Nymphalinae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Nymphalinae
Tribus: Coeini – Junoniini – Kallimini – Melitaeini – Nymphalini – Victorinini

Genera (incertae sedis): Kallimoides – Rhinopalpa – Vanessula
Name

Nymphalinae Swainson, 1827
References

Wahlberg, N., Brower, A.V.Z. & Nylin, S. 2005. Phylogenetic relationships and historical biogeography of tribes and genera in the subfamily Nymphalinae (Lepidoptera: Nymphalidae). Biological Journal of the Linnean Society 86(2): 227–251. DOI: 10.1111/j.1095-8312.2005.00531.x Open access. Reference page.

Vernacular names
日本語: タテハチョウ亜科
Nederlands: schoenlappers
中文: 蛺蝶亞科

The Nymphalinae are a subfamily of brush-footed butterflies (family Nymphalidae). Sometimes, the subfamilies Limenitidinae, and Biblidinae are included here as subordinate tribe(s), while the tribe Melitaeini is occasionally regarded as a distinct subfamily.

Systematics

The traditionally recognized tribes of Nymphalinae are here listed in the presumed phylogenetic sequence:[1]

Coeini (six or seven genera)
Nymphalini – anglewings, tortoiseshells and relatives (about 15 genera, two are fossil)
Kallimini (about five genera)
Victorinini (four genera, formerly in Kallimini)
Junoniini (about five genera)
Melitaeini – fritillaries (about 25 genera)

Genera incertae sedis are:

Rhinopalpa – the wizard
Kallimoides Shirôzu & Nakanishi, 1984 – African leaf butterfly
Vanessula Dewitz, 1887

In addition to these, Crenidomimas is sometimes placed here, but may belong in the Limenitidinae, genus Euryphura.

The Late Eocene fossil genus Lithodryas might belong here, but it is generally assigned to the Lycaenidae.
References

See references in Savela (2010)