Coccidae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Paraneoptera
Superordo: Condylognatha
Ordo: Hemiptera
Subordo: Sternorrhyncha
Superfamilia: Coccoidea

Familia: Coccidae
Subfamiliae: Cardiococcinae – Ceroplastinae – Cissococcinae – Coccinae – Cyphococcinae – Eriopeltinae – Eulecaniinae – Filipiinae – Myzolecaniinae –
Overview of genera (163)
A

Acantholecanium – Acanthopulvinaria – Akermes – Alecanium – Alecanochiton – Alecanopsis – Alichtensia – Allopulvinaria – Anapulvinaria – Anopulvinaria – Antandroya – Anthococcus – Aphenochiton – Austrolecanium – Austrolichtensia – Avricus – Aztecalecanium
B

Bodenheimera – Bombacoccus
C

Cajalecanium – Cardiococcus – Ceronema – Ceroplastes – Ceroplastodes – Chlamydolecanium – Cissococcus – Coccus – Conofilippia – Couturierina – Cribrolecanium – Cribropulvinaria – Cryptinglisia – Cryptostigma – Crystallotesta – Ctenochiton – Cyclolecanium – Cyphococcus
D

Dermolecanium – Dicyphococcus – Didesmococcus – Differococcus – Discochiton – Drepanococcus
E

Edwallia – Epelidochiton – Ericeroides – Ericerus – Eriopeltis – Etiennea – Eucalymnatus – Eulecanium – Eumashona – Eutaxia – Exaeretopus
FG

Filippia – Fistulococcus – Foldilecanium
H

Hadzibejliaspis – Hallicoccus – Halococcus – Hemilecanium – Houardia
I

Idiosaissetia – Inglisia – Insularicoccus
K

Kalasiris – Kenima – Kilifia – Kozaricoccus
L

Lagosinia – Lecaniococcus – Lecanochiton – Lecanopsis – Leptopulvinaria – Lichtensia – Loemica – Luzulaspis
M

Maacoccus – Magnococcus – Mallococcus – Mametia – Mariacoccus – Marsipococcus – Megalecanium – Megalocryptes – Megapulvinaria – Megasaissetia – Melanesicoccus – Membranaria – Mesembryna – Mesolecanium – Messinea – Metaceronema – Metapulvinaria – Millericoccus – Milviscutulus – Mitrococcus – Myzolecanium

Nemolecanium – Neolecanium – Neolecanochiton – Neoplatylecanium – Neosaissetia
P

Palaeolecanium – Paracardiococcus – Paractenochiton – Parafairmairia – Paralecanium – Parapulvinaria – Parasaissetia – Parthenolecanium – Peculiaricoccus – Pendularia – Perilecanium – Pharangococcus – Philephedra – Phyllostroma – Physokermes – Platinglisia – Platylecanium – Platysaissetia – Plumichiton – Poaspis – Podoparalecanium – Poropeza – Pounamococcus – Prionococcus – Prococcus – Protopulvinaria – Pseudalichtensia – Pseudokermes – Pseudophilippia – Pseudopulvinaria – Psilococcus – Pulvinaria – Pulvinariella – Pulvinarisca – Pulvinella
R

Rhizopulvinaria – Rhodococcus – Richardiella
S

Saccharolecanium – Saissetia – Schizochlamidia – Scythia – Sphaerolecanium – Stenolecanium – Stictolecanium – Stotzia – Suareziella – Symonicoccus
T

Takahashia – Tectopulvinaria – Tillancoccus – Torarchus – Toumeyella – Trijuba
UV

Udinia – Umbonichiton – Umwinsia – Vinsonia – Vitrococcus – Vittacoccus
WX

Waricoccus – Waxiella – Xenolecanium

All genera in overview are current and taken from Catalogue of Life 29th November 2018 edition.
Name

Coccidae Fallén, 1814
References

Fallén, C.F. 1814. Specimen novam Hemiptera disponendi methodum exhibens. Berling, Lundae. 26 pp. Reference page.
Caballero, A., Ramos-Portilla, A.A. & Kondo, T. 2017. Scale insects (Hemiptera: Coccomorpha) on sugarcane in Colombia, with description of a new species of Tillancoccus Ben-Dov (Coccidae). Zootaxa 4258(5): 490–500. DOI: 10.11646/zootaxa.4258.5.6. Reference page.
Choi, J., Soysouvanh, P., Lee, S. & Hong, K-J. 2018. Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos. Zootaxa 4460(1): 1–62. DOI: 10.11646/zootaxa.4460.1.1 Paywall Reference page.
Garonna, A.P., Scarpato, S., Vicinanza, F. & Espinosa, B. 2015: First report of Toumeyella parvicornis (Cockerell) in Europe (Hemiptera: Coccidae). Zootaxa 3949(1): 142–146. DOI: 10.11646/zootaxa.3949.1.9. Preview (PDF) Full article (PDF) Reference page.
Gullan, P.J.; Hodgson, C.J. 1998: A new genus of Australian soft scale insect (Hemiptera: Coccidae) with species on Capparis (Capparaceae) and Doryphora (Monimiaceae) from New South Wales. Proceedings of the Linnean Society of New South Wales, 119: 197–217.
Henderson, R.C. 1995: Lectotype designations and definitions for five species of New Zealand Coccidae (Hemiptera: Homoptera). New Zealand journal of zoology, 22: 105–108. DOI: 10.1080/03014223.1995.9518026
Hodgson, C.J.; Henderson, R.C. 2000: Coccidae (Insecta: Hemiptera: Coccoidea). Fauna of New Zealand, (41)
Hodgson, C.J.; Henderson, R.C. 2004: Coccidae (Insecta: Hemiptera: Coccoidea): adult males, pupae and prepupae of indigenous species. Fauna of New Zealand, (51)
Hodgson, C.J. & Williams, D.J. 2018. Revision of the soft scale genus Paralecanium (Hemiptera: Coccomorpha: Coccidae) with the introduction of three new genera and twenty new species. Zootaxa 4443(1): 1–162. DOI: 10.11646/zootaxa.4443.1.1 Paywall Reference page.
Kondo, T. 2010: Description of a new coccid (Hemiptera, Coccidae) on avocado (Persea americana Mill.) from Colombia, South America. ZooKeys, 42: 37–45. ISSN: 1313-2970 (online) ISSN: 1313-2989 (print) DOI: 10.3897/zookeys.42.377
Kondo, T.; Gullan, P.J. 2010: The Coccidae (Hemiptera: Coccoidea) of Chile, with descriptions of three new species and transfer of Lecanium resinatum Kieffer & Herbst to the Kerriidae. Zootaxa, 2560: 1–15. Preview PDF
Kondo, T. & G. Pellizzari, 2011: Description of a new species of Toumeyella Cockerell (Hemiptera, Coccidae) from Mexico, with a taxonomic key to Mexican species. Revista Brasileira de Entomologia 55 (2): 229–233. Abstract and full article: DOI: 10.1590/S0085-56262011005000019.
Łagowska, B., Martin, J.H. & Hodgson, C.J. 2017. A new species of Marsipococcus Cockerell & Bueker (Hemiptera: Coccomorpha: Coccidae) from Malaysia. Zootaxa 4358(2): 295–310. DOI: 10.11646/zootaxa.4358.2.4. Reference page.
Lin, Y-P., Ding, Z.Y., Gullan, P.J. & Cook, L.G. 2017. A newly recognised Australian endemic species of Austrolecanium Gullan & Hodgson 1998 (Hemiptera: Coccidae) from Queensland. Zootaxa 4272(1): 119–130. DOI: 10.11646/zootaxa.4272.1.6. Full article (PDF) Reference page.

Vernacular names
English: soft scales
日本語: カタカイガラムシ科

The Coccidae are a family of scale insects belonging to the superfamily Coccoidea. They are commonly known as soft scales, wax scales or tortoise scales. The females are flat with elongated oval bodies and a smooth integument which may be covered with wax. In some genera they possess legs but in others, they do not, and the antennae may be shortened or missing. The males may be winged or wingless.[1]

Genera

There are >1,100 spp. in 171 genera worldwide.

Subfamily Myzolecaniinae
Akermes
Alecanium
Alecanopsis
Cribolecanium
Cryptostigma
Cyclolecanium
Halococcus
Houardia
Megasaissetia
Myzolecanium
Neolecanium
Paractenochiton
Pseudophilippia
Richardiella
Torarchus
Toumeyella
Xenolecanium[2]
Antecerococcus
Ceroplastes
Coccus Linnaeus, 1758
Eucalymnatus Cockerell, 1901
Eulecanium
Kilifia De Lotto, 1965
Lecanium
Luzulaspis Targioni Tozzetti, 1868
Metaceronema
Milviscutulus
Paralecanium Cockerell in Cockerell & Parrott 1899
Parasaissetia Takahashi 1955
Parthenolecanium Šulc 1908
Physokermes Targioni Tozzetti 1868
Protopulvinaria Cockerell 1894
Pulvinaria Targioni-tozzetti, 1867
Saissetia Deplanche, 1858