Agra

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Adephaga

Familia: Carabidae
Subfamilia: Harpalinae
Supertribus: Lebiitae
Tribus: Lebiini
Subtribus: Agrina
Genus: Agra
Species: A. holzschuhi – A. absurdis – A. aculeata – A. acuspina – A. acutidens – A. acutoides – A. aeneipennis – A. aeneola – A. aeris – A. aeroides – A. alinahui – A. alternata – A. alvarengai – A. amabilis – A. amoena – A. andina – A. andrewesi – A. anguinea – A. anthrax – A. ardoini – A. ariasi – A. arrowi – A. asymetrica – A. aterrima – A. atlas – A. atriperna – A. attenuata – A. aurata – A. aurea – A. aureoviridis – A. auricula – A. aurifera – A. auripennis – A. auronitens – A. aurora – A. aurovittata – A. azurea – A. azureipennis – A. baccii – A. bahiae – A. baleni – A. barrensis – A. basilewskyi – A. bci – A. belize – A. besckei – A. bicolor – A. bicoloripes – A. bicostata – A. biexcavata – A. biexcisa – A. bifaria – A. biolat – A. biseriatella – A. blumax – A. boggianii – A. bogotana – A. boliviana – A. bonsi – A. bonvouloiri – A. bonvouloirii – A. borgmeieri – A. brentoides – A. brevicollis – A. brevicornis – A. brevithorax – A. brunnea – A. brunneipennis – A. brunneitarsis – A. brunneoviridis – A. brunnescens – A. brunnipennis – A. buqueti – A. cachimbana – A. cachimbo – A. cadabra – A. cajennensis – A. calamitas – A. caligata – A. caligo – A. callidoides – A. campana – A. cancellata – A. capitata – A. castanea – A. castaneipes – A. castanoptera – A. catascopoides – A. catbellae – A. catenulata – A. catie – A. cauca – A. cavei – A. ce – A. cephalota – A. chalcea – A. chalcoptera – A. championi – A. chapada – A. chaudoiri – A. chaudoirii – A. chlorocera – A. chocha – A. chryseis – A. chrysis – A. chrysopteryx – A. cicatricosa – A. clavata – A. clavipes – A. cobra – A. cochlearis – A. coerulea – A. colasi – A. coleps – A. columbiana – A. communis – A. confusa – A. conhormigas – A. conicollis – A. constans – A. coptoptera – A. cornicola – A. costaricensis – A. crassicornis – A. crassipes – A. crebrefoveata – A. crebrepunctata – A. cribriceps – A. cribricollis – A. cruciaria – A. csikif – A. csikii – A. cuneolus – A. cuprea – A. cupreola – A. cupripennis – A. cuproptera – A. cyanea – A. cyanescens – A. cyaneucnemes – A. cyanippe – A. cyanosticta – A. cyanoviridis – A. cynthia – A. cytherea – A. dable – A. dafne – A. darlingtoni – A. dation – A. dax – A. delgadoi – A. demerasae – A. denticulata – A. dentipennis – A. diagonalis – A. dimidiata – A. dominula – A. dora – A. dorazul – A. driades – A. dryas – A. duckworlhorum – A. duckworthorum – A. duplicata – A. ecaligis – A. ega – A. elaina – A. eowilsoni – A. eponine – A. erythrocera – A. erythropus – A. eucera – A. eucnemes – A. eurypelma – A. exarata – A. excavata – A. exsculpta – A. fada – A. falcon – A. fallax – A. falsisagax – A. famula – A. feisthameli – A. feisthamelii – A. felix – A. femoralis – A. femorata – A. filicornis – A. filiformis – A. fimbriata – A. flava – A. flavipes – A. foraminosa – A. formicaria – A. fortuna – A. foveella – A. foveigera – A. foveipennis – A. foveolata – A. fryi – A. fugax – A. fulvicauda – A. funebris – A. fuscipes – A. fuseipes – A. gaudiola – A. gemmata – A. geniculata – A. geographica – A. geographiea – A. germaini – A. giesberti – A. glauca – A. goryi – A. gounellei – A. goyasensis – A. goyazella – A. grace – A. gracilis – A. graminea – A. granodeoro – A. gravis – A. guatemalena – A. guyanensis – A. hespenheide – A. hilaris – A. hirta – A. honesta – A. hovorei – A. howdenorum – A. huambana – A. humboldti – A. humeralis – A. hyalina – A. hypsophila – A. ichabod – A. ictina – A. ignobilis – A. imaginis – A. immersa – A. inbio – A. inca – A. incerta – A. incisa – A. infuscata – A. inops – A. inpa – A. insidiosa – A. intermedia – A. invicta – A. iota – A. iquitosana – A. iris – A. itatiaya – A. iycisca – A. janzeni – A. jedlickai – A. jimwappes – A. julie – A. katewinsletae – A. kayae – A. ketschuana – A. klugii – A. lacrymosa – A. laeticolor – A. laetipes – A. lamproptera – A. lata – A. laticeps – A. latifemoris – A. latipes – A. lavernae – A. leprieuri – A. liebkei – A. ligulata – A. lilu – A. limulus – A. lindae – A. littleorum – A. liv – A. lobata – A. longelytrata – A. longicornis – A. longula – A. loricata – A. luctuosa – A. luehea – A. lugubris – A. lutea – A. lycisca – A. macra – A. macracantha – A. macrodera – A. magdalena – A. magnifica – A. maia – A. maiae – A. manu – A. maracay – A. mathani – A. mauritii – A. max – A. maxli – A. maya – A. megaera – A. megera – A. melanogona – A. melanogqna – A. memnon – A. metallescens – A. metallica – A. mexicana – A. mime – A. minasiana – A. minasianus – A. mira – A. mirabilis – A. misella – A. mixta – A. mnemosine – A. mniszechi – A. modesta – A. moerens – A. moesta – A. moira – A. monteverde – A. monticola – A. moritzi – A. moritzii – A. muehlei – A. mulieris – A. multifoveolata – A. multiplicata – A. multipunctata – A. multisetosa – A. mustela – A. negrei – A. nevermanni – A. nex – A. nicki – A. nickiana – A. nigra – A. nigrarima – A. nigrella – A. nigripes – A. nigritarsis – A. nigritula – A. nigriventris – A. nigroaenea – A. nigrocyanea – A. nigrotibiata – A. nigroviridis – A. nodicornis – A. nola – A. not – A. notcatie – A. notichlora – A. notiocyanea – A. notpusilla – A. nova – A. novoaurora – A. nox – A. oblongopunctata – A. obscura – A. obscuripes – A. obscuritibiis – A. occipitalis – A. ocellata – A. ohausi – A. oiapoquensis – A. olfersi – A. olivella – A. olivencana – A. oliviella – A. optima – A. orabrocha – A. orinocensis – A. osculatii – A. othello – A. ovicollis – A. oxyptera – A. pacifica – A. pala – A. pallens – A. pallida – A. pallipes – A. palmata – A. paloma – A. panamensis – A. para – A. paradoxa – A. paratax – A. parumfoveata – A. pavonina – A. pearsoni – A. peccata – A. pehlkei – A. pennyi – A. perexcisa – A. perforata – A. perinvicta – A. perkinsorum – A. perrinae – A. pertzeli – A. peruana – A. phaearlhra – A. phaearthra – A. phaenicodera – A. phaenoptera – A. phaeogena – A. phaeogona – A. phainops – A. phallica – A. phite – A. phoenicodera – A. pia – A. pichincha – A. picipes – A. piligera – A. piranha – A. pitilla – A. plantipedis – A. platyscelis – A. plaumanni – A. plebeja – A. polita – A. prasina – A. prodigiosa – A. proxima – A. pseuderythropus – A. pseudoboliviana – A. pseudolaetipes – A. pseudolutea – A. pseudomoesta – A. pseudopusilla – A. pseudorufitarsis – A. pseudovarians – A. pujoli – A. pulchella – A. pulchra – A. pulla – A. punctata – A. punctatella – A. punctatostriata – A. puncticeps – A. puncticollis – A. punctigera – A. punctulata – A. pupilla – A. purpurea – A. pusilla – A. quadricephala – A. quadrilamata – A. quadriseriata – A. quadrispinosa – A. quararibea – A. quesada – A. raffaellae – A. reflexidens – A. regina – A. regularis – A. relucens – A. resplendens – A. rhomboides – A. risseri – A. rondonia – A. rosea – A. rosettae – A. rotundangula – A. rubra – A. rubricephala – A. rubricollis – A. rubripes – A. rubrocuprea – A. rubrofemorata – A. rubroviolacea – A. rufarima – A. rufescens – A. ruficornis – A. rufipes – A. rufitarsis – A. rufiventris – A. rufoaenea – A. rufonigra – A. rugosostriata – A. rutilipennis – A. sagax – A. sahlbergii – A. saltatrix – A. samiria – A. santarema – A. santarosa – A. saramax – A. sasquatch – A. satipo – A. saundersi – A. saundersii – A. schwarzeneggeri – A. scrobipennis – A. seabrae – A. semifulva – A. semiviridis – A. seriefoveata – A. serra – A. servatorum – A. seticollis – A. setifemoris – A. setigera – A. sexdentata – A. sigillata – A. similis – A. simillima – A. simplex – A. sirena – A. sironyx – A. smaragdina – A. smaragdinipennis – A. smaragdula – A. smurf – A. soccata – A. solanoi – A. solimoes – A. solisi – A. soror – A. sparsepunctata – A. sphenarion – A. spina – A. spinicauda – A. spinipennis – A. spinosa – A. splendida – A. steinbachi – A. sternitica – A. stockwelli – A. strandi – A. strangulata – A. striatifemoris – A. striatopunctata – A. subcoerulea – A. subgemmata – A. sublaevicollis – A. sublaevigata – A. subpallipes – A. subtilicornis – A. subtilis – A. superba – A. suprema – A. surinama – A. suturalis – A. tarapoto – A. tarapotoana – A. tarnieri – A. tenuis – A. terebrata – A. tessera – A. tetraspina – A. thiemei – A. thomsoni – A. tibialis – A. tingo – A. tingomaria – A. tiputini – A. titan – A. titschacki – A. tremolerasi – A. tricarinata – A. tricolor – A. tricuspidata – A. tridentata – A. triseriata – A. tristis – A. trochanterica – A. truquii – A. tubercolata – A. tuitis – A. tumatumari – A. turrialba – A. tuxtlas – A. ubicki – A. urania – A. valentina – A. vanemdeni – A. varceicola – A. variabilis – A. varians – A. variipes – A. varioligera – A. variolosa – A. vate – A. vation – A. venatrix – A. venezuelana – A. venustula – A. vesedes – A. vicina – A. vidua – A. violacea – A. violante – A. virgata – A. viridicollis – A. viridipennis – A. viridipleuris – A. viridipunctata – A. viridisticta – A. viridistriata – A. viridula – A. vulgaris – A. winnie – A. xingu – A. yeti – A. yoda – A. yodella – A. yola – A. zapotal – A. zellibori – A. zischkai – A. zona – A. zumbado – A. zuniga
Name

Agra Fabricius, 1801
Type species: Agra aenea Fabricius, 1801
Fixation: subsequent designation by Erwin (1982)

Synonyms

Agridia Chaudoir, 1861
Type species: Agridia platyscelis Chaudoir, 1861
Fixation: subsequent designation by Erwin (1982)

Agra grace

References

Erwin, T.L. 1978: Systematic, natural history, and zoogeographic notes on the genus Agra Fabricius, with a description of a new species from Panama (Coleoptera: Carabidae: Lebiini). Coleopterists bulletin, 32(4): 261–268. JSTOR
Erwin, T.L. 1982: Agra, arboreal beetles of Neotropical forests: erythropus group systematics (Carabidae). Systematic entomology, 7(1): 39–71. DOI: 10.1111/j.1365-3113.1982.tb00125.x
Erwin, T.L. 1982: Agra, arboreal beetles of Neotropical forests: platyscelis-group systematics (Carabidae). Systematic entomology, 7(2): 185–210. DOI: 10.1111/j.1365-3113.1982.tb00131.x
Erwin, T.L. 1983: Agra, arboreal beetles of Neotropical forests: famula group and formicaria group systematics (Carabidae). Systematic entomology, 8(3): 263–292. DOI: 10.1111/j.1365-3113.1983.tb00482.x
Erwin, T.L. 1984: Agra, arboreal beetles of Neotropical forests: palmata-group systematics (Carabidae). Systematic entomology, 9(1): 9–48. DOI: 10.1111/j.1365-3113.1984.tb00499.x
Erwin, T.L. 1986: Agra, arboreal beetles of Neotropical forests: mixta group, virgata group, and ohausi group systematics (Carabidae). Systematic entomology, 11(3): 293–316. DOI: 10.1111/j.1365-3113.1986.tb00183.x
Erwin, T.L. 1993(1991): Agra, arboreal beetles of Neotropical forests: rufoaenea and quararibea group systematics (Carabidae). Revista peruana de Entomologia, 34: 15–28. [not seen]
Erwin, T.L. 1996: Arboreal beetles of neotropical forests: Agra Fabricius, the cayennensis complex (Coleoptera: Carabidae: Lebiini: Calleidina). Annales zoologici fennici, 33(1): 17–21. PDF
Erwin, T.L. 1998: Evolution at the equator arboreal and alticolous beetles and their taxon pulses with descriptions of a new Agra subclade and its species (Coleoptera: Carabidae: Lebiini). Pp. 491–510 in: Ball, G.E.; Casale, A.; Vigna Taglianti, A. (eds.) Phylogeny and classifi cation of Caraboidea (Coleoptera: Adephaga). Proceedings of a Symposium (28 August, 1996, Florence, Italy), XX International Congress of Entomology Museo Regionale Scienze Naturali - Torino, Atti. Torino. [not seen]
Erwin, T.L. 2000: Arboreal beetles of Neotropical forests: Agra Fabricius, the novaurora complex (Coleoptera: Carabidae: Lebiini: Agrina). Smithsonian contributions to zoology, (608)
Erwin, T.L. 2000: Arboreal beetles of Neotropical forests: Agra Fabricius, a taxonomic supplement for the platyscelis group with new species and distribution records (Coleoptera: Carabidae, Lebiini, Agrina). Coleopterists bulletin, 54(1): 90–119. DOI: 10.1649/0010-065X(2000)054[0090:ABONFA]2.0.CO;2 JSTOR
Erwin, T.L. 2000: Arboreal beetles of Neotropical forests: taxonomic supplement for the Agra virgata and ohausi groups with a new species and additional distribution records (Coleoptera: Carabidae). Coleopterists bulletin, 54(2): 251–262. DOI: 10.1649/0010-065X(2000)054[0251:ABONFT]2.0.CO;2 JSTOR
Erwin, T.L. 2002: The beetle family Carabidae of Costa Rica: twenty-nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina). Zootaxa 119: 1–68. Abstract & excerpt PDF Reference page.
Erwin, T.L. 2010: Agra, arboreal beetles of Neotropical forests: pusilla group and piranha group systematics and notes on their ways of life (Coleoptera, Carabidae, Lebiini, Agrina). ZooKeys, 66: 1–28. ISSN: 1313-2970 (online) ISSN: 1313-2989 (print) DOI: 10.3897/zookeys.66.684
Erwin, T.L.; Pogue, M.G. 1988: Agra, arboreal beetles of Neotropical forests: biogeography and the forest refugium hypothesis (Carabidae). Pp. 161–188 in: Heyer, W.R.; Vanzolini, P.E. (eds.) Neotropical Distribution Patterns: Proceedings of a 1987 Workshop. Proceedings of the Brazilian Academy of Sciences. [not seen]

Vernacular names
English: elegant canopy beetles

Agra is a genus of beetles in the family Carabidae, the ground beetles. There are over 500 described species, but there are well over 1000 specimens in collections that have not yet been described.[1] The common name elegant canopy beetles has been used for genus Agra.[1]

Beetles of this genus have narrow heads and long, constricted necks. The tarsomeres, or the "feet" of the beetle, are wide and equipped with pads, allowing the beetle to grip leaves as it moves about and rests on plants. Females have "telescopic" reproductive organs, which they use to deposit eggs deep in the substrate.[1]

Adults of some species are probably predators of other arthropods. Some have also been seen drinking sap and eating pollen. Adult beetles are nocturnal and attracted to lights. They rest on the undersides of leaves with their bodies aligned along the midrib, legs and antennae tucked in. The larvae develop in burrows and under tree bark, and are likely predatory.[1]

Adult beetles have anti-predator secretions that are offensive to predators; they have been noted to repel bats.[1]

Species include:[2]

Agra absurdis Liebke, 1938
Agra aculeata Chaudoir, 1854
Agra acuspina Straneo, 1979
Agra acutidens Straneo, 1965
Agra acutoides Straneo, 1982
Agra aeneipennis Chaudoir, 1861
Agra aeneola Bates, 1883
Agra aeris Erwin, 2000
Agra aeroides Erwin, 1983
Agra alinahui Erwin, 2000
Agra alternata Klug, 1834
Agra alvarengai Straneo, 1982
Agra amabilis Chaudoir, 1863
Agra amoena Chaudoir, 1861
Agra andina Liebke, 1938
Agra andrewesi Liebke, 1938
Agra anguinea Bates, 1865
Agra anthrax Erwin, 1986
Agra ardoini Straneo, 1982
Agra ariasi Erwin, 1982
Agra arrowi Liebke, 1938
Agra asymetrica Straneo, 1979
Agra aterrima Klug, 1824
Agra atlas Erwin, 1984
Agra atriperna Erwin, 1984
Agra attenuata Klug, 1824
Agra aurata Bates, 1865
Agra aurea Liebke, 1940
Agra aureoviridis Straneo, 1982
Agra auricula Liebke, 1940
Agra aurifera Liebke, 1940
Agra auripennis Liebke, 1940
Agra auronitens Bates, 1883
Agra aurora Liebke, 1940
Agra aurovittata Chaudoir, 1850
Agra azurea Chaudoir, 1861
Agra azureipennis Erwin, 1982
Agra baccii (Straneo, 1958)
Agra bahiae Straneo, 1965
Agra baleni Steinheil, 1875
Agra barrensis Straneo, 1955
Agra basilewskyi Straneo, 1955
Agra bci Erwin, 2000
Agra belize Erwin, 1984
Agra besckei Liebke, 1940
Agra bicolor Straneo, 1979
Agra bicoloripes Straneo, 1955
Agra bicostata Bates, 1865
Agra biexcavata Straneo, 1958
Agra biexcisa Chaudoir, 1866
Agra bifaria Chaudoir, 1866
Agra biolat Erwin, 2000
Agra biseriatella Straneo, 1955
Agra blumax Erwin, 1983
Agra boggianii Straneo, 1955
Agra bogotana Straneo, 1979
Agra boliviana Liebke, 1951
Agra bonsi Straneo, 1979
Agra bonvouloiri Chaudoir, 1866
Agra borgmeieri Liebke, 1940
Agra brentoides Latreille & Dejean, 1823
Agra brevicollis Klug, 1824
Agra brevicornis Bates, 1865
Agra brevithorax Straneo, 1965
Agra brunnea Liebke, 1940
Agra brunneipennis Gory, 1833
Agra brunneitarsis Straneo, 1958
Agra brunneoviridis Straneo, 1979
Agra brunnescens Straneo, 1965
Agra buqueti Gory, 1833
Agra cachimbana Straneo, 1965
Agra cachimbo Erwin, 1984
Agra cadabra Erwin, 1986
Agra cajennensis (Olivier, 1795)
Agra calamitas Erwin, 1986
Agra caligata Liebke 1951
Agra caligo Erwin, 1982
Agra callidoides Straneo, 1979
Agra campana Erwin, 1983
Agra cancellata Dejean 1831
Agra capitata Straneo, 1958
Agra castanea Straneo, 1965
Agra castaneipes Bates, 1883
Agra castanoptera Straneo, 1965
Agra catascopoides Straneo, 1965
Agra catbellae Erwin, 2002
Agra catenulata Klug, 1824
Agra catie Erwin, 2002
Agra cauca Erwin, 1998
Agra cavei Erwin, 1984
Agra ce Erwin, 2010
Agra cephalota Liebke 1938
Agra chalcea Klug, 1834
Agra chalcoptera Klug, 1834
Agra championi Bates, 1883
Agra chapada Erwin, 1987
Agra chaudoiri Bates, 1865
Agra chlorocera Chaudoir, 1863
Agra chocha Erwin, 1986
Agra chrysis Bates, 1865
Agra chrysopteryx Bates, 1878
Agra cicatricosa Chaudoir, 1861
Agra clavata Liebke, 1951
Agra clavipes Klug, 1834
Agra cobra Erwin, 1982
Agra cochlearis Liebke, 1938
Agra coeps Erwin, 1982
Agra coerulea Chaudoir, 1854
Agra colasi Straneo, 1965
Agra columbiana Chaudoir, 1861
Agra communis Straneo, 1965
Agra confusa Chaudoir, 1854
Agra conhormigas Erwin, 2000
Agra conicollis Straneo, 1965
Agra constans Erwin, 1984
Agra coptoptera Chaudoir, 1861
Agra cornicola Liebke, 1940
Agra costaricensis Liebke, 1940
Agra crassicornis Liebke, 1940
Agra crassipes Liebke, 1951
Agra crebrefoveata Straneo, 1965
Agra crebrepunctata Straneo, 1955
Agra cribriceps Straneo, 1965
Agra cribricollis Chaudoir, 1861
Agra cruciaria Erwin, 2010
Agra csikii Liebke, 1940
Agra cuneolus Erwin, 1983
Agra cuprea Klug, 1824
Agra cupreola Chaudoir, 1847
Agra cupripennis Dejean, 1831
Agra cuproptera Liebke, 1940
Agra cyanea Chaudoir, 1861
Agra cyanescens Lucas, 1857
Agra cyaneucnemes Erwin, 1984
Agra cyanippe Bates, 1891
Agra cyanosticta Klug, 1834
Agra cyanoviridis Straneo, 1979
Agra cynthia Buquet, 1835
Agra cytherea J.Thomson, 1857
Agra dable Erwin, 2002
Agra dafne Straneo, 1965
Agra darlingtoni Straneo, 1955
Agra dation Erwin, 1987
Agra dax Erwin, 2000
Agra delgadoi Erwin, 2002
Agra demerasae Erwin, 1984
Agra denticulata Steinheil 1875
Agra dentipennis Liebke, 1951
Agra diagonalis Straneo, 1966
Agra dimidiata Chevrolat 1856
Agra dominula Bates, 1865
Agra dora Erwin, 1984
Agra dorazul Erwin, 1984
Agra driades Straneo, 1965
Agra dryas Erwin, 1982
Agra duckworlhorum Erwin, 2000
Agra duplicata Liebke, 1938
Agra ecaligis Erwin, 1982
Agra ega Erwin, 1982
Agra elaina Bates, 1883
Agra eowilsoni Erwin, 1998
Agra eponine Erwin, 2000
Agra erythrocera Brulle 1837
Agra erythropus Dejean 1825
Agra eucera Erwin, 1984
Agra eucnemes Erwin, 1984
Agra eurypelma Bates, 1883
Agra exarata Klug, 1824
Agra excavata Klug, 1824
Agra exsculpta Liebke, 1951
Agra fada Chevrolat 1856
Agra falcon Erwin, 2000
Agra fallax Liebke, 1951
Agra falsisagax Erwin, 1982
Agra famula Liebke, 1940
Agra feisthameli Buquet, 1835
Agra felix Liebke, 1951
Agra femoralis Chaudoir, 1861
Agra femorata Klug, 1824
Agra filicornis Straneo, 1955
Agra filiformis Dejean, 1831
Agra fimbriata Straneo, 1965
Agra flava Straneo, 1966
Agra flavipes Straneo, 1982
Agra foraminosa Liebke, 1938
Agra forluna Erwin, 1983
Agra formicaria Thomson, 1857
Agra foveella Liebke, 1937
Agra foveigera Chaudoir, 1861
Agra foveipennis Chaudoir, 1866
Agra foveolata Chaudoir, 1850
Agra fryi Chaudoir, 1866
Agra fugax Erwin, 2002
Agra fulvicauda Bates, 1883
Agra funebris Straneo, 1965
Agra fuseipes Straneo, 1979
Agra gaudiola Bates, 1865
Agra gemmata Klug, 1824
Agra geniculata Klug, 1824
Agra geographiea Straneo, 1965
Agra germaini Liebke, 1938
Agra giesberti Erwin, 2002
Agra glauca Liebke, 1951
Agra goryi Chaudoir, 1847
Agra gounellei Liebke, 1938
Agra goyasensis Straneo, 1965
Agra goyazella Erwin, 1984
Agra grace Erwin, 2010
Agra gracilis Lucas 1857
Agra graminea Bates, 1865
Agra granodeoro Erwin, 2002
Agra gravis Straneo, 1966
Agra guatemalena Csiki, 1932
Agra guyanensis (Chaudoir, 1863)
Agra hespenheide Erwin, 2000
Agra hilaris Liebke, 1938
Agra hirta Straneo, 1965
Agra honesta Chaudoir, 1854
Agra hovorei Erwin, 2000
Agra howdenorum Erwin, 1982
Agra huambana Straneo, 1982
Agra humboldti Liebke, 1951
Agra humeralis Straneo, 1966
Agra hyalina Chaudoir, 1861
Agra hypsophila Straneo, 1966
Agra ichabod Erwin, 2002
Agra ictina Bates, 1883
Agra ignobilis Chaudoir, 1872
Agra imaginis Erwin, 1986
Agra immersa Klug, 1824
Agra inbio Erwin, 2000
Agra inca Erwin, 1986
Agra incerta Straneo, 1979
Agra incisa Liebke, 1938
Agra infuscata Klug, 1824
Agra inops Straneo, 1979
Agra inpa Erwin, 1983
Agra insidiosa Bates, 1883
Agra intermedia Straneo, 1958
Agra invicta Erwin, 1982
Agra iota Erwin, 1984
Agra iquitosana Erwin, 1982
Agra iris Liebke, 1938
Agra itatiaya Erwin, 1986
Agra iycisca Buquet 1835
Agra janzeni Erwin, 2002
Agra jedlickai Liebke, 1938
Agra jimwappes Erwin, 2002
Agra julie Erwin, 2002
Agra katewinsletae Erwin, 2002
Agra kayae Erwin, 1984
Agra ketschuana Liebke, 1951
Agra klugii Brulle, 1837
Agra lacrymosa Straneo, 1965
Agra laeticolor Bates, 1878
Agra laetipes Straneo, 1965
Agra lamproptera Chaudoir, 1847
Agra lata Erwin, 1987
Agra laticeps Bates, 1865
Agra latifemoris Straneo, 1982
Agra latipes Chaudoir, 1861
Agra lavernae Erwin, 1978
Agra leprieuri Buquet 1835
Agra liebkei Straneo, 1958
Agra ligulata Liebke, 1940
Agra lilu Erwin, 2000
Agra limulus Erwin, 1982
Agra lindae Erwin, 2000
Agra littleorum Erwin, 1984
Agra liv Erwin, 2002
Agra lobata Straneo, 1982
Agra longelytrata Straneo, 1965
Agra longicornis Liebke 1938
Agra longula Chaudoir, 1866
Agra loricata Liebke 1940
Agra luctuosa Straneo, 1979
Agra luehea Erwin, 1983
Agra lugubris Straneo, 1979
Agra lutea Liebke 1938
Agra macra Steinheil 1875
Agra macracantha Bates, 1883
Agra macrodera Chaudoir, 1866
Agra magdalena Erwin, 1987
Agra magnifica Erwin, 2000
Agra maia Erwin, 2010
Agra manu Erwin, 2000
Agra maracay Erwin, 2000
Agra mathani Straneo, 1965
Agra mauritii Straneo, 1982
Agra max Erwin, 2010
Agra maxli Erwin, 1982
Agra maya Liebke 1951
Agra megaera J,Thomson 1857
Agra melanogqna Chaudoir, 1861
Agra memnon Erwin, 1987
Agra metallescens Chaudoir, 1847
Agra mexicana Buquet 1835
Agra mime Erwin, 2000
Agra minasianus Erwin, 2010
Agra mira Liebke 1938
Agra mirabilis Straneo, 1966
Agra misella Straneo, 1965
Agra mixta Straneo, 1965
Agra mnemosine Straneo, 1965
Agra mniszechi Erwin, 1982
Agra modesta Straneo, 1979
Agra moerens Chaudoir, 1861
Agra moesta Chaudoir, 1861
Agra moira Erwin, 1983
Agra monteverde Erwin, 2002
Agra monticola Liebke, 1938
Agra moritzi Chaudoir, 1861
Agra muehlei Straneo, 1982
Agra mulieris Liebke 1951
Agra multifoveolata Steinheil, 1875
Agra multiplicata Klug, 1824
Agra multipunctata Straneo, 1965
Agra multisetosa Bates, 1883
Agra mustela Bates, 1865
Agra negrei Straneo, 1966
Agra nevermanni Liebke, 1938
Agra nex Erwin, 2000
Agra nicki Schweiger, 1952
Agra nickiana Straneo, 1955
Agra nigra Liebke, 1951
Agra nigrarima Erwin, 1984
Agra nigrella Straneo, 1965
Agra nigripes Chaudoir, 1847
Agra nigritarsis Straneo, 1982
Agra nigritula Straneo, 1955
Agra nigriventris Chaudoir, 1861
Agra nigroaenea Chaudoir, 1854
Agra nigrocyanea Straneo, 1955
Agra nigrotibiata Straneo, 1965
Agra nigroviridis Straneo, 1982
Agra nodicornis Straneo, 1965
Agra nola Erwin, 1986
Agra not Erwin, 2002
Agra notcatie Erwin, 2002
Agra notichlora Erwin, 1984
Agra notiocyanea Erwin, 1984
Agra notpusilla Erwin, 2010
Agra nova Straneo, 1982
Agra novoaurora Erwin, 2000
Agra nox Erwin, 1984
Agra oblongopunctata Chevrolat 1835
Agra obscura Chaudoir, 1861
Agra obscuripes Chaudoir, 1854
Agra obscuritibiis Straneo, 1965
Agra occipitalis Bates, 1865
Agra ocellata Liebke, 1938
Agra ohausi Liebke, 1938
Agra oiapoquensis Erwin, 1982
Agra olfersi Straneo, 1966
Agra olivella Bates, 1883
Agra olivencana Erwin, 1982
Agra optima Bates, 1865
Agra orabrocha Erwin, 1984
Agra orinocensis Erwin, 2000
Agra osculatii Guerin-Meneville, 1855
Agra othello Erwin, 2000
Agra ovicollis Chaudoir, 1861
Agra oxyptera Chaudoir, 1861
Agra pacifica Bates, 1891
Agra pala Liebke, 1937
Agra pallens Lucas 1857
Agra pallida Liebke, 1937
Agra pallipes Liebke, 1938
Agra palmata Steinheil, 1875
Agra paloma Erwin, 1984
Agra panamensis Bates, 1878
Agra para Erwin, 1987
Agra paradoxa Straneo, 1979
Agra paratax Erwin, 2000
Agra parumfoveata Straneo, 1965
Agra pavonina Straneo, 1965
Agra pearsoni Erwin, 1984
Agra peccata Liebke, 1940
Agra pehlkei Liebke, 1938
Agra pennyi Erwin, 1982
Agra perexcisa Straneo, 1965
Agra perforata Liebke, 1938
Agra perinvicta Erwin, 1982
Agra perkinsorum Erwin, 1986
Agra perrinae Straneo, 1982
Agra pertzeli Liebke, 1951
Agra peruana Liebke, 1951
Agra phaearlhra Chaudoir, 1866
Agra phaenicodera (Bates, 1865)
Agra phaenoptera Chaudoir, 1854
Agra phaeogena Bates, 1865
Agra phainops Erwin, 1986
Agra phallica Erwin, 2002
Agra phite Erwin, 1987
Agra pia Liebke, 1940
Agra pichincha Erwin, 2000
Agra picipes Klug, 1834
Agra piligera Straneo, 1982
Agra piranha Erwin, 2010
Agra pitilla Erwin, 2002
Agra plantipedis Liebke, 1951
Agra platyscelis (Chaudoir, 1861)
Agra plaumanni Liebke, 1940
Agra plebeja Chaudoir, 1872
Agra polita Lucas 1857
Agra prasina Liebke, 1940
Agra prodigiosa Liebke, 1938
Agra proxima Straneo, 1982
Agra pseuderythropus Erwin, 1982
Agra pseudoboliviana Straneo, 1965
Agra pseudolaetipes Straneo, 1979
Agra pseudolutea Straneo, 1982
Agra pseudomoesta Straneo, 1965
Agra pseudopusilla Erwin, 2010
Agra pseudorufitarsis Straneo, 1965
Agra pseudovarians Straneo, 1965
Agra pujoli Straneo, 1965
Agra pulchella Chaudoir, 1861
Agra pulchra Straneo, 1982
Agra pulla Chaudoir, 1866
Agra punctata Straneo, 1955
Agra punctatella Straneo, 1958
Agra punctatostriata Chaudoir, 1861
Agra puncticeps Straneo, 1982
Agra puncticollis Dejean, 1825
Agra punctigera Straneo, 1958
Agra punctulata Liebke, 1951
Agra pupilla Liebke, 1940
Agra purpurea Bates, 1883
Agra quadricephala Straneo, 1955
Agra quadrilamata Straneo, 1965
Agra quadriseriata Lansberge, 1866
Agra quadrispinosa Chaudoir, 1872
Agra quararibea Erwin, 1993
Agra quesada Erwin, 2002
Agra raffaellae Straneo, 1982
Agra reflexidens Chaudoir, 1861
Agra regina Liebke 1951
Agra regularis Klug, 1834
Agra relucens Schweiger 1952
Agra resplendens Chaudoir, 1866
Agra rhomboides Erwin, 1982
Agra risseri Erwin, 2010
Agra rondonia Erwin, 2000
Agra rosea Liebke, 1940
Agra rosettae Straneo, 1960
Agra rotundangula Straneo, 1955
Agra rubra Erwin, 1987
Agra rubricephala (Straneo, 1982)
Agra rubricollis (Chaudoir, 1863)
Agra rubripes Straneo, 1955
Agra rubrocuprea Bates, 1865
Agra rubrofemorata Straneo, 1958
Agra rubroviolacea Straneo, 1965
Agra rufarima Erwin, 1984
Agra rufescens Klug, 1824
Agra ruficornis Klug, 1824
Agra rufipes Fabrlcius, 1801
Agra rufitarsis Straneo, 1955
Agra rufiventris Bates, 1883
Agra rufoaenea Chevrolat, 1835
Agra rufonigra Straneo, 1965
Agra rugosostriata Chaudoir, 1854
Agra rutilipennis Laporte De Castelnau, 1835
Agra sagax Liebke, 1940
Agra sahlbergii Chaudoir, 1854
Agra saltatrix Erwin, 1982
Agra samiria Erwin, 2000
Agra santarema Chaudoir, 1866
Agra santarosa Erwin, 2002
Agra saramax Erwin, 1993
Agra sasquatch Erwin, 1982
Agra satipo Erwin, 1984
Agra saundersi Bates, 1865
Agra schwarzeneggeri Erwin, 2002
Agra scrobipennis Chaudoir, 1863
Agra seabrae Erwin, 1982
Agra semifulva Bates, 1883
Agra semiviridis (Straneo, 1968)
Agra seriefoveata Chaudoir, 1877
Agra serra Erwin, 1984
Agra servatorum Erwin, 2000
Agra seticollis Straneo, 1982
Agra setifemoris Straneo, 1982
Agra setigera Liebke, 1938
Agra sexdentata Straneo, 1982
Agra sigillata Liebke, 1940
Agra similis Schweiger, 1952
Agra simillima Straneo, 1965
Agra simplex Liebke, 1940
Agra sirena Erwin, 2002
Agra sironyx Erwin, 1984
Agra smaragdina Chaudoir, 1866
Agra smaragdinipennis (Steinheil, 1875)
Agra smaragdula Liebke, 1938
Agra smurf Erwin, 2000
Agra soccata Bates, 1883
Agra solanoi Erwin, 2002
Agra solimoes Erwin, 2000
Agra solisi Erwin, 2002
Agra soror Chaudoir, 1866
Agra sparsepunctata Straneo, 1982
Agra sphenarion Erwin, 1982
Agra spina Erwin, 1983
Agra spinicauda Straneo, 1982
Agra spinipennis Chaudoir, 1850
Agra spinosa Liebke, 1940
Agra splendida Dejean 1829
Agra steinbachi Liebke, 1951
Agra sternitica Straneo, 1982
Agra stockwelli Erwin, 1984
Agra strandi Liebke, 1940
Agra strangulata Chaudoir, 1863
Agra striatifemoris Liebke, 1940
Agra striatopunctata Chaudoir, 1866
Agra subcoerulea Straneo, 1965
Agra subgemmata Straneo, 1965
Agra sublaevicollis Straneo, 1965
Agra sublaevigata Straneo, 1965
Agra subpallipes Straneo, 1982
Agra subtilicornis Straneo, 1982
Agra subtilis Straneo, 1982
Agra superba Erwin, 2000
Agra suprema Erwin, 2000
Agra surinama Csiki, 1932
Agra suturalis Lucas, 1857
Agra tarapoto Erwin, 1984
Agra tarapotoana Erwin, 1982
Agra tarnieri Chaudoir, 1861
Agra tenuis Chaudoir, 1863
Agra terebrata Straneo, 1966
Agra tessera Erwin, 1983
Agra tetraspina Straneo, 1982
Agra thiemei Liebke, 1940
Agra thomsoni Liebke, 1940
Agra tibialis Chaudoir, 1861
Agra tingo Erwin, 2000
Agra tingomaria Erwin, 1984
Agra tiputini Erwin, 2010
Agra titan Erwin, 1982
Agra titschacki Liebke, 1951
Agra tremolerasi Liebke, 1940
Agra tricarinata Straneo, 1965
Agra tricolor Liebke, 1940
Agra tricuspidata Straneo, 1979
Agra tridentata (Olivier, 1795)
Agra triseriata Chaudoir, 1861
Agra tristis Dejean 1831
Agra trochanterica Straneo, 1965
Agra truquii Chaudoir, 1866
Agra tubercolata Straneo, 1965
Agra tuitis Erwin, 1987
Agra tumatumari Erwin, 1982
Agra turrialba Erwin, 2002
Agra tuxtlas Erwin, 2000
Agra ubicki Erwin, 2002
Agra urania Straneo, 1965
Agra valentina Bates, 1865
Agra vanemdeni Straneo, 1955
Agra varceicola Erwin, 1982
Agra variabilis Straneo, 1982
Agra varians Chaudoir, 1861
Agra variipes Liebke, 1951
Agra varioligera Chaudoir, 1863
Agra variolosa Klug, 1824
Agra vate Erwin, 1986
Agra vation Erwin, 1983
Agra venatrix Liebke, 1951
Agra venezuelana Straneo, 1982
Agra venustula Straneo, 1958
Agra vesedes Erwin, 1984
Agra vicina Chaudoir, 1847
Agra vidua Straneo, 1965
Agra violacea Straneo, 1966
Agra violante Liebke, 1940
Agra virgata Chevrolat, 1856
Agra viridicollis Straneo, 1965
Agra viridipennis Straneo, 1958
Agra viridipleuris Straneo, 1965
Agra viridipunctata Chaudoir, 1854
Agra viridisticta Chaudoir, 1861
Agra viridistriata Schweiger 1952
Agra viridula Straneo, 1965
Agra vulgaris Straneo, 1965
Agra winnie Erwin, 2002
Agra xingu Erwin, 1984
Agra yeti Erwin, 1982
Agra yoda Erwin, 1982
Agra yodella Erwin, 1982
Agra yola Erwin, 2000
Agra zapotal Erwin, 2000
Agra zellibori Straneo, 1966
Agra zischkai Straneo, 1955
Agra zona Erwin, 1983
Agra zumbado Erwin, 2002
Agra zuniga Erwin, 2002

References

Erwin, Terry L. (2010). "Agra, arboreal beetles of Neotropical forests: pusilla group and piranha group systematics and notes on their ways of life (Coleoptera, Carabidae, Lebiini, Agrina)". ZooKeys (66): 1–28. doi:10.3897/zookeys.66.684. PMC 3088417. PMID 21594029.
"Agra Fabricius, 1801". Carabidae of the World. 2011. Retrieved 6 Jul 2011.