Coccus

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Paraneoptera
Superordo: Condylognatha
Ordo: Hemiptera
Subordo: Sternorrhyncha
Superfamilia: Coccoidea

Familia: Coccidae
Subfamilia: Coccinae
Genus: Coccus
Species: C. acaciae – C. acrossus – C. aequale – C. africanus – C. almoraensis – C. alpinus – C. anneckei – C. antidesmae – C. asiaticus – C. australe – C. brasiliensis – C. bromeliae – C. cajani – C. cambodiensis – C. cameronensis – C. capparidis – C. caudatus – C. caviramicolus – C. celatus – C. circularis – C. colemani – C. cryptus – C. deformosum – C. delottoi – C. diacopeis – C. discrepans – C. duartei – C. ehretiae – C. elatensis – C. erion – C. erythrinae – C. ficicola – C. giliomeei – C. guerinii – C. gymnospori – C. heckrothi – C. hesperidum – C. hordeolum – C. illuppalamae – C. incisus – C. inquilinum – C. insolens – C. inyangombae – C. jaculator – C. koleos – C. kosztarabi – C. lambirensis – C. latioperculatum – C. leurus – C. lidgetti – C. litzeae – C. lizeri – C. longulus – C. lumpurensis – C. macarangae – C. macarangicolus – C. malloti – C. melaleucae – C. microogenes – C. milanjianus – C. moestus – C. muiri – C. multisetus – C. murex – C. narcodes – C. nyika – C. oogenes – C. ophiorrhizae – C. opimum – C. paradeformosum – C. penangensis – C. phalaridis – C. planchonii – C. praetermissus – C. pruni – C. pseudelongatus – C. pseudohesperidum – C. pseudomagnoliarum – C. pseudotumuliferus – C. purpuratus – C. ramakrishnai – C. resinatum – C. rhodesiensis – C. rubellus – C. saltuarius – C. schini – C. secretus – C. sectilis – C. smaragdinus – C. sociabilis – C. sordidus – C. stipulaeformis – C. subacutus – C. subhemisphaericus – C. synapheae – C. takanoi – C. tangandae – C. tenebricophilum – C. trichodes – C. tumuliferus – C. viridis – C. viridulus

Nomen dubium: C. phalaridis
Name

Coccus Linnaeus, 1758

Type species: Coccus hesperidum Linnaeus, 1758, by subsequent designation under Plenary Powers in ICZN, 1985: 139

References

Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Tomus I. Holmiæ (Stockholm): impensis direct. Laurentii Salvii. 824 pp. DOI: 10.5962/bhl.title.542 BHL p. 455 BHL Reference page.
ICZN 1985. Opinion 1303. Coccus Linnaeus, 1758 and Parthenolecanium Šulc, 1908 (Insecta, Hemiptera, Homoptera): type species designated. Bulletin of Zoological Nomenclature 42(2): 139–141. BHL Reference page.
Choi, J. & Lee, S. 2018. Review of the genus Coccus Linnaeus from Korea, with description of a new species (Hemiptera, Coccomorpha, Coccidae). ZooKeys 734: 121–135. DOI: 10.3897/zookeys.734.22774. Reference page.
Gullan, P.J., Kondo, T., Fiala, B. & Quek, S-P. 2018. Taxonomy of coccids (Hemiptera: Coccidae: Coccus L.) associated with Crematogaster ants (Hymenoptera: Formicidae) in the stems of Macaranga plants (Euphorbiaceae) in Southeast Asia. Zootaxa 4521(1): 1–51. DOI: 10.11646/zootaxa.4521.1.1 Open access pdf Reference page.
Łagowska, B. & Hodgson, C.J. 2019 On some new and old soft scale insects (Hemiptera: Coccomorpha: Coccidae) from Africa, with description of a new Coccus species and introduction of a new genus. Zootaxa 4612(3): 373–386. DOI: 10.11646/zootaxa.4612.3.4. Reference page.
Lin, Y-P., Tanaka, H., Kondo, T. & Cook, L.G. 2017. A newly recognised species that has been confused with the global polyphagous pest scale insect, Coccus hesperidum Linnaeus (Hemiptera: Coccomorpha: Coccidae). Zootaxa 4320(3): 571–591. DOI: 10.11646/zootaxa.4320.3.10. Full article (PDF) Reference page.
Williams, D.J.; Ben-Dov, Y. 2009: A review of species names combined with the genus name Coccus Linnaeus (Hemiptera: Sternorrhyncha: Coccoidea). Zootaxa, 2285: 1–64. Abstract & excerpt

Links

Coccus Taxon details on Fauna Europaea
Scale Net
ZooBank: D7AFF012-1B2A-4F9B-BA23-795DCC253C10

Coccus is a genus of scale insects in the family Coccidae. Several species, such as Coccus viridis, a major pest of coffee, are major agricultural pests.[1] The type species is Coccus hesperidum Linnaeus.
Species

Coccus acaciae (Newstead, 1917)[2]
Coccus bromeliae Bouché, 1833[3]
Coccus capparidis (Green, 1904)[4]
Coccus celatus De Lotto, 1960[5]
Coccus gymnospori (Green, 1908)[6]
Coccus hesperidum Linnaeus, 1758, the brown soft scale[7]
Coccus longulus (Douglas, 1887)[8]
Coccus penangensis Morrison, 1921[9]
Coccus pseudelongatus (Brain, 1920)[10]
Coccus pseudohesperidum (Cockerell, 1895)[11]
Coccus pseudomagnoliarum (Kuwana, 1914)[12]
Coccus formicarii (Green, 1896)[13]
Coccus viridis (Green, 1889)[14]

Former species

Coccus aceris Fabricius, 1794 synonym for Eulecanium tiliae (Linnaeus, 1758)
Coccus alni Modeer, 1778 synonym for Eulecanium tiliae (Linnaeus, 1758)
Coccus arens Hodgson, 1968 synonym for Coccus capparidis (Green, 1904)
Coccus arundinariae Sanders, 1906 synonym for Maacoccus arundinariae (Green, 1904)
Coccus bauhini Targioni Tozzetti, 1867 synonym for Kermes ilicis (Linnaeus, 1758)
Coccus bicruciatus Sanders, 1906 synonym for Maacoccus bicruciatus (Green, 1904)
Coccus cacti Cockerell, 1893 synonym for Dactylopius coccus Costa, 1829
Coccus celticum Takahashi, 1955 synonym for Coccus longulus (Douglas, 1887)
Coccus celtium Sasscer, 1911 synonym for Coccus longulus (Douglas, 1887)
Coccus coffeae Kirkaldy, 1902 synonym for Saissetia coffeae (Walker, 1852)
Coccus cypraeola Dalman, 1826 synonym for Eulecanium tiliae (Linnaeus, 1758)
Coccus deltae Corseuil & Barbosa, 1971 synonym for Parthenolecanium perlatum (Cockerell, 1898)
Coccus diversipes Cockerell, 1905 synonym for Kilifia diversipes (Cockerell, 1905)
Coccus elongatus Sanders, 1909 synonym for Coccus longulus (Douglas, 1887)
Coccus fagi Baerensprung, 1849 ambiguous synonym for Cryptococcus fagisuga Lindinger, 1936
Coccus fagi Walker, 1852 ambiguous synonym for Cryptococcus fagisuga Lindinger, 1936
Coccus ficus Fernald, 1903 synonym for Coccus longulus (Douglas, 1887)
Coccus frontalis Sanders, 1906 synonym for Coccus longulus (Douglas, 1887)
Coccus fuscus Gmelin, 1790 synonym for Eulecanium tiliae (Linnaeus, 1758)
Coccus iceryi Tao et al., 1983 synonym for Pulvinaria iceryi (Signoret, 1869)
Coccus ilicis Linnaeus, 1758 synonym for Kermes ilicis (Linnaeus, 1758)
Coccus impar Bodkin, 1917 synonym for Mesolecanium impar (Cockerell, 1898)
Coccus lacca Kerr, 1782 synonym for Kerria lacca (Kerr, 1782)
Coccus mali Schrank, 1781 synonym for Eulecanium tiliae (Linnaeus, 1758)
Coccus marsupialis Sanders, 1906 synonym for Marsipococcus marsupialis (Green, 1904)
Coccus namunakuli Ali, 1971 synonym for Parlatoria namunakuli (Green, 1922)
Coccus perlatus Fernald, 1903 synonym for Parthenolecanium perlatum (Cockerell, 1898)
Coccus polonicus Linnaeus, 1758 synonym for Porphyrophora polonica (Linnaeus, 1758)
Coccus pyri Schrank, 1801 synonym for Eulecanium tiliae (Linnaeus, 1758)
Coccus radicum Beckmann, 1790 synonym for Porphyrophora polonica (Linnaeus, 1758)
Coccus rubi Schrank, 1801 synonym for Eulecanium tiliae (Linnaeus, 1758)
Coccus salicum Fabricius, 1781 synonym for Eulecanium tiliae (Linnaeus, 1758)
Coccus tiliae Linnaeus, 1758 synonym for Eulecanium tiliae (Linnaeus, 1758)
Coccus xylostei Schrank, 1801 synonym for Eulecanium tiliae (Linnaeus, 1758)

References

Ananthakrishnan, Taracad Narayanan (2004). "Family Coccidae (soft scales, wax scales, tortoise scales)". In Nayar, K. Karunakaran; Ananthakrishnan, Taracad Narayanan; David, B. Vasantharaj (eds.). General and applied entomology (2nd ed.). New Delhi: Tata McGraw-Hill. p. 437. ISBN 978-0-07-043435-6.
Ben-Dov, Yair (1977). "Taxonomy of the long brown scale Coccus longulus (Douglas), stat. n.(Homoptera: Coccidae)". Bulletin of Entomological Research. 67 (1): 89–95. doi:10.1017/S0007485300010919.
Ben-Dov, Yair; Cox, Jennifer M. (1990). "The identity of five species of scale insects (Hem., Homoptera, Coccoidea), living on ornamental plants, originally described by PF Bouché". Entomologist's Monthly Magazine. 126: 79–84.
Sanders, James Glossbrenner (1906). Catalogue of recently described Coccidae. United States Department of Agriculture Technical Series, number 12, part 1. Washington, D.C.: Bureau of Entomology, United States Department of Agriculture. p. 372.
De Lotto, Giovanni (1960). "The Green Scales of Coffee in Africa South of the Sahara (Homoptera, Coccidae)". Bulletin of Entomological Research. 51 (2): 389–403. doi:10.1017/S0007485300058041.
Sanders, James Glossbrenner (1909). The euonymus scale (Chionaspis euonymi Comstock). United States Department of Agriculture Circular 114. Washington, D.C.: Bureau of Entomology, United States Department of Agriculture. Google Books - no preview
Bartlett, Blair Ralph (1978). "Coccidae". In Clausen, Curtis P. (ed.). Introduced Parasites and Predators of Arthropod Pests and Weeds: A world review. Agriculture Handbook No. 480. Washington, D.C.: Agricultural Research Service, United States Department of Agriculture. pp. 57–74, view=1up, seq=68 page 60.
Hall, Wilfrid John (1922). Observations on the Coccidae of Egypt. Technical and Scientific Service Bulletin, number 22. Cairo: Ministry of Agriculture, Egypt, Government Press.
Ben-Dov, Yair (1993). A Systematic Catalogue of the Soft Scale Insects of the World (Homoptera: Coccoidea: Coccidae): With Data on Geographical Distribution, Host Plants, Biology, and Economics Importance. Flora & Fauna Handbook, No. 9. Gainesville, Florida: CRC Press. p. 85. ISBN 978-1-877743-13-9.
De Lotto, Giovanni (1957). "On some Ethiopian species of the genus Coccus (Homoptera: Coccoidea: Coccidae)". Journal of the Entomological Society of Southern Africa. 20: 295–314.
Fernald, Maria Elizabeth Smith (1903). A catalogue of the Coccidae of the world. Bulletin of the Hatch Experiment Station of the Massachusetts Agricultural College, Number 88. Amherst, Massachusetts: Carpenter and Morehouse. p. 173, Item 860. book at Google Books
Bartlett 1978, p. 61
Lin, Yen-Po; Kondo, Takumasa; Gullan, Penny; Cook, Lyn G. (2013). "Delimiting genera of scale insects: molecular and morphological evidence for synonymising Taiwansaissetia Tao, Wong and Chang with Coccus Linnaeus (Hemiptera: Coccoidea: Coccidae)". Systematic Entomology. 38 (2): 249–264. doi:10.1111/j.1365-3113.2012.00664.x.
Fernald 1903, p. 174