Agraphydrus

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Hydrophiloidea

Familia: Hydrophilidae
Subfamilia: Acidocerinae
Genus: Agraphydrus
Species: A. abrasus – A. activus – A. acutus – A. aethiopicus – A. agilis – A. ampullatus – A. andringitra – A. angustatus – A. anhuianus – A. arduus – A. atripalpis – A. attenuatus – A. audax – A. batak – A. bicoloratus – A. bilardoi – A. biltoni – A. biprojectus – A. brevilobatus – A. calvus – A. camerunensis – A. cantonensis – A. chinensis – A. comes – A. confusus – A. congolensis – A. conicus – A. connexus – A. contractus – A. coomani – A. coronarius – A. decipiens – A. fasciatus – A. fikaceki – A. flavescens – A. flavipes – A. fontis – A. forcipatus – A. fujianensis – A. gereckei – A. globipenis – A. goldschmidti – A. gracilipalpis – A. igneus – A. insidiator – A. jilanzhui – A. longipalpus – A. longipenis – A. madagascarensis – A. malkini – A. masatakai – A. matoposensis – A. montanus – A. niger – A. occultus – A. orientalis – A. palawanensis – A. pauper – A. pelingeni – A. politus – A. puzhelongi – A. reductus – A. reticulicep – A. rhodesiensis – A. rivalis – A. robustus – A. schoenmanni – A. scutifer – A. setifer – A. splendens – A. tenuipalpis – A. thaiensis – A. tumidus – A. umbrosus – A. uncinatus – A. usambaraensis – A. vadoni – A. variabilis – A. wangmiaoi – A. yunnanensis – A. zetteli
Name

Agraphydrus Régimbart, 1903

Type species: Agraphydrus punctatellus Régimbart, 1903
Synonymy

Gymnhelochares Orchymont 1932: 692
Type species: Helochares geminus Armand D'Orchymont, 1932
Pseudohelochares Satô 1960: 76
Type species: Pseudohelochares narusei Masataka Satô, 1960
Pseudopelthydrus Jia 1998: 225
Type species: Pseudopelthydrus longipalpus Feng-Long Jia, 1998
Megagraphydrus Hansen, 1999: 137(synonymized by Minoshima, Komarek & Ôhara, 2015)
Type species: Megagraphydrus siamensis Hansen, 1999

References
Additional references

Hansen, M. 1999: Fifteen new genera of Hydrophilidae (Coleoptera), with remarks on the generic classification of the family. Entomologica scandinavica, 30: 121–172.
Jia, F.-L. 1998: A new genus Pseudopelthydrus gen. n. from Hainan Island, China (Coleoptera: Hydrophilidae: Hydrophilinae). Chinese journal of entomology 18(4): 225–230. abstract and pdf Reference page.
Komarek, A. 2019. Taxonomic revision of Agraphydrus RÉGIMBART, 1903 III. Southeast Asia (except Philippines) and Australian Region (Coleoptera: Hydrophilidae: Acidocerinae. Koleopterologische Rundschau 89: 151–316. Full article (PDF). Reference page.
Komarek, A. 2020. Taxonomic revision of Agraphydrus RÉGIMBART, 1903 IV. Africa, Western Asia, and redescription of the genus .(Coleoptera: Hydrophilidae: Acidocerinae). Koleopterologische Rundschau 90: 127–200. Full article (PDF). Reference page.
Komarek, A. & Freitag, H. 2020. Taxonomic revision of Agraphydrus RÉGIMBART, 1903 V. Philippine species and their first DNA barcodes (Coleoptera: Hydrophilidae: Acidocerinae). Koleopterologische Rundschau 90: 201–242. Full article (PDF). Reference page.
Komarek, A. & Hebauer, F. 2018. Taxonomic revision of Agraphydrus Régimbart, 1903 I. China and Taiwan (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4452(1): 1–101. DOI: 10.11646/zootaxa.4452.1.1 Reference page.
Minoshima, Y.N., Komarek, A. & Ôhara, M. 2015. A revision of Megagraphydrus Hansen (Coleoptera, Hydrophilidae): synonymization with Agraphydrus Régimbart and description of seven new species. Zootaxa 3930(1): 1–63. DOI: 10.11646/zootaxa.3930.1.1. Reference page. NOTE: New species listed above are directly from here

Agraphydrus is a genus of water scavenger beetle in the family Hydrophilidae represented by 205 described species.[1] It is distributed across the Afrotropical, Australasian, and Indomalayan realms.

Taxonomy

The genus Agraphydrus was described for the first time by Régimbart in 1903.[2]

It currently contains 205 described species, partly thanks to a series revisionary work developed by Albretch Komarek.[3][4][5][6][7]

Description

Small beetles (1.4–4.8 mm), pale/yellowish to dark brown in coloration, eyes not emarginate, labrum exposed, maxillary palps moderately long. Elytra without sutural striae, not laterally explanate; the elytral punctation ranges from very fine and subtle to coarse.

A complete diagnosis was presented by Girón and Short,[1] but also see Komarek and collaborators.[3][4][5][6][7]
Habitat

According to Girón and Short,[1] based on annotations by Komarek and collaborators:[3][4][5][6][7]

Agraphydrus can be found in an extremely broad range of habitats, from rivers, streams and forest pools, to hygropetric environments around waterfalls or seepages over rocks; a few species have been collected in terrestrial habitats by sifting moss and leaves from near water bodies, or in the gravel along the bank of a river; in many cases specimens have been found associated with floating vegetation, mosses and algae.[1]
Species

Agraphydrus abrasus Komarek and Freitag, 2020 [6]
Agraphydrus activus Komarek and Hebauer, 2018 [7]
Agraphydrus acutus Komarek, 2020 [5]
Agraphydrus aethiopicus Komarek, 2020 [5]
Agraphydrus agilis Komarek and Hebauer, 2018 [7]
Agraphydrus albescens (Régimbart, 1903) [8]
Agraphydrus ampullatus Komarek and Freitag, 2020 [6]
Agraphydrus anacaenoides Komarek, 2019 [4]
Agraphydrus anatinus Komarek, 2018 [3]
Agraphydrus andamanicus Komarek, 2018 [3]
Agraphydrus andringitra Komarek, 2020 [5]
Agraphydrus angulatus Komarek, 2019 [4]
Agraphydrus angustatus Komarek, 2020 [5]
Agraphydrus angustipenis Komarek, 2018 [3]
Agraphydrus anhuianus (Hebauer, 2000) [9]
Agraphydrus annapurnensis Komarek, 2018 [3]
Agraphydrus arduus Komarek and Hebauer, 2018 [7]
Agraphydrus ater Komarek, 2018 [3]
Agraphydrus atripalpis Komarek, 2020 [5]
Agraphydrus attenuatus (Hansen, 1999) [10]
Agraphydrus audax Komarek and Hebauer, 2018 [7]
Agraphydrus avita (Hansen, 1997) [11]
Agraphydrus bacchusi Komarek, 2019 [4]
Agraphydrus balkeorum Komarek, 2019 [4]
Agraphydrus batak Komarek and Freitag, 2020 [6]
Agraphydrus bhutanensis Komarek, 2018 [3]
Agraphydrus bicoloratus Komarek, 2020 [5]
Agraphydrus bilardoi Komarek, 2020 [5]
Agraphydrus biltoni Komarek, v [5]
Agraphydrus biprojectus Minoshima, Komarek, and Ôhara, 2015 [12]
Agraphydrus borneensis Komarek, 2019 [4]
Agraphydrus boukali Komarek, 2018 [3]
Agraphydrus brevilobatus Komarek and Freitag, 2020 [6]
Agraphydrus brevipenis Komarek, 2019 [4]
Agraphydrus burmensis Komarek, 2019 [4]
Agraphydrus calvus Komarek and Hebauer, 2018 [7]
Agraphydrus camerunensis Komarek, 2020 [5]
Agraphydrus cantonensis Komarek and Hebauer, 2018 [7]
Agraphydrus carinatulus Komarek, 2019 [4]
Agraphydrus cervus Komarek, 2019 [4]
Agraphydrus ceylonensis Komarek, 2018 [3]
Agraphydrus chinensis Komarek and Hebauer, 2018 [7]
Agraphydrus cinnamum Komarek, 2018 [3]
Agraphydrus clarus Komarek, 2019 [4]
Agraphydrus comes Komarek and Hebauer, 2018 [7]
Agraphydrus communis Komarek, 2018 [3]
Agraphydrus confusus Komarek and Hebauer, 2018 [7]
Agraphydrus congolensis Komarek, 2020 [5]
Agraphydrus conicus Komarek and Hebauer, 2018 [7]
Agraphydrus connexus Komarek and Hebauer, 2018 [7]
Agraphydrus constrictus Komarek, 2018 [3]
Agraphydrus contractus Komarek and Hebauer, 2018 [7]
Agraphydrus coomani (d'Orchymont, 1927) [13]
Agraphydrus coronarius Minoshima, Komarek, and Ôhara, 2015 [12]
Agraphydrus crassipenis Komarek, 2018 [3]
Agraphydrus dapengensis Yang & Jia, 2021 [14]
Agraphydrus decipiens Minoshima, Komarek, and Ôhara, 2015 [12]
Agraphydrus delineatus Komarek, 2019 [4]
Agraphydrus elongatus Ribera, Hernando, and Cieslak, 2019 [15]
Agraphydrus engkari Komarek, 2019 [4]
Agraphydrus excisus Komarek, 2019 [4]
Agraphydrus exedis (d'Orchymont, 1937) [16]
Agraphydrus exiguus Komarek, 2019 [4]
Agraphydrus falcatus Komarek, 2018 [3]
Agraphydrus fasciatus Komarek and Hebauer, 2018 [7]
Agraphydrus fikaceki Komarek and Hebauer, 2018 [7]
Agraphydrus flavescens Komarek, 2020 [5]
Agraphydrus flavipes Komarek, 2020 [5]
Agraphydrus flavonotus Komarek, 2018 [3]
Agraphydrus floresinus Komarek, 2019 [4]
Agraphydrus fontis Komarek, 2020 [5]
Agraphydrus forcipatus Komarek and Hebauer, 2018 [7]
Agraphydrus fortis Komarek, 2018 [3]
Agraphydrus fujianensis Komarek and Hebauer, 2018 [7]
Agraphydrus geminus (d'Orchymont, 1932) [17]
Agraphydrus gereckei Komarek, 2020 [5]
Agraphydrus gilvus Komarek, 2018 [3]
Agraphydrus glaber Komarek, 2018 [3]
Agraphydrus globipenis Komarek and Hebauer, 2018 [7]
Agraphydrus goldschmidti Komarek, 2020 [5]
Agraphydrus gracilipalpis Komarek, 2018 [3]
Agraphydrus hamatus Komarek, 2019 [4]
Agraphydrus hanseni (Satô and Yoshitomi, 2004) [18]
Agraphydrus heinrichi Komarek, 2018 [3]
Agraphydrus helicopter Komarek, 2019 [4]
Agraphydrus hendrichi Komarek, 2019 [4]
Agraphydrus heterochromatus Komarek, 2019 [4]
Agraphydrus hortensis Komarek, 2019 [4]
Agraphydrus hygropetricus Komarek, 2018 [3]
Agraphydrus igneus Komarek and Hebauer, 2018 [7]
Agraphydrus imitans Komarek, 2019 [4]
Agraphydrus indicus (d'Orchymont, 1932) [17]
Agraphydrus inflatus Komarek, 2018 [3]
Agraphydrus infuscatus Komarek, 2019 [4]
Agraphydrus insidiator Minoshima, Komarek, and Ôhara, 2015 [12]
Agraphydrus ishiharai (Matsui, 1994) [19]
Agraphydrus jaechi (Hansen, 1999) [10]
Agraphydrus jankodadai Komarek, 2019 [4]
Agraphydrus jilanzhui Komarek and Hebauer, 2018 [7]
Agraphydrus kallar Komarek, 2018 [3]
Agraphydrus kathapa Komarek, 2019 [4]
Agraphydrus kempi (d'Orchymont, 1922)
Agraphydrus khasiensis Komarek, 2018 [3]
Agraphydrus kodaguensis Komarek, 2018 [3]
Agraphydrus komareki Yang & Jia, 2021 [14]
Agraphydrus laocaiensis Komarek, 2019 [4]
Agraphydrus latus Komarek, 2019 [4]
Agraphydrus longipalpus (Jia, 1998) [20]
Agraphydrus longipenis Komarek and Hebauer, 2018 [7]
Agraphydrus lunaris Komarek, 2019 [4]
Agraphydrus luteilateralis (Minoshima and Fujiwara, 2009) [21]
Agraphydrus madagascarensis Komarek, 2020 [5]
Agraphydrus maehongsonensis Komarek, 2019 [4]
Agraphydrus malayanus (Hebauer, 2000) [9]
Agraphydrus malkini Komarek, 2020 [5]
Agraphydrus manfredjaechi Komarek, 2019 [4]
Agraphydrus masatakai Minoshima, Komarek, and Ôhara, 2015 [12]
Agraphydrus matoposensis Komarek, 2020 [5]
Agraphydrus mazzoldii Komarek, 2019 [4]
Agraphydrus meghalayanus Komarek, 2018 [3]
Agraphydrus microphthalmus Komarek, 2019 [4]
Agraphydrus minutissimus (Kuwert, 1890) [22]
Agraphydrus mirabilis Komarek, 2019 [4]
Agraphydrus montanus Minoshima, Komarek, and Ôhara, 2015 [12]
Agraphydrus muluensis Komarek, 2019 [4]
Agraphydrus musculus Komarek, 2019 [4]
Agraphydrus namthaensis Komarek, 2019 [4]
Agraphydrus nanus Komarek, 2018 [3]
Agraphydrus narusei (Satô, 1960) [23]
Agraphydrus nemorosus Komarek, 2019 [4]
Agraphydrus nepalensis Komarek, 2018 [3]
Agraphydrus niger Komarek, 2018 [3]
Agraphydrus nigroflavus Komarek, 2019 [4]
Agraphydrus obesus Komarek, 2019 [4]
Agraphydrus obscuratus Komarek, 2018 [3]
Agraphydrus obsoletus Komarek, 2018 [3]
Agraphydrus occultus Komarek and Freitag, 2020 [6]
Agraphydrus ogatai Minoshima, 2016 [24]
Agraphydrus orbicularis Komarek, 2019 [4]
Agraphydrus orientalis (d'Orchymont, 1932) [17]
Agraphydrus palawanensis Komarek and Freitag, 2020 [6]
Agraphydrus pallidus Komarek, 2019 [4]
Agraphydrus papuanus Komarek, 2019 [4]
Agraphydrus pauculus (Knisch, 1924) [25]
Agraphydrus pauper Komarek, 2020 [5]
Agraphydrus pelingeni Komarek and Freitag, 2020 [6]
Agraphydrus penangensis Komarek, 2019 [4]
Agraphydrus piceus Komarek, 2019 [4]
Agraphydrus politus (Hansen, 1999) [10]
Agraphydrus praecipuus (d'Orchymont, 1937) [26]
Agraphydrus protentus Komarek, 2018 [3]
Agraphydrus pseudoniger Yang & Jia, 2021 [14]
Agraphydrus pullus Komarek, 2018 [3]
Agraphydrus punctatellus Régimbart, 1903 [2]
Agraphydrus punctulatus Komarek, 2018 [3]
Agraphydrus puzhelongi (Jia, 2010) [27]
Agraphydrus pygmaeus (Knisch, 1924) [25]
Agraphydrus raucus Komarek, 2019 [4]
Agraphydrus reductus Komarek and Hebauer, 2018 [7]
Agraphydrus regularis (Hansen, 1999) [10]
Agraphydrus reticulatus Komarek, 2019 [4]
Agraphydrus reticuliceps Komarek and Hebauer, 2018 [7]
Agraphydrus rhodesiensis Komarek, 2020 [5]
Agraphydrus rhomboideus Komarek, 2019 [4]
Agraphydrus rivalis Komarek, 2020 [5]
Agraphydrus robustus Komarek and Hebauer, 2018 [7]
Agraphydrus rostratus Komarek, 2018 [3]
Agraphydrus rugosus Komarek, 2018 [3]
Agraphydrus sabulosus Yang & Jia, 2021 [14]
Agraphydrus sarawakensis Komarek, 2019 [4]
Agraphydrus schoedli Komarek, 2019 [4]
Agraphydrus schoenmanni Komarek and Hebauer, 2018 [7]
Agraphydrus scintillans Komarek, 2019 [4]
Agraphydrus scutifer Komarek, 2020 [5]
Agraphydrus setifer Komarek and Hebauer, 2018 [7]
Agraphydrus shaverdoae Komarek, 2019 [4]
Agraphydrus siamensis (Hansen, 1999) [10]
Agraphydrus sipekorum Komarek, 2018 [3]
Agraphydrus skalei Komarek, 2019 [4]
Agraphydrus spadix Komarek, 2019 [4]
Agraphydrus spinosus Komarek, 2019 [4]
Agraphydrus splendens Komarek and Hebauer, 2018 [7]
Agraphydrus stagnalis (d'Orchymont, 1937)
Agraphydrus stramineus Komarek, 2019 [4]
Agraphydrus sucineus Komarek, 2019 [4]
Agraphydrus sundaicus Komarek, 2019 [4]
Agraphydrus tamdao Komarek, 2019 [4]
Agraphydrus taprobanensis Komarek, 2018 [3]
Agraphydrus tenuipalpis Komarek and Freitag, 2020 [6]
Agraphydrus thaiensis Minoshima, Komarek, and Ôhara, 2015 [12]
Agraphydrus tristis Komarek, 2019 [4]
Agraphydrus tulipa Komarek, 2019 [4]
Agraphydrus tumidus Komarek, 2020 [5]
Agraphydrus tumulosus Komarek, 2018 [3]
Agraphydrus umbrosus Komarek and Hebauer, 2018 [7]
Agraphydrus uncinatus Komarek and Hebauer, 2018 [7]
Agraphydrus usambaraensis Komarek, 2020 [5]
Agraphydrus uvaensis (Hebauer, 2000) [9]
Agraphydrus vadoni Komarek, 2020 [5]
Agraphydrus variabilis Komarek and Hebauer, 2018 [7]
Agraphydrus vietnamensis Komarek, 2019 [4]
Agraphydrus villiersi (Balfour-Browne, 1958) [28]
Agraphydrus wangmiaoi Komarek and Hebauer, 2018 [7]
Agraphydrus yunnanensis Komarek and Hebauer, 2018 [7]
Agraphydrus zetteli Komarek and Freitag, 2020 [6]

References

Girón, Jennifer C.; Short, Andrew Edward Z. (2021-06-18). "The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species". ZooKeys (1045): 1–236. doi:10.3897/zookeys.1045.63810. ISSN 1313-2970. PMC 8233300. PMID 34228772.
Régimbart, MA (1903). "Coléoptères aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 – Mai 1901)". Annales de la Société Entomologique de France. 72: 1–51.
Komarek, A. (2018). "Taxonomic revision of AgraphydrusRégimbart, 1903 II. The Indian Subcontinent" (PDF). Koleopterologische Rundschau. 88: 103–204.
Komarek, A. (2019). "Taxonomic revision of AgraphydrusRégimbart, 1903 III. Southeast Asia (except Philippines) and Australian Region (Coleoptera: Hydrophilidae: Acidocerinae)" (PDF). Koleopterologische Rundschau. 89: 151–316.
Komarek, A. (2020). "Taxonomic revision of AgraphydrusRégimbart, IV. Africa, Western Asia, and redescription of the genus (Coleoptera: Hydrophilidae: Acidocerinae)" (PDF). Koleopterologische Rundschau. 90: 127–200.
Komarek, A.; Freitag, H. (2020). "Taxonomic revision of AgraphydrusRégimbart, 1903 V. Philippine species and their first DNA barcodes" (PDF). Koleopterologische Rundschau. 90: 201–242.
Komarek, A.; Hebauer, F. (2018-07-30). "Taxonomic revision of Agraphydrus Régimbart, 1903 I. China and Taiwan (Coleoptera: Hydrophilidae: Acidocerinae)". Zootaxa. 4452 (1): 1–101. doi:10.11646/zootaxa.4452.1.1. ISSN 1175-5334. PMID 30314249. S2CID 52975164.
Régimbart, M. (1903). "Coléoptères aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophi-lidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 – Mai 1901)". Annales de la Société Entomologique de France. 72: 1–51.
Hebauer, F. (2000). "The genus Megagraphydrus Hansen, 1999, with description of new species (Coleoptera: Hydrophilidae)". Acta Coleopterologica. 16: 14–22.
Hansen, Michael (1999-01-01). "Fifteen new genera of Hydrophilidae (Coleoptera), with remarks on the generic classification of the family". Insect Systematics & Evolution. 30 (2): 121–172. doi:10.1163/187631200X00228. ISSN 1876-312X.
Hansen, M. (1997). "A new subfamily for a remarkable new genus and species of Hydrophilidae from New Guinea (Coleoptera: Hydrophilidae)". Annales Zoologici. 47: 107–110.
Minoshima, Y. N.; Komarek, A.; Ôhara, M. (2015). "A revision of Megagraphydrus Hansen (Coleoptera, Hydrophilidae): synonymization with Agraphydrus Régimbart and description of seven new species". Zootaxa. 3930: 1–63. PMID 25781810.
d’Orchymont, A. (1927). "Papers on Malayan aquatic biology. V. Notes on the Hydrophilidae in the Federated Malay States Museums". Journal of the Federated Malay States Museums. 13 (4): 246–252.
Yang, Zhen-ming; Jia, Fenglong; Jiang, Lu; Guo, Qiang (2021-05-18). "Four new species of Agraphydrus Régimbart, 1903 with additional faunistic record from China (Coleoptera, Hydrophilidae, Acidocerinae)". Deutsche Entomologische Zeitschrift. 68 (1): 189–205. doi:10.3897/dez.68.66200. ISSN 1860-1324. S2CID 236330170.
Ribera, I.; Hernando, C.; Cieslak, A. (2019). "Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae" (PDF). Acta Entomologica Musei Nationalis Pragae. 59: 253–272. doi:10.2478/aemnp-2019-0021. hdl:10230/44839. S2CID 198381025.
d’Orchymont, A. (1937). "Descriptions of three new Hydrophilidae from India". Records of the Indian Museum. 39: 29–31.
d’Orchymont, A. (1932). "Zur Kenntnis der Kolbenwasserkäfer (Palpicornia) von Sumatra, Java und Bali". Archiv für Hydrobiologie. Supplement Band IX: 623–714. [pl. XIV–XVII.]
Satô, M.; Yoshitomi, H. (2004). "Discovery of a second representative of the genus Horelophopsis (Coleoptera, Hydrophilidae) from the Ryukyu archipelago, Japan". Elytra. 32: 41–49.
Matsui, E. (1994). "Three new species of the genus Enochrus from Japan and Taiwan (Coleoptera: Hydrophilidae)". Transactions of the Shikoku Entomological Society. 20: 215–220.
Jia, F. (1998). "A new genus Pseudopelthydrus gen. n. from Hainan Island, China (Coleoptera: Hydrophilidae: Hydrophilinae)". Chinese Journal of Entomology. 18: 225–230.
Minoshima, Y. N.; Fujiwara, J. (2009). "Discovery of water scavenger beetle genus Megagraphydrus Hansen (Coleoptera, Hydrophilidae) from Japan, with description of a new species" (PDF). Elytra. 37: 53–64.
Kuwert, A. (1890). "Bestimmungs-Tabellen der europäischen Coleopteren. XX. Heft. Hydrophilidae. II. Abteilung: Sphaeridiini und Helophorini". Verhandlungen des naturforschenden Vereins in Brünn. 28: 159–328.
Satô, M. (1960). "One new genus and two new species of the subtribe Helocharae from Japan (Coleoptera: Hydrophilidae)". Transactions of the Shikoku Entomological Society. 6: 76–80.
Minoshima, Yûsuke N. (2016). "Taxonomic review of Agraphydrus from Japan (Coleoptera: Hydrophilidae: Acidocerinae)". Entomological Science. 19 (4): 351–366. doi:10.1111/ens.12213. ISSN 1479-8298. S2CID 89394202.
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Jia, Fenglong (2010-06-08). "Megagraphydrus puzhelongi, sp. n., a new water scavenger beetle from China (Coleoptera: Hydrophilidae: Hydrophilinae)". Zootaxa. 2498 (1): 65. doi:10.11646/zootaxa.2498.1.6. ISSN 1175-5334.
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