Bradysia

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Antliophora
Ordo: Diptera
Subordo: Nematocera
Infraordo: Bibionomorpha
Superfamilia: Sciaroidea

Familia: Sciaridae
Genus: Bradysia
Species (570):
A

B. abnormalis – B. acerba – B. acerpontia – B. acinacauda – B. acutimedialis – B. acutiungula – B. aequalis – B. aequans – B. aequispina – B. affinis – B. agrestis – B. aitkeni – B. aktru – B. albanensis – B. albochaeta – B. aleica – B. alpicola – B. altaica – B. alutacea – B. amurensis – B. angustata – B. angustipennis – B. angustoocularis – B. angustostylus – B. antehilaris – B. antemorio – B. antica – B. apicalba – B. aprica – B. arcana – B. arcuans – B. arcula – B. ascenda – B. asiatica – B. atrcornea – B. atrorubens – B. atrospina – B. auriculariae – B. auriculata – B. austera –
B

B. baishanzuna – B. basalimedia – B. basiangustata – B. basilatissima – B. bellingeri – B. bellstedti – B. bicolor – B. bicuspidata – B. bidentata – B. bijca – B. bilobata – B. bishopi – B. bispina – B. bispinifera – B. bituberculata – B. boliviensis – B. boninensis – B. borori – B. brachyflagellata – B. brachystyla – B. brachytoma – B. breviallata – B. brevifurcata – B. brevipalpis – B. brevispina – B. browni – B. brunnipes – B. bulbiformis – B. bulbigera – B. bulbostyla –
C

B. caldaria – B. campbellensis – B. campylura – B. capillata – B. cardiocrivora – B. cauta – B. cellarum – B. centidens – B. chamei – B. chandleri – B. chenjinae – B. chikuni – B. chilensis – B. chloroantennata – B. chlorocornea – B. choui – B. chunguii – B. chunmeiae – B. ciliocera – B. cinerascens – B. clara – B. clavistylis – B. clavulata – B. cohilaris – B. columbia – B. commixta – B. compacta – B. condensa – B. confinis – B. congata – B. conglomerata – B. conicopalpi – B. conspersa – B. consumpta – B. crassicera – B. crinita – B. cryptacantha – B. ctenoura – B. cubiticaudata – B. cucumeris – B. cuneiforma – B. curvipedicula – B. cuspidalis –
D

B. dactylina – B. dalmatina – B. dazhulana – B. defossa – B. delectabilis – B. desolata – B. diabolica – B. dichaeta – B. dictyophorae – B. difformis – B. dilucida – B. diluta – B. dimelanata – B. diplostida – B. disjuncta – B. distincta – B. diversiabdominalis – B. diversispina – B. dolichoflagella – B. dolosa – B. drakenbergensis – B. dubia – B. dureti – B. dux –
E

B. elegans – B. elobata – B. entraqueensis – B. ericia – B. evanescens – B. excelsa – B. exigua – B. expolita –
F

B. falcata – B. fallaciosa – B. familiaris – B. fatigans – B. felti – B. fenestralis – B. filigera – B. fimbriata – B. flavida – B. flavipila – B. floribunda – B. florida – B. fluminensis – B. fochi – B. fontinalis – B. forattinii – B. forcipulata – B. fugaca – B. fuliginosa – B. fulvescens – B. fulvicauda – B. fumida – B. fumigans – B. funga – B. fungicola – B. furcata – B. furcatina – B. fusca – B. fuscescens – B. fuscothorax – B. fuscovirgata –
G

B. gabyae – B. gemellata – B. geninispina – B. gibbosa – B. giraudii – B. gonata – B. grandicellaris – B. gratiosa – B. guadunana – B. guayanensis – B. gutianshana –
H

B. hamata – B. hartii – B. hastata – B. helleri – B. helveolus – B. henana – B. hexacantha – B. hexamelanata – B. hexamera – B. heydemanni – B. hilaris – B. hildae – B. hirsutiseta – B. holsatica – B. hortensis – B. hortulana – B. humiliflagella – B. huoditangana – B. hupleuroti – B. hyalescens – B. hyalichaeta – B. hygida –
I

B. iberiana – B. iberica – B. impatiens – B. incidera – B. individua – B. infuscatipennis – B. insolita – B. insulana – B. insularis – B. introflexa – B. inundata – B. inusitata – B. inversa – B. iridipennis – B. ismayi –
J

B. janduisi – B. jianglensis – B. johannseni – B. jucunda –
K

B. kaigangi – B. kaihuana – B. kairensis – B. kaiseri – B. kassebeeri – B. kirstenae – B. kopetdagica – B. kraussi –
L

B. laboriosa – B. lapponica – B. lasiphlepia – B. latiala – B. latiflagellomera – B. latimedialis – B. latipennis – B. latiponsa – B. latispinosa – B. latiterga – B. laurencei – B. lembkei – B. leptocera – B. leptoptera – B. leucopeza – B. lii – B. lilienthalae – B. liubana – B. lobatula – B. lobosa – B. longiaciculata – B. longicauda – B. longicolla – B. longicubitalis – B. longimedia – B. longimentula – B. longinguans – B. longioculosa – B. longiradiomedialis – B. longiseta – B. longispina – B. longistyla – B. longistylia – B. longitibalis – B. longitoma – B. longqishana – B. longyuwana – B. loricata – B. loriculata – B. loudoni – B. loxostyla – B. lucichaeta – B. lucida – B. luhi – B. luodiana – B. lutaria – B. luteicauda – B. luteolineata –
M

B. mabiusi – B. macclurei – B. macfarlanei – B. macroptera – B. macrotrichata – B. macrura – B. madens – B. maggiaensis – B. magnifica – B. malitiosa – B. matogrossensis – B. mecocteniuni – B. media – B. medialis – B. mediterranea – B. megamelanata – B. meigeni – B. meinacusi – B. melaina – B. melanota – B. melanura – B. melina – B. mellea – B. mesochra – B. mexicana – B. microcteniuni – B. microspina – B. minima – B. minitriacantha – B. minorlobus – B. minpleuroti – B. minuens – B. minuscula – B. minutissima – B. moerens – B. moesta – B. molokaiensis – B. monacantha – B. monospina – B. munda – B. mutua –
N

B. nedongana – B. neocampestris – B. neopraecox – B. neoreflexa – B. neotropica – B. nervosa – B. nicolae – B. nigricans – B. nigrisensilla – B. nigrispina – B. nigroantennata – B. nigrochaeta – B. ningshana – B. nitida – B. nitidicollis – B. nocturna – B. noduspina – B. nomica – B. novalobata – B. nubilans – B. nudilobata –
O

B. obesa – B. obscuricostalis – B. obsolescens – B. obtusa – B. oculosa – B. odoriphaga – B. opaca – B. optata – B. ovata –
P

B. pallidipes – B. pallidiventris – B. pallipes – B. palpiclavata – B. paracantha – B. paradichaeta – B. paralobata – B. paranocturna – B. parareflexa – B. pararufescens – B. parasita – B. parilis – B. paucidens – B. paulistana – B. pauperata – B. pectinata – B. pectoralis – B. pedisequa – B. penna – B. pentadactyla – B. peraffinis – B. perfecta – B. phalerata – B. phili – B. picea – B. picticornis – B. pilata – B. pilularia – B. placida – B. planistylata – B. pollicis – B. polonica – B. polychaeta – B. ponticula – B. portoricensis – B. posthilaris – B. postlobata – B. postrufescens – B. praecox – B. praehilaris – B. praemonticola – B. praemorio – B. praepaupera – B. pratincola – B. procera – B. profunda – B. promissa – B. propinguans – B. protohilaris – B. pruinosula – B. pseudocampestris – B. pseudodalmatina – B. pseudohilaris – B. pseudopolonica – B. pteropleurostigma – B. pugiata – B. pumila – B. pustulispina –
Q

B. qinlingana – B. quadrata – B. quadridentata – B. quadrisetosa – B. quadrispinistylata – B. quadrispinosa – B. quasiradicum – B. quinquespina –
R

B. radicicola – B. radicum – B. ravensburgensis – B. rectinervis – B. reflexa – B. regens – B. regulariformis – B. regularis – B. reticulata – B. reynoldsi – B. robusticalcar – B. rostrata – B. rubelloi – B. rubra – B. rubrascuta – B. rufescens – B. ruginosa –
S

B. sachalinensis – B. salina – B. sandalimorpha – B. santorina – B. sarcinispina – B. sauteri – B. semihilaris – B. semirufescens – B. senilis – B. serrata – B. seticornis – B. setigera – B. sexdentata – B. siberica – B. sibirica – B. sicelidis – B. siciliensis – B. siegeli – B. siliquaris – B. silvestrii – B. silvosa – B. simulans – B. simulatrix – B. smithae – B. snyderi – B. songxiana – B. sordida – B. soudeki – B. spatitergum – B. spherostyla – B. spinata – B. spinea – B. spinellosa – B. spinostyla – B. spissa – B. splendida – B. strenua – B. stricta – B. strigata – B. stupenda – B. subaffinis – B. subamoena – B. subangustata – B. subaprica – B. subbetuleti – B. subbispinifera – B. subbrunnipes – B. subconfinis – B. subconiferoterga – B. subcuneostylata – B. subfungicola – B. subgiraudi – B. subgrandis – B. subhilaris – B. subiridipennis – B. submarginata – B. submoesta – B. submonticola – B. submorio – B. subpallidipes – B. subrufescens – B. subsantorina – B. subscabricornis – B. subspinea – B. subvernalis – B. surinamensis –
T

B. tenebricosa – B. tenuicalcar – B. tenuicauda – B. thailandina – B. tianzei – B. tibiaspinosa – B. tilicola – B. tirolensis – B. tobiasi – B. tomentosa – B. tongmaiana – B. triacanthifera – B. triaristata – B. tridonta – B. trifurca – B. trigonospina – B. trinidadensis – B. triseta – B. trisetosa – B. trispinifera – B. trispinosa – B. tristans – B. tritici – B. trivialis – B. trivittata – B. truncorum – B. tuberculata – B. tuberosa – B. tumidicauda – B. tumulta – B. tungmuguana – B. tupinamba – B. turgida –
U

B. umesaoi – B. uncipleuroti – B. unguicauda – B. unguispinata – B. urticae –
V

B. vagans – B. validolobata – B. vaneyi – B. varians – B. variopalpa – B. venezuelensis – B. ventricosa – B. vergens – B. vernalis – B. verruca – B. virgulata –
W

B. wanzhii – B. watsoni – B. weiningana – B. willinki –
X

B. xenoreflexa – B. xianyingi – B. xiaochengi – B. xizangana –
Y

B. yangi – B. yucollectia – B. yuhuacavernaria – B. yungata –
Z

B. zetterstedti – B. zheana – B. zhenzhongi – B. zingiberis – B. zizhongi – B. zonata

[source: Catalogue of Life: 2012 Annual Checklist plus (22) B. aequispina – B. aktru – B. aleica – B. arcula – B. centidens – B. clavistylis – B. filigera – B. fontinalis – B. gemellata – B. gibbosa – B. iberiana – B. iberica – B. kirstenae – B. melina – B. nudilobata – B. paucidens – B. perfecta – B. planistylata – B. regulariformis – B. seticornis – B. spissa – B. stricta
]

Add (1): B. quercina
Name

Bradysia Winnertz, 1867
References

Winnertz, J. 1867. Beitrag zu einer Monographie der Sciarinen 1–187. W. Braunmüller. DOI: 10.5962/bhl.title.14230 Open access. Reference page.
Köhler, A. & Menzel, F. 2013. New records of black fungus gnats (Diptera: Sciaridae) from New Caledonia, with the description of two new Bradysia species and an updated checklist. Zootaxa 3718(1): 63–72. DOI: 10.11646/zootaxa.3718.1.5 Reference page.
Menzel, F. & Ziegler, J. 2002. New records of Diptera from the Hohe Tauern National Park (Austria), including information on flower visiting and the description of two new black fungus gnat species (Sciaridae). Studia dipterologica 8(2): 355–409.
Mohrig, W., Kauschke, E. & Broadley, A. 2018. New black fungus gnats (Diptera: Sciaridae) from Eastern Australia. Zootaxa 4450(2): 203–241. DOI: 10.11646/zootaxa.4450.2.3 Open access PDF Reference page.
Vilkamaa, P., Hippa, H. & Mohrig, W. 2012. The genus Bradysia Winnertz (Diptera, Sciaridae) in New Caledonia, with the description of thirteen new species. Zootaxa 3489: 25–44. Preview Reference page.
Yang, X., Shi, K., Heller, K., Menzel, F., Huang, J.H. & Wu, H. 2019. Morphology and DNA barcodes of two species of Bradysia Winnertz from China (Diptera, Sciaridae), with the description of Bradysia minorlobus Yang, Shi & Huang sp. n. Zootaxa 4612(1): 85–94. DOI: 10.11646/zootaxa.4612.1.5 Reference page.

Bradysia is a genus of fungus gnat in the family Sciaridae.[3][4] They are commonly known as darkwinged fungus gnats.[3] They are considered a major pest in greenhouse agriculture because they thrive in the moist conditions common inside greenhouses and feed on the plants being grown within.[5] Bradysia is a large genus containing over 500 living species,[6] with at least 65 species found in North America[3] and 172 in Europe.[2]

Bradysia species are a major pollinator of plants such as Aspidistra elatior.[7]
Species

Selected species of Bradysia include:[6][8][9]

B. affinis (Zetterstedt, 1838)
B. alpicola (Winnertz, 1867)
B. amoena (Winnertz, 1867)
B. angustipennis (Winnertz, 1867)
B. angustoocularis Mohrig & Krivosheina, 1989
B. aprica (Winnertz, 7) C
B. arcula Vilkamaa, Salmela & Hippa, 2007
B. atracornea Mohrig & Menzel, 1992
B. austera Menzel et al., 2006
B. bellingeri Shaw, 1953
B. biformis (Lundbeck, 1898)
B. bispina (Fisher, 1938)
B. brevispina Tuomikoski, 1960
B. browni (Shaw, 1935)
B. caldaria (Linyner, 1895)
B. cellarum Frey, 1948
B. chlorocornea Mohrig & Menzel, 1992
B. cinerascens (Grzegorzek, 1884)
B. confinis (Winnertz, 1867)
B. coprophila (Lintner, 1895)
B. cucumeris (Johannsen, 1912)
B. cuneiforma Komarova, 1997
B. dichaeta (Shaw, 1941)
B. diluta (Johannsen, 1912)
B. distincta (Staeger, 1840)
B. dux (Johannsen, 1912)
B. ericia (Pettey, 1918)
B. excelsa Menzel & Mohrig, 1998
B. expolit (Coquillett, 1900)
B. falcata (Pettey, 1918)
B. fatigans (Johannsen, 1912)
B. felti (Pettey, 1918)
B. fenestralis (Zetterstedt, 1838)
B. flavipila Tuomikoski, 1960
B. fochi (Pettey, 1918)
B. forcipulata (Lundbeck, 1898)
B. fugaca Mohrig & Mamaev, 1989
B. fulvicauda (Felt, 1898)
B. fumida (Johannsen, 1912)
B. fungicola (Winnertz, 1867)
B. giraudii (Egger, 1862)
B. groenlandica (Holmgren, 1872)
B. hamata (Pettey, 1918)
B. hartii (Johannsen, 1912)
B. hastata (Johannsen, 1912)
B. heydemanni (Lengersdorf, 1955)
B. hilariformis Tuomikoski, 1960
B. hygida Sauaia & Alves, 1968
B. fungicola (Winnertz, 1867)
B. impatiens (Johannsen, 1912)
B. inusitata Tuomikoski, 1960
B. iridipennis (Zetterstedt, 1838)
B. ismayi Menzel et al., 2006[10]
B. johannseni Enderlein, 1912
B. jucunda (Johannsen, 1912)
B. kaiseri (Shaw, 1941)
B. lapponica (Lengersdorf, 1926)
B. lembkei Mohrig & Menzel, 1990
B. lobosa (Pettey, 1918)
B. longicubitalis (Lengersdorf, 1924)
B. longimentula Sasakawa, 1994
B. longispina (Pettey, 1918)
B. loriculata Mohrig, 1985
B. macclurei (Shaw, 1941)
B. macfarlanei (Jones, 1920)
B. macroptera (Pettey, 1918)
B. mellea (Johannsen, 1912)
B. mesochra (Shaw, 1941)
B. moesta Frey, 1948
B. munda (Johannsen, 1912)
B. mutua (Johannsen, 1912)
B. neglecta (Johannsen, 1912)
B. nemoralis (Meigen, 1818)
B. nervosa (Meigen, 1818)
B. nitidicollis (Meigen, 1818)
B. nigripes (Meigen, 1830)
B. nigrispina Menzel et al., 2006
B. nomica Mohrig & Rsschmann, 1996
B. normalis Frey, 1948
B. ocellaris (Comstock, 1882)
B. odoriphaga Yang & Zhang, 1985
B. ovata (Pettey, 1918)
B. pallipes (Fabricius, 1787)
B. paradichaeta (Shaw, 1941)
B. parilis (Johannsen, 1912)
B. pauperata (Winnertz, 1867)
B. pectoralis (Staeger, 1840)
B. penna (Pettey, 1918)
B. peraffinis Tuomikoski, 1960
B. petaini (Pettey, 1918)
B. picea (Rubsaamen, 1894)
B. pilata (Pettey, 1918)
B. placida (Winnertz, 1867)
B. pollicis (Pettey, 1918)
B. polonica (Lengersdorf, 1929)
B. praecox (Meigen, 1818)
B. procera (Winnertz, 1868)
B. prolifica (Felt, 1898)
B. protohilaris Mohrig & Krivosheina, 1983
B. quadrispinistylata Alam, 1988
B. quadrispinosa (Pettey, 1918)
B. quadrispinosa (Pettey, 1918)
B. reflexa Tuomikoski, 1960
B. sachalinensis Mohrig & Krivosheina, 1989
B. scabricornis Tuomikoski, 1960
B. sexdentata (Pettey, 1918)
B. silvestrii (Kieffer, 1910)
B. similigibbosa Köhler & Menzel, 2013
B. smithae Menzel & Heller, 2005
B. spinata (Pettey, 1918)
B. splendida Mohrig & Krivosheina, 1989
B. strenua (Winnertz, 1867)
B. strigata (Staeger, 1840)
B. subaprica Mohrig & Krivosheina, 1989
B. subgrandis (Shaw, 1941)
B. subrufescens Mohrig & Krivosheina, 1989
B. subvernalis Mohrig & Heller, 1992
B. tilicola (Loew, 1850)
B. trifurca (Pettey, 1918)
B. trispinifera Mohrig & Krivosheina, 1979
B. tritici (Coquillett, 1895)
B. trivialis (Johannsen, 1912)
B. trivittata (Staeger, 1840)
B. trivittata (Staeger, 1840)
B. unguicauda (Malloch, 1923)
B. urticae Mohrig & Menzel, 1992
B. vagans (Winnertz, 1868)
B. varians (Johannsen, 1912)
B. vernalis (Winnertz, 1868)
B. zetterstedti Mohrig & Menzel, 1993

Data sources: C = Catalogue of Life,[6] I = ITIS,[8] N = NCBI[9]
References

"Genus Bradysia". BugGuide. Iowa State University Department of Entomology. Retrieved 2019-08-17.
"Bradysia Winnertz, 1867". Fauna Europaea. Retrieved 2019-08-18.
"darkwinged fungus gnats - Bradysia spp". entnemdept.ufl.edu. Retrieved 2019-08-18.
Köhler, Arne; Menzel, Frank (2013). "New records of Black Fungus Gnats (Diptera: Sciaridae) from New Caledonia, with the description of two new Bradysia species and an updated checklist". Zootaxa. Magnolia Press. 3718 (1): 63–72. doi:10.11646/zootaxa.3718.1.5. PMID 26258208. S2CID 23947245.a
Cloyd, Raymond (2015-04-09). "Ecology of Fungus Gnats (Bradysia spp.) in Greenhouse Production Systems Associated with Disease-Interactions and Alternative Management Strategies". Insects. 6 (2): 325–332. doi:10.3390/insects6020325. ISSN 2075-4450. PMC 4553482. PMID 26463188.
"Browse Bradysia". Catalogue of Life. Retrieved 2019-08-18.
Suetsugu, Kenji; Sueyoshi, Masahiro (2018-01-01). "Subterranean flowers of Aspidistra elatior are mainly pollinated by not terrestrial amphipods but fungus gnats" (PDF). Ecology. 99 (1): 244–246. doi:10.1002/ecy.2021. ISSN 1939-9170. PMID 29136275.
"Bradysia". Integrated Taxonomic Information System. Retrieved 2019-08-17.
"Bradysia". NCBI Taxonomy Browser. Retrieved 2019-08-17.