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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Caenogastropoda
Ordo: Neogastropoda
Superfamilia: Conoidea

Familia: Drilliidae
Genus: Drillia


Drillia is a genus of small sea snails, marine gastropod mollusks in the family Drilliidae.[1][2]
General characteristics

The shell is turriculated, with an elevated spire, longitudinal ribs, and usually revolving striae. The spiral sculpture of the shell is rather smooth or shows a wide spacing of incised lines. The body whorl is usually short. The columella has a posterior callus. The outer lip is thick, but not varicose, nor dentate within, flexuous, with a well-marked posterior anal sinus near (but not reaching) the suture, and an anterior constriction or sinuosity. The siphonal canal is short, curved and usually narrow. Some species show an interrupted pink banding.

The animal has tentacles approaching at their bases and eyes near their extremities.[3]
Species

Species within the genus Drillia include:

Drillia acapulcana (Lowe, 1935)[4]
Drillia adusta (G.B. I Sowerby, 1834)
† Drillia aegyptiaca Cossmann, 1901
Drillia aerope (Dall, 1919)[5]
Drillia albicostata (Sowerby I, 1834)[6]
Drillia albomaculata (Adams C. B., 1845)[7]
Drillia alcyonea Melvill & Standen, 1901[8]
† Drillia allionii Bellardi, 1875
Drillia altispira Sysoev, 1996[9]
Drillia amblytera (Bush, 1893)[10]
† Drillia ammoni Cossmann & Pissarro, 1900
Drillia angolensis Odhner, 1923[11]
Drillia annielonae Nolf & Verstraeten, 2007[12]
Drillia aquatilis (L.A. Reeve, 1845)
Drillia armilla Barnard, 1958[13]
Drillia asra Thiele, 1925[14]
Drillia athyrma Melvill, J.C. & R. Standen, 1901
Drillia audax Melvill & Standen, 1903[15]
Drillia ballista Von Maltzan, 1883[16]
Drillia barkliensis H. Adams, 1869 [17]
† Drillia bonneti Cossmann, 1900 [18]
† Drillia boussaci Cossmann, 1913 [19]
† Drillia brocchii (Bellardi & Michelotti, 1841)[20]
Drillia bruchia Barnard, 1958[21]
Drillia bruuni Knudsen, 1952[22]
Drillia caffra (Smith E. A., 1882)[23]
† Drillia calvimontensis Cossman, 1889 [24]
Drillia capta E.A. Smith, 1899
Drillia captiva E.A. Smith, 1899
Drillia cecchii Jousseaume, 1891[25]
Drillia clionellaeformis (Weinkauff & Kobelt, 1875)
Drillia collina Barnard, 1958
Drillia concolor Smith E.A., 1877
Drillia connelli Kilburn, 1988
† Drillia constantinensis Cossmann & Pissarro, 1900 [26]
Drillia corusca Reeve, 1843
Drillia cratista Melvill J.C., 1927
Drillia cunninghamae McLean & Poorman, 1971[27]
† Drillia cureti Cossmann, 1919 [28]
Drillia cuyoensis Bozzetti, 2020
Drillia dakarensis Knudsen, 1956[29]
Drillia dejecta Smith E.A., 1888
Drillia diasi Barnard, 1958[30]
Drillia dives Melvill, J.C. & R. Standen, 1903
† Drillia douvillei Basse, 1932 [31]
Drillia dovyalis Barnard, 1969[32]
Drillia dunkeri (Weinkauff, 1876)[33]
Drillia eborea A.A. Gould, 1860
† Drillia edulcorata Cossmann & Pissarro, 1900 [26]
Drillia enna (Dall, 1918)
Drillia erepta Barnard, 1969
Drillia euchroes Melvill, J.C., 1912
† Drillia ferenuda Cossmann, 1900 [18]
Drillia fijiensis E.A. Smith, 1888
Drillia fraga H.B. Preston, 1908
Drillia ghyooti Nolf, 2008[34]
Drillia gibberulus (Hervier, 1896)[35]
Drillia griffithii (Reeve, 1843)[36]
Drillia havanensis (Dall, 1881)[37]
Drillia idalinae Bernard & Nicolay, 1984[38]
Drillia inchoata (Sturany, R., 1903)
Drillia indra Thiele, 1925[39]
Drillia inexspectata K. Martin, 1895
Drillia inornata McLean & Poorman, 1971[40]
Drillia investigatoris Smith E. A., 1899[41]
† Drillia ischnocolpa Cossmann & Pissarro, 1900 [26]
Drillia janseni H. Strebel, 1905
Drillia katiae Nolf, 2006[42]
Drillia knudseni Tippett, 2006[43]
Drillia kophameli Strebel, 1905[44]
Drillia latisulcus Barnard, 1958
Drillia lea Thiele, 1925[45]
† Drillia leptoides (de Boury, 1899) [46]
Drillia levanderi R. Sturany, 1905
Drillia lignaria (Sowerby III, 1903)[47]
Drillia macilenta (Melvill, 1923)[48]
Drillia macleani Tucker J., 1992[49]
Drillia maculomarginata Kilburn & Stahlschmidt, 2012
Drillia meridiana Perugia & Prelle, 2012
Drillia monodi Knudsen, 1952[50]
Drillia nodilirata E.A. Smith, 1877
Drillia ochroleuca J.C. Melvill & E.R. Sykes, 1897
Drillia oleacina (Dall, 1881)[51]
Drillia oliverai Kilburn & Stahlschmidt, 2012
Drillia omanensis Melvill, J.C. & R. Standen, 1901
Drillia patriciae Bernard & Nicolay, 1984[52]
Drillia philotima J.C. Melvill & R. Standen, 1903
Drillia phymaticus R.B. Watson, 1885
Drillia poecila Sysoev & Bouchet, 2001[53]
Drillia potti (R. Sturany, 1903)
Drillia prattii E.A. Smith, 1877
Drillia pselia Barnard, 1958
Drillia pyramidata (Kiener, 1840)[54]
Drillia quadrasi O. Böttger, 1895
Drillia recordata (E.R. Sykes, 1905)
Drillia regia (Habe & Murakami, 1970)
Drillia rimata E.A. Smith, 1888
Drillia rosacea (Reeve, 1845)[55]
Drillia roseola (Hertlein & Strong, 1955)[56]
Drillia rosolina (Marrat, 1877)[57]
Drillia rubrozonata M.M. Schepman, 1913
Drillia rufescens R.W. Dunker, 1871
Drillia sesquitertia E.C. Von Martens, 1904
Drillia siebenrocki (Sturany, 1900)
Drillia sikesi H.B. Preston, 1908
Drillia sinuosa (Montagu, 1803)
Drillia sowerbyi W.H. Turton, 1932
Drillia spirostachys Kilburn, 1988[58]
Drillia suxdorfi Strebel, 1905
† Drillia terebra (Basterot, 1825)
Drillia tholos Barnard, 1958
Drillia tripter Von Maltzan, 1883[59]
Drillia tumida McLean & Poorman, 1971[60]
† Drillia turrella (Lamarck, 1804) [61]
Drillia umbilicata Gray, 1838[62]
Drillia valida McLean & Poorman, 1971[63]
Drillia worthingtoni E.A. Smith, 1904

Nomen dubium

Drillia signa Bartsch, 1915

Species inquirenda

Drillia ganjamensis Preston, 1910
Drillia mariesi Melvill & Standen, 1897

Nomen nudum

Drillia alata H. Adams & A. Adams, 1853

Species brought into synonymy
This is a long list of synonyms of Drillia
References

Drillia Gray, 1838. Retrieved through: World Register of Marine Species on 8 July 2012.
P. Bouchet; Yu. I. Kantor; A. Sysoev; N. Puillandre (2011). "A new operational classification of the Conoidea (Gastropoda)". Journal of Molluscan Studies. 77 (3): 273–308. doi:10.1093/mollus/eyr017.
George Washington Tryon, Manual of Conchology vol. VI p. 155; 1884
Drillia acapulcana (Lowe, 1935). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia aerope (Dall, 1919). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia albicostata (Sowerby I, 1834). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia albomaculata (Adams C. B., 1845). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia alcyonea Melvill & Standen, 1901. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia altispira Sysoev, 1996. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia amblytera (Bush, 1893). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia angolensis Odhner, 1923. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia annielonae Nolf & Verstraeten, 2007. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia armilla Barnard, 1958. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia asra Thiele, 1925. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia audax Melvill & Standen, 1903. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia ballista Von Maltzan, 1883. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia barkliensis. Retrieved through: World Register of Marine Species on 13 April 2010.
Cossmann (M.), 1900 Faune pliocènique de Karikal (Inde française). Journal de Conchyliologie, t. 48, vol. 1, p. 14-66
Cossmann (M.), 1913 Étude comparative de fossiles miocéniques recueillis à la Martinique et à l'Isthme de Panama. Journal de Conchyliologie, t. 61, vol. 1, p. 1-64
Cossmann (M.), 1896 Essais de Paléoconchologie comparée (2ème livraison), p. 1-179
Drillia bruchia Barnard, 1958. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia bruuni Knudsen, 1952. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia caffra (Smith E. A., 1882). Retrieved through: World Register of Marine Species on 13 April 2010.
Holotype at MNHN, Paris
Drillia cecchii Jousseaume, 1891. Retrieved through: World Register of Marine Species on 13 April 2010.
Cossmann (M.) & Pissarro (G.), 1900 Faune éocènique du Cotentin. 1er article. Bulletin de la Société Géologique de Normandie, t. 19, p. 19-75
Drillia cunninghamae McLean & Poorman, 1971. Retrieved through: World Register of Marine Species on 13 April 2010.
Cossmann (M.), 1919 - Supplément aux mollusques éocèniques de la Loire-Inférieure. Bulletin de la Société des Sciences Naturelles de l'Ouest de la France, sér. 3, t. 5, p. 53-141
Drillia dakarensis Knudsen, 1956. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia diasi Barnard, 1958. Retrieved through: World Register of Marine Species on 13 April 2010.
Basse (E.), 1932 - Paléontologie de Madagascar. XVIII: Faune malacologique du Crétacé supérieur du Sud-Ouest de Madagascar. Annales de Paléontologie, t. 21, p. 91-168
Drillia dovyalis Barnard, 1969. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia dunkeri Knudsen, 1952. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia ghyooti Nolf, 2008. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia gibberulus (Hervier, 1896). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia griffithii (Reeve, 1843). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia havanensis (Dall, 1881). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia idalinae Bernard & Nicolay, 1984. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia indra Thiele, 1925. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia inornata McLean & Poorman, 1971. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia investigatoris Smith E. A., 1899. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia katiae Nolf, 2006. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia knudseni Tippett, 2006. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia kophameli Strebel, 1905. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia lea Thiele, 1925. Retrieved through: World Register of Marine Species on 13 April 2010.
Cossmann (M.) & Pissarro (G.), 1913 Iconographie complète des coquilles fossiles de l'Éocène des environs de Paris, t. 2, p. pl. 46-65
Drillia lignaria (Sowerby III, 1903). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia macilenta (Melvill, 1923). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia macleani Tucker J., 1992. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia monodi Knudsen, 1952. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia oleacina (Dall, 1881). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia patriciae Bernard & Nicolay, 1984. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia poecila Sysoev & Bouchet, 2001. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia pyramidata (Kiener, 1840). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia rosacea (Reeve, 1845). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia roseola (Hertlein & Strong, 1955). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia rosolina (Marrat, 1877). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia spirostachys Kilburn, 1988. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia tripter Von Maltzan, 1883. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia tumida McLean & Poorman, 1971. Retrieved through: World Register of Marine Species on 13 April 2010.
(M.) & Pissarro (G.), 1913 - Iconographie complète des coquilles fossiles de l'Éocène des environs de Paris, t. 2, p. pl. 46-65
Drillia umbilicata Gray, 1838. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia valida McLean & Poorman, 1971. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia actinocycla Dall & Simpson, 1901. Retrieved through: World Register of Marine Species on 13 April 2010.
Dall, W.H. (1890b) Contributions to the Tertiary fauna of Florida with especial reference to the Miocene silex-beds of Tampa and the Pliocene beds of the Caloosahatchie River. Part I. Pulmonate, opisthobranchiate and orthodont gastropods. Transactions of the Wagner Free Institute of Science, 3, 1–200
Drillia bealiana (Schwengel & McGinty, 1942). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia blacki (Petuch, 2004). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia blakensis Tippett, 2007. Retrieved through: World Register of Marine Species on 13 April 2010.
Cossmann (M.) & Pissarro (G.), 1913 Iconographie complète des coquilles fossiles de l'Éocène des environs de Paris, t. 2, p. pl. 46-65
Drillia cydia (Bartsch, 1943). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia detecta (Dall, 1881). Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia dolorosa Thiele, 1925. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia echinata. Retrieved through: World Register of Marine Species on 13 April 2010.
Drillia enae (Bartsch, 1934). Retrieved through: World Register of Marine Species on 13 April 2010.
Cossmann (M.), 1902 Mollusques éocèniques de la Loire-Inférieure. tome 2, fascicule 2. Bulletin de la Société des Sciences naturelles de l'Ouest de la France, sér. 2, t. 2 , vol. 1, p. 5-159
Drillia flavidula. Retrieved through: World Register of Marine Species on 13 April 2010.
Cossmann (M.) & Pissarro (G.), 1900 - Faune éocènique du Cotentin. 1er article. Bulletin de la Société Géologique de Normandie, t. 19, p. 19-75
Drillia incerta (Smith E. A., 1877). Retrieved through: World Register of Marine Species on 13 April 2010.
Tucker, J.K. 2004 Catalog of recent and fossil turrids (Mollusca: Gastropoda). Zootaxa 682:1-1295
Cossmann (M.), 1896 Mollusques éocèniques de la Loire-Inférieure. Tome 1, fascicule 2. Bulletin de la Société des Sciences Naturelles de l'Ouest de la France, t. 6, vol. 4, p. 180-246
Drillia nicklesi Knudsen, 1956. Retrieved through: World Register of Marine Species on 13 April 2010.
Cossmann (M.), 1896 - Mollusques éocèniques de la Loire-Inférieure. Tome 1, fascicule 2. Bulletin de la Société des Sciences Naturelles de l'Ouest de la France, t. 6, vol. 4, p. 180-246

Drillia wolfei Tippett, 1995. Retrieved through: World Register of Marine Species on 13 April 2010.

Cossmann (M.), 1896 Mollusques éocèniques de la Loire-Inférieure. Tome 1, fascicule 2. Bulletin de la Société des Sciences Naturelles de l'Ouest de la France, t. 6, vol. 4, p. 180-246
Cossmann (M.), 1896 Essais de Paléoconchologie comparée (2ème livraison), p. 1-179
Cossmann (M.), 1919 Supplément aux mollusques éocèniques de la Loire-Inférieure. Bulletin de la Société des Sciences Naturelles de l'Ouest de la France, sér. 3, t. 5, p. 53-141
Lamy, Ed., 1934. Coquilles marines recueillies par M. E. Aubert de Rüe dans l'Amérique du Sud. Bulletin du Muséum d'Histoire naturelle 6: 432-435
Vaught, K.C. (1989). A classification of the living Mollusca. American Malacologists: Melbourne, FL (USA). ISBN 0-915826-22-4. XII, 195 pp.
Tucker, J.K. 2004 Catalog of recent and fossil turrids (Mollusca: Gastropoda). Zootaxa 682:1-1295.
Tippett, D.L. (2006), Taxonomic notes on some Indo-Pacific and West African Drillia species (Conoidea: Drilliidae) = Notas taxonómicas sobre unas especies de Drillia del Indo-Pacifico y África occidental (Conoidea: Drilliidae); Iberus: Revista de la Sociedad Espänola de Malacología 24(1): 13-21

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