Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Vetigastropoda
Ordo: Lepetellida
Superfamilia: Fissurelloidea
Familia: Fissurellidae
Subfamilia: Emarginulinae
Genera: Agariste – Altrix – Arginula – Buchanania – Clathrosepta – Clypidina – Emarginella – Emarginula – Fissurisepta – Laeviemarginula – Laevinesta – Manganesepta – Montfortula – Parmaphorella – Pupillaea – Rimulanax – Rixa – Scelidotoma – Scutus – Stromboli – Tugali – Tugalina – Vacerrena – †Agariste – †Entomella
Name
Emarginulinae Children, 1834
Synonyms
Clypidinidae Golikov & Starobogatov, 1989
Emarginulidae Children, 1834 (original rank)
Fissurellideinae Pilsbry, 1890
References
Links
Emarginulinae in the World Register of Marine Species
The subfamily Emarginulinae, common name keyhole limpets and slit limpets, is a taxonomic subfamily of limpet-like sea snails, marine gastropod molluscs in the family Fissurellidae, the keyhole limpets and slit limpets.[1]
The subfamily consists of the following tribes:[2]
Tribe Emarginulini Children, 1834 - synonyms: Rimulidae, Anton, 1838; Zeidoridae Naef, 1913; Hemitominae Kuroda, Habe & Oyama, 1971; Clypidinidae Golikov & Starobogatov, 1989
Tribe Diodorini Odhner, 1932: this tribe has been raised to the rank of subfamily Diodorinae Odhner, 1932 [3]
Tribe Fissurellideini Pilsbry, 1890
Tribe Scutini Christiaens, 1973
Of the subfamilies in Fissurellidae, the subfamily Emarginulinae is the oldest: its earliest known species date back to the Mesozoic.[4]
The various tribes belong to this subfamily because of the unifying features in their radula. Several tribes were formerlysynonym of subfamilies, because of the differences in their shell. The Emargulini have a caplike shell with a slit in the margin but without an apical opening (foramen). The Scutini have a flat shield-shaped shell with a vague indentation at the back. The other tribes have a caplike shell with a round, oval or triangular apical opening.
The species of this subfamily can be found attached to rocks or coral. They are Herbivores.
Genera
Agariste Monterosato, 1892
Altrix Palmer, 1942
Arginula Palmer, 1937
Buchanania Lesson, 1831
Clathrosepta McLean & Geiger, 1998
Clypidina Gray, 1847
Emarginella Pilsbry, 1891
Emarginula Lamarck, 1801
† Entomella Cossmann, 1888
Fissurisepta Seguenza, 1863
Laeviemarginula Habe, 1953
Laevinesta Pilsbry & McGinty, 1952
Manganesepta McLean & Geiger, 1998
Montfortula Iredale, 1915
† Palaeoloxotoma Hansen, 201
Parmaphorella Strebel, 1907
Pupillaea Gray, 1835
Rixa Iredale, 1924
Scelidotoma McLean, 1966
Scutus Montfort, 1810
Stromboli Berry, 1954
Tugali Gray, 1843
Tugalina Habe, 1953
Vacerrena Iredale, 1958
Taxa inquirenda
Clypidella Swainson, 1840
Rimulanax Iredale, 1924 (possible synonym of Puncturella)
Synonyms
Aviscutum Iredale, 1940synonym of Scutus Montfort, 1810
Entomella Cotton, 1945synonym of Emarginula Lamarck, 1801 (invalid: junior homonym of Entomella Cossmann, 1888; Notomella is a replacement name)
† Loxotoma P. Fischer, 1885 synonym of † Palaeoloxotoma Hansen, 2019 (invalid: junior homonym of Loxotoma Zeller, 1854 [Lepidoptera]; Palaeoloxotoma is a replacement name)
Nannoscutum Iredale, 1937synonym of Scutus Montfort, 1810
Notomella Cotton, 1957synonym of Emarginula Lamarck, 1801
Parmophoridea Wenz, 1938synonym of Parmaphorella Strebel, 1907 (unnecessary replacement name)
Parmophorus Blainville, 1817synonym of Scutus Montfort, 1810 (unnecessary substitute name for Scutus)
Plagiorhytis P. Fischer, 1885synonym of Montfortula Iredale, 1915 (Invalid: junior homonym of Plagiorhytis Chaudoir, 1848)
Rimularia Bronn, 1838synonym of Rimula Defrance, 1827 (unjustified emendation of Rimula Defrance, 1827)
Scutum P. Fischer, 1885synonym of Scutus Montfort, 1810 (invalid: unjustified emendation of Scutus)
Semperia Crosse, 1867synonym of Emarginula Lamarck, 1801
Subzeidora Iredale, 1924synonym of Emarginula (Subzeidora) Iredale, 1924 represented as Emarginula Lamarck, 1801
Tugalia Gray, 1857synonym of Tugali Gray, 1843
Vacerra Iredale, 1924synonym of Vacerrena Iredale, 1958 (invalid: junior homonym of Vacerra Godman, 1900 [Lepidoptera];Vacerrena is a replacement name)
Variegemarginula McLean, 2011synonym of Montfortula Iredale, 1915
References
Bouchet, P.; Gofas, S. (2012). Emarginulinae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=224510 on 2013-01-01
Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)
Aktipis S.W., Boehm E. & Giribet G. (2011) Another step towards understanding the slit-limpets (Fissurellidae, Fissurelloidea, Vetigastropoda, Gastropoda): a combined five-gene molecular phylogeny. Zoologica Scripta 40: 238-259
McLean, James H.; R.N. Kilburn (September 1986). "Propodial elaboration in Southern African and Indian Ocean Fissurellidae (Mollusca, Prosobranchia) with description of two new genera and one new species" (PDF). Contributions in Science. 379: 1–12. doi:10.5962/p.208123. S2CID 135095660. Archived from the original (PDF) on 2010-07-21. Retrieved 2010-07-19.
Children J.G. (1834). [Mollusca]. pp. 88–118, in: Synopsis of the contents of the British Museum, ed. 28: 88–118. [Authorship attributed to Children after G. Steiner & A.R. Kabat, 2001, Zoosystema, 23(3): 454-45]
Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1–2): 1-526
External links
Pilsbry, H. A. (1890-1891). Manual of conchology, structural and systematic, with illustrations of the species. (1)12: Stomatellidae, Scissurellidae, Pleurotomariidae, Haliotidae, Scutellinidae, Addisoniidae, Cocculinidae, Fissurellidae, pp. 1-192, pls 1-45 [1890], 193-323, pls 46-65 [1891]. Philadelphia, Conchological Section, Academy of Natural Sciences
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