Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Cladus: Telluraves
Cladus: Australaves
Ordo: Passeriformes
Subordo: Passeri
Infraordo: Corvida
Superfamilia: Meliphagoidea
Familia: Acanthizidae
Genus: Calamanthus
Species: Calamanthus campestris Subspecies: C. c. campestris - C. c. dorrie - C. c. ethelae - C. c. hartogi - C. c. isabellinus - C. c. rubiginosus - C. c. wayensis - C. c. winiam
Name
Calamanthus campestris (Gould, 1841)
Vernacular names
References
Proceedings of the Zoological Society of London (1840) Pt8 no.95 p. 171
The rufous fieldwren (Calamanthus campestris) also known as the desert wren or sandplain wren[2] is a species of insectivorous bird in the family Acanthizidae, endemic to Australia.
Kerang District, Lake Tyrell -Victoria, Australia
Taxonomy
The rufous fieldwren is one of 63 species of the Australasian Warbler family Acanthizidae.[3] Which include gerygones, pardalotes, bristlebirds and thornbills.
Subspecies
Calamanthus campestris includes the following subspecies:
C. c. winiam - (Campbell, AJ. & Campbell, AG., 1927) from s.e Australian mallee.
C. c. campestris - (Gould, 1841) from s. South Australia and Nullabor Plain.
C. c. rubiginosus - (Campbell, AJ., 1899)
C. c. dorrie - (Mathews, 1912)
C. c. hartogi - (Carter, 1916)
C. c. wayensis - (Mathews, 1912)
C. c. isabellinus - (North, 1896)
C. c. montanellus (Milligan, 1903)
There is both historical and contemporary contention to Calamanthus campestris montanellus' split from a subspecies of the rufous fieldwren to a taxonomically recognised species: the Western Fieldwren.
The 1926 Royal Australasian Ornithological Union Checklist[4] recognised four individual fieldwren species; the striated fieldwren (fuliginosus), the rufous fieldwren(campestris), the rusty frieldwren(isabellinus) and the rock fieldwren (montanellus). This was contended at the time by Mathews (1912), who's 'official' bird reference list documented just striated and rufous fieldwrens.[5]
The International Ornithological Committee's World Bird List v.12.1[4] and IUCN's Red List[1] both list the Western Fieldwren as its own species, where as the Handbook of Australian, New Zealand and Antarctic Birds (HANZAB)[2] supports Parker & Eckert (1983),[6] and Christidis & Boles (1994)[7] in only recognising C. campestris and C. fuliginosus as the two fieldwren species.
Current in print field guides highlight this division with the CSIRO The Australian Bird Guide (first printed 2018 and revised edition) only recognises the striated fieldwren and the rufous fieldwren.[8] Whilst Pizzey & Knight's The Field Guide to the Birds of Australia (7th edition) includes the Western fieldwren: C. montanellus as the third species.[8]
Description
The rufous fieldwren is a small terrestrial bird, measuring 11.5–13.5 cm (4.5–5.3 in) long, 15.5–19.5 cm (6.1–7.6 in) wingspan,[2] 11–16 g (0.38–0.56 oz) weight.[8]
The eight subspecies are divisible into five groups by their physical traits, characterised by the prominent colour of their mantle, back, scapulars, underbody and degree of striation throughout their plumage.
Subspecies C.c. campestris and C.c. rubiginosus is predominantly grey backed with a slight rufous wash, a cream toned underbody with dark striations.
Subspecies C.c. winiam features a grey back, thick streaking on the underside.
Subspecies C.c. Isabellinus and C.c. wayenis rich rufous-brown upper parts, wish washed cinnamon underparts, has very fine, if not obsolete streaking.
Island Subspecies C.c. Dorrie and C.c. Hartogi have pale-grey upper parts, white underneath with moderate striations.
C.c montanellus is the darkest form, resembling closely to the striated fieldwren Olive grey above with a pale yellow underside.[2]
Females of all subspecies are descriptively similar, only differing through a slightly duller supercillium. Sexual dimorphism is most prominent in subspecies C. montantellus with the chin, throat and supercillium being white in males and yellowish-white and duller in females.[2]
Rufous fieldwren
Both sexes are known to sing, which is a cheerful, melodius and clear 'whirr-whirr-chick-chick-whirr-ree-ree' whistled from atop low laying shrubs.[9][8][2] Singing only occurs during their breeding season, which in some sub-populations can occur year-round. Their song can typically be heard at day break and for about 30 minutes after the sun has set. Alarm calls consist of sharp 'chrr-r-r' and 'han-han-han'[2]
Rufous fieldwrens are usually observed singularly or in pairs, but can be seen in small family groups of up to five and are often mingled in inter-species flocks with similarly sized inland thornbills, shy heathwrens and slender-billed thornbills. Their tail always strongly cocked including when hopping along ground, though sometimes when running they will flatten their tail out horizontally.[2]
Striated fieldwren
Unlikely to be confused with similar species if clearly spotted, though the rufous fieldwren does share similar traits to the striated fieldwren. Best distinguishing features are differences in habitat, with the rufous fieldwren preferring low shrub or heathland, where as the striated fieldwren occupies rank herbage or cutting-grass swamps. Though can overlap through coastal melaleuca heathland.[8][2]
The rufous fieldwren has noticeably greyer upperparts with whiter underparts and has a distinctive longer and finer decurved bill than its striated counterpart whose bill is straighter and diagnostically wedge-shaped in profile, with much heavier streaking through plumage than rufous fieldwrens[2][9][8]
Distribution and Habitat
Populations
Occurring in multiple sub-populations, the subspecies of the Rufous Fieldwren occur across the four southern states of mainland Australia.
Habitat
Due to the widespread but disjunct range of the rufous fieldwren sub-populations, there is a diverse variety of habitat that the rufous field wren occupies. Populations to the West preference dry and open woodlands and coastal heaths, mainly in the temperate and sub-tropical zone of Western Australia. On the North-west Nullabor Plains, rufous fieldwren populations occur throughout pearl bluebush shrubland, clumps of ray flower and usually throughout a diverse variety of shrubs such as mallee paperbark, mallee honey myrtle and dwarf sheoaks.
Subspecies occurring through the arid and semi-arid regions of central Australia mainly occur through chenopod shrublands and heathlands gravitating towards vegetation stands of Atriplex and Sclerolaena saltbush, Maireana bluebush and Sarcocornia samphire shrublands and heathlands.[2]
Eastern populations occur throughout gibber plains, saline or brackish wetlands with a preference for rocky areas sparesly vegetated with spinifex groundcover and eucalypts.[10] Rufous field wren are occasionally observed in the Little Desert, Victoria amongst stunted mallee eucalypts and spinifex grasslands.
Behaviour
Breeding
Breeding has been recorded from June to April with egg laying and incubation timing varying geographically. Sub populations in Western Australia have been observed breeding year round. Nesting sites are scattered throughout the rufous fieldwren's entire range, though nest records are sparse, representing sites from North-West Victoria through to West coast Western Australia.[2]
Nesting sites are predominately ground-based, under the canopy of shrubs or grass tussocks, though nests been found out in the open on mounds of soil, also above ground in the branches of low-laying vegetation; usually saltbush, bluebush, samphire or occasionally spinifex grass.[2]
Nests are globular or domed in construction, measuring 15.2 cm (5.98 in) external length, 10.2 cm (4.01 in) external diameter, 3.8 cm (1.49 in) entrance diameter, and made largely with strips of bark, or fine dry grass. Nests are lined with softer materials such as wool, fur, feathers, plant down and insect cocoons. A nest observed from subspecies C. winiam was lined with the feathers of Australian magpies, brown falcons, quails and duck down.[2]
Females sit closely and are likely to be the sole incubator, with egg incubation periods largely unknown.[2] Clutch sizes are similar across all subspecies with 3–4 eggs, with occasionally as little as two being laid.[11]
The smooth and glossy eggs are oval to rounded-oval in shape measuring 19–22 mm (0.74–0.86 in) in length, and 14.5–18 mm (0.57–0.7 in) diameter width. There is geographical variation in egg colouration with inland subspecies' eggs being uniformly pale chocolate coloured, grading to a darker toned cap. Subspecies closer to the coast observed to have buff salmon, chocolate-red and chestnut-brown egg colourations. Subspecies C. montanellous' eggs closely resemble eggs of striated fieldwren and feature deeper toned flecked markings along egg surface.
Both parents attend to and feed fledglings, who are dependant ≤17 days. Nestling success not well documented, though birds can be quick to abandon nest and young, or dismantle nests when disturbed by human observers. Rufous fieldwren's nests can be prone to brood parasitism by pallid cuckoos, black-eared cuckoos, horsfield's and shining bronze cuckoos.[2]
Feeding
Little is known of the rufous fieldwren's diet and feeding behaviours outside of casual observations.[2] They are predominate insectivores[11][8] feeding largely on beetles, wasps and small grasshoppers. They will occasionally also eat small spiders, snails and seeds from Setaria viridis and Erodium cygnorum[2].
Threats and Human Interaction
Degradation of habitat, largely from over-grazing of livestock, clearing land for agriculture and the invasion of weeds in Western Australia's Wheatbelt has caused steady decline and isolation in rufous field-wren sub populations from their former range. Rufous fieldwrens will persist and inhabit an area until the last remnant of cover remaining have been removed, with windbreaks occasionally providing suitable habitat.[2]
Isolated population's survival are dependent on the maintenance of their current remnant habitat selective planting to link vegetation a viable solution, as threats of fires on the remaining habitat can lead to localised extinctions. Despite the presence of livestock, feral cats and occasional large-scale landscape fires, subspecies C. Hartogi persist on Dirk Hartog Island.[2]
Conservation Status
The International Union for Conservation of Nature's Red List of Threatened Species lists the Rufous fieldwren as Least Concern.[1]
References
BirdLife International (2017). "Calamanthus campestris". The IUCN Red List of Threatened Species. doi:10.2305/IUCN.UK.2017-1.RLTS.T103691268A112512396.en.
Higgins, P.J; Peter, J.M. (2002). Handbook of Australian, New Zealand and Antarctic Birds. Vol. 6: Pardalotes to shrike-thrushes. Melbourne: Oxford University Press. pp. 258–269.
Gill, F.; Donsker, D.; Rasmussen, P. (eds.). IOC World Bird List 10.2. doi:10.14344/IOC.ML.10.2.
Royal Australasian Ornithologists Union Checklist Committee (1926). "Official Checklist of the Birds of Australia". EMU.
Mathews, G.M. (1912). "A Reference-List to the Birds of Australia". Novitates Zoologicae. 18: 171–446. doi:10.5962/bhl.part.1694.
Parker, S.A.; Eckert, H.J. (1983). "Remarks on the taxonomy of the genus Calamanthus (fieldwrens)". South Australian Ornithologist. 29: 65–75.
Christidis, L.; Boles, W.E. (1994). The Taxonomy and Species of Birds of Australia and Its Territories. Royal Australasian Ornithologists Union.
Menkhorst, Peter; Rogers, Danny; Clarke, Rohan; Davies, Jeff; Marsack, Peter; Franklin, Kim (2020). The Australian Bird Guide (Revised ed.). Melbourne: CSIRO. p. 348.
Pizzey, G; F, Knight (2003). The Field Guide to the Birds of Australia (7th ed.).
Campbell, I.; Woods, S.; Leseburg, N. (2014). Birds of Australia: A Photographic Guide. Princeton University Press.
Pizzey, Graham; Knight, Frank; Menkhorst, Peter (2003). Graham Pizzey & Frank Knight The field guide to the birds of Australia. Pymble, N.S.W: HarperCollins.
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