Goliathus goliatus

Goliathus goliatus , Photo: Michael Lahanas

Cladus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Subclassis: Pterygota
Divisio: Neoptera
Subdivisio: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea
Familia: Scarabaeidae
Subfamilia: Cetoniinae
Tribus: Goliathini
Genus: Goliathus
Species: Goliathus goliatus

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The Goliath beetles (named after the biblical giant Goliath) are among the largest insects on Earth, if measured in terms of size, bulk and weight. They are members of subfamily Cetoniinae, within the family Scarabaeidae. Goliath beetles can be found in many of Africa's tropical forests, where they feed primarily on tree sap and fruit. Little appears to be known of the larval cycle in the wild, but in captivity, Goliathus beetles have been successfully reared from egg to adult using protein-rich foods such as commercial cat and dog food. Goliath beetles measure from 60–110 millimetres (2.4–4.3 in) for males and 50–80 millimetres (2.0–3.1 in) for females, as adults, and can reach weights of up to 80–100 grams (2.8–3.5 oz) in the larval stage, though the adults are only about half this weight. The females range from a dark brown to silky white, but the males are normally brown/white/black or black/white.


Habitat

All Goliathus species are native to Africa. They are primarily tropical, although one species, G. albosignatus, is localized in the more temperate southeastern portion of Africa. It is in the continent's equatorial rain forests that the Goliath beetles have reached their greatest diversity.

Life cycle
Goliathus giganteus (Goliath Beetle)

Goliathus larvae are somewhat unusual among cetoniine scarabs in that they have a greater need for high protein foods than do those of most other genera. Pellets of dry or soft dog or cat food (buried in the rearing substrate on a regular schedule) provide a suitable diet for Goliathus larvae in captivity. However, a substrate of somewhat moistened, decayed leaves and wood should still be provided in order to create a suitable medium for larval growth. The young stage larvae (1st instar) will eat some of this material. Even under optimum conditions, the larvae take a number of months to mature fully because of the great size they attain. They are capable of growing up to 250 millimetres (9.8 in) in length, and reaching weights in excess of 100 grams (3.5 oz). When maximum size is reached, the larva constructs a rather thin walled, hardened cell of sandy soil in which it will undergo metamorphosis (pupation) to the adult state. Once building of this cocoon is completed, the larva transforms to the pupal stage, which is an intermediate phase between the larval and adult stages. During the pupal duration, the insect's tissues are broken down and re-organized into the form of the adult beetle. Once metamorphosis is complete, the insect sheds its pupal skin and undergoes a period of hibernation as an adult beetle until the dry season ends. When the rains begin, the beetle breaks open its cocoon, locates a mate, and the entire life cycle starts over again. The adult beetles feed on materials rich in sugar, especially tree sap and fruit. Under captive conditions, adults can sometimes live for about a year after emerging from their pupal cells. Longevity in the wild is likely to be shorter on average due to factors such as predators and weather. The adult phase concentrates solely on reproduction, and once this function is performed, the time of the adult beetle is limited, as is true for the vast majority of other insect species.

Description

Goliath beetles, like almost all other beetles, possess a reinforced first pair of wings (called elytra) which act as protective covers for their secondary pair of wings and abdomen. Only the second pair of wings (which are large and membranous) are actually used for flying. When not in use, they are kept completely folded beneath the elytra. Each of the beetle's legs ends in a pair of sharp claws (called tarsi) which provide a strong grip useful for climbing on tree trunks and branches. Males have a Y-shaped horn on the head which is used as a pry bar in battles with other males over feeding sites or mates. Females are without a horn, and instead have a wedge-shaped head which assists in burrowing when they lay eggs. Apart from their massive size, Goliathus beetles are strikingly patterned as well. Prominent markings common to all of the Goliathus species are the sharply contrasting black vertical stripes on the pronotum (thoracic shield).

References

* Gilbert Lachaume: The Beetles of the World, volume 3, Goliathini 1, 1983, Sciences Nat, Venette. [1]

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