setornis cannot be excluded from the "Hypsipetes group", and an all-out merge would turn the resultant "genus" Ixos into an ill-defined "wastebin taxon". The erroneous inclusion of I. virescens in Hypsipetes has caused the Nicobar bulbul to be listed under its invalid junior synonym H. nicobariensis rather than the valid names H. virescens or I. nicobariensis.[3]
Extant species

There are five extant species in the genus Ixos:[4][5]

Nicobar bulbul (Ixos nicobariensis)
Mountain bulbul (Ixos mcclellandii)
Streaked bulbul (Ixos malaccensis)
Javan bulbul (Ixos virescens)[5] (the type species)
Sumatran bulbul (Ixos sumatranus)[5] (split from I. virescens)

Former species

Formerly, some authorities also considered the following species (or subspecies) as species within the genus Ixos:

Cream-striped bulbul (as Ixos leucogrammicus)[6]
Spot-necked bulbul (as Ixos tympanistrigus)[7]
Grey-headed bulbul (as Ixos Fisquetti)[8]
Scaly-breasted bulbul (as Ixos squamatus)[9]
Red-whiskered bulbul (pyrrhotis) (as Ixos pyrrhotis)[10]
Red-whiskered bulbul (emeria) (as Ixos emeria)[11]
Chinese red-whiskered bulbul (as Ixos monticola)[12]
Brown-breasted bulbul (andersoni) (as Ixus Andersoni)[13]
Light-vented bulbul (hainanus) (as Ixus hainanus)[14]
White-eared bulbul (as Ixos leucotis)[15]
Sooty-headed bulbul (germani) (as Ixus Germani)[16]
White-spectacled bulbul (as Ixus xanthopygos)[17]
Upper Guinea bulbul (as Ixos inornatus)[18]
Dark-capped bulbul (as Ixos tricolor)[19]
Pale-eyed bulbul (as Ixus Davisoni)[20]
Yellow-throated bulbul (as Ixos xantholaemus)[21]
Yellow-vented bulbul (gourdini) (as Ixos gourdini)[22]
Spectacled bulbul (as Ixos erythropthalmos)[23]
Yellow-throated leaflove (as Ixus flavicollis)[24]
Yellow-bellied bulbul (as Ixos phaeocephalus)[25]
Sulphur-bellied bulbul (as Ixos palawanensis)[26]
Seram golden bulbul (as Ixos affinis)[27]
Philippine bulbul (as Ixos philippinus)[28]
Mindoro bulbul (as Ixos mindorensis)[29]
Visayan bulbul (as Ixos guimarasensis)[30]
Zamboanga bulbul (as Ixos rufigularis)[31]
Streak-breasted bulbul (as Ixos siquijorensis)[32]
Yellowish bulbul (as Ixos everetti)[33]
Brown-eared bulbul (as Ixos amaurotis)[34]

Footnotes

Temminck, Coenraad Jacob (1838) [1825]. Nouveau recueil de planches coloriées d'oiseaux, pour servir de suite et de complément aux planches enluminées de Buffon (in French). Volume 3, Livraison 64. Paris: F.G. Levrault. Plate 382, Fig. 1. The 5 volumes were originally issued in 102 livraison (parts), 1820-1839. For the dates see: Dickinson, E.C. (2001). "Systematic notes on Asian birds. 9. The Nouveau recueil de planches coloriées of Temminck & Laugier (1820–1839)". Zoologische Verhandelingen, Leiden. 335: 7–53.
Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 209. ISBN 978-1-4081-2501-4.
Gregory (2000), Pasquet et al. (2001), Moyle & Marks (2006)
"Bulbuls « IOC World Bird List". www.worldbirdnames.org. Retrieved 2017-03-10.
"Species Updates – IOC World Bird List". Retrieved 2021-05-29.
"Pycnonotus leucogrammicus - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-08.
"Pycnonotus tympanistrigus - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-08.
Eydoux F, L FASouleyet (1841). Voyage Autour du Monde sur la Corvette La Bonite. Paris: Arthus Bertrand. pp. 86–88.
"Pycnonotus squamatus - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-12.
"Pycnonotus jocosus pyrrhotis - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-12.
Giebel, Christoph (1877-01-01). Thesaurus Ornithologiae: bd. Pachycephala-Zosterops. Brockhaus.
"Pycnonotus jocosus monticola - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-12.
"Pycnonotus xanthorrhous andersoni - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-13.
"Pycnonotus sinensis hainanus - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-13.
"Pycnonotus leucotis - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-24.
"Pycnonotus aurigaster germani - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-26.
"Pycnonotus xanthopygos - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-26.
"Pycnonotus barbatus inornatus - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-27.
"Pycnonotus tricolor - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-27.
"Pycnonotus finlaysoni davisoni - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-29.
Jerdon, Thomas Claverhill (1863-01-01). The Birds of India: Being a Natural History of All the Birds Known to Inhabit Continental India, with Descriptions of the Species, Genera, Families, Tribes, and Orders, and a Brief Notice of Such Families as are Not Found in India, Making it a Manual of Ornithology Specially Adapted for India. Author.
"Pycnonotus goiavier gourdini - Avibase". avibase.bsc-eoc.org. Retrieved 2017-03-30.
"Pycnonotus erythropthalmos - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-01.
Catalogue of the Birds in the British Museum: Passeriformes, or perching birds. Cichlomorphœ: pt. III-IV, containing the ... family Timeliidœ (babbling-thrushes) by R.B. Sharpe. order of the Trustees. 1881-01-01.
"Alophoixus phaeocephalus - Avibase". avibase.bsc-eoc.org. Retrieved 2017-05-29.
"Iole palawanensis - Avibase". avibase.bsc-eoc.org. Retrieved 2017-06-05.
"Thapsinillas [affinis, mystacalis or longirostris] (Golden Bulbul) - Avibase". avibase.bsc-eoc.org. Retrieved 2017-10-18.
"Hypsipetes philippinus (Philippine Bulbul) - Avibase". avibase.bsc-eoc.org. Retrieved 2017-11-03.
"Hypsipetes mindorensis (Mindoro Bulbul) - Avibase". avibase.bsc-eoc.org. Retrieved 2017-11-04.
"Hypsipetes guimarasensis (Visayan Bulbul) - Avibase". avibase.bsc-eoc.org. Retrieved 2017-11-05.
"Hypsipetes rufigularis (Zamboanga Bulbul) - Avibase". avibase.bsc-eoc.org. Retrieved 2017-11-05.
"Hypsipetes siquijorensis (Streak-breasted Bulbul) - Avibase". avibase.bsc-eoc.org. Retrieved 2017-11-05.
"Hypsipetes everetti (Yellowish Bulbul) - Avibase". avibase.bsc-eoc.org. Retrieved 2017-11-06.

"Hypsipetes amaurotis (Brown-eared Bulbul) - Avibase". avibase.bsc-eoc.org. Retrieved 2017-11-06.

References

Gregory, Steven M. (2000): Nomenclature of the Hypsipetes Bulbuls (Pycnonotidae). Forktail 16: 164–166. PDF fulltext
Moyle, Robert G. & Marks, Ben D. (2006): Phylogenetic relationships of the bulbuls (Aves: Pycnonotidae) based on mitochondrial and nuclear DNA sequence data. Mol. Phylogenet. Evol. 40(3): 687–695. doi:10.1016/j.ympev.2006.04.015 (HTML abstract)
Pasquet, Éric; Han, Lian-Xian; Khobkhet, Obhas & Cibois, Alice (2001): Towards a molecular systematics of the genus Criniger, and a preliminary phylogeny of the bulbuls (Aves, Passeriformes, Pycnonotidae). Zoosystema 23(4): 857–863. PDF fulltext

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