Heptapteridae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Osteichthyes
Classis: Actinopterygii
Subclassis: Neopterygii
Infraclassis: Teleostei
Superordo: Ostariophysi
Ordo: Siluriformes
Familia: Heptapteridae
Genera: Acentronichthys - Brachyglanis - Brachyrhamdia - Cetopsorhamdia - Chasmocranus - Gladioglanis - Goeldiella - Heptapterus - Horiomyzon - Imparales - Imparfinis - Leptorhamdia - Mastiglanis - Myoglanis - Nannoglanis - Nemuroglanis - Pariolius - Phenacorhamdia - Phreatobius - Pimelodella - Rhamdella - Rhamdia - Rhamdioglanis - Rhamdiopsis - Taunayia

References

* Froese, R. and D. Pauly. Editors. 2006. FishBase, version (02/2006). [1]

Heptapteridae is a family of catfish that originate from Mexico to South America.[2] Heptapteridae is derived from Greek, hepta meaning seven and pteron meaning fin.[3]

The diversity of this family is poorly known and there are many species yet to be described.[2] So far some 190 species have been described. This family is equivalent to the previously recognized Rhamdiinae, a subfamily of the family Pimelodidae.[2] However, molecular evidence shows that this family is a part of the superfamily Pimelodoidea along with Pimelodidae, Pseudopimelodidae, and Conorhynchos.[4]

The skin of these fish is usually naked (scaleless). They exhibit three pairs of barbels. They have a large adipose fin, and their caudal fin is deeply forked. However, there are no external characteristics that are unique to this family that allow it to be differentiated from Pimelodidae.[2]

Heptapteridae includes a few troglobitic species in the genera Pimelodella, Rhamdia, and Taunayia, as well as in an undescribed genus.[5]

Species

There are 189 species in 24 genera:[6]

* Genus Acentronichthys Eigenmann & Eigenmann, 1889.
o Acentronichthys leptos Eigenmann & Eigenmann, 1889.
* Genus Brachyglanis Eigenmann, 1912.
o Brachyglanis frenatus Eigenmann, 1912.
o Brachyglanis magoi Fernández-Yépez, 1967.
o Brachyglanis melas Eigenmann, 1912.
o Brachyglanis microphthalmus Bizerril, 1991.
o Brachyglanis nocturnus Myers, 1928.
o Brachyglanis phalacra Eigenmann, 1912.
* Genus Brachyrhamdia Myers, 1927.
o Brachyrhamdia heteropleura (Eigenmann, 1912).
o Brachyrhamdia imitator Myers, 1927.
o Brachyrhamdia marthae Sands & Black, 1985.
o Brachyrhamdia meesi Sands & Black, 1985.
o Brachyrhamdia rambarrani (Axelrod & Burgess, 1987).
* Genus Cetopsorhamdia Eigenmann & Fisher, 1916.
o Cetopsorhamdia boquillae Eigenmann & Fisher, 1922.
o Cetopsorhamdia filamentosa Fowler, 1945.
o Cetopsorhamdia iheringi Schubart & Gomes, 1959.
o Cetopsorhamdia insidiosa (Steindachner, 1915).
o Cetopsorhamdia molinae Miles, 1943.
o Cetopsorhamdia nasus Eigenmann & Fisher, 1916.
o Cetopsorhamdia orinoco Schultz, 1944.
o Cetopsorhamdia phantasia Stewart, 1985.
o Cetopsorhamdia picklei Schultz, 1944.
* Genus Chasmocranus Eigenmann, 1912.
o Chasmocranus brachynemus Gomes & Schubart, 1958.
o Chasmocranus brevior Eigenmann, 1912.
o Chasmocranus chimantanus Inger, 1956.
o Chasmocranus longior Eigenmann, 1912.
o Chasmocranus lopezi Miranda Ribeiro, 1968.
o Chasmocranus peruanus Eigenmann & Pearson, 1942.
o Chasmocranus quadrizonatus Pearson, 1937.
o Chasmocranus rosae Eigenmann, 1919.
o Chasmocranus surinamensis (Bleeker, 1862).
o Chasmocranus truncatorostris Borodin, 1927.
* Genus Gladioglanis Ferraris & Mago-Leccia, 1989.
o Gladioglanis conquistador Lundberg, Bornbusch & Mago-Leccia, 1991.
o Gladioglanis machadoi Ferraris & Mago-Leccia, 1989.
* Genus Goeldiella Eigenmann & Norris, 1900.
o Goeldiella eques (Müller & Troschel, 1848).
* Genus Heptapterus Bleeker, 1858.
o Heptapterus bleekeri Boeseman, 1953.
o Heptapterus fissipinnis Miranda Ribeiro, 1911.
o Heptapterus multiradiatus Ihering, 1907.
o Heptapterus mustelinus (Valenciennes, 1835).
o Heptapterus stewarti Haseman, 1911.
o Heptapterus sympterygium Buckup, 1988.
o Heptapterus tapanahoniensis Mees, 1967.
o Heptapterus tenuis Mees, 1986.
* Genus Horiomyzon Stewart, 1986.
o Horiomyzon retropinnatus Stewart, 1986.
* Genus Imparfinis Eigenmann & Norris, 1900.
o Imparfinis borodini Mees & Cala, 1989.
o Imparfinis cochabambae (Fowler, 1940).
o Imparfinis guttatus (Pearson, 1924).
o Imparfinis hasemani Steindachner, 1915.
o Imparfinis hollandi Haseman, 1911.
o Imparfinis lineatus (Bussing, 1970).
o Imparfinis longicauda (Boulenger, 1887).
o Imparfinis microps Eigenmann & Fisher, 1916.
o Imparfinis minutus (Lütken, 1874).
o Imparfinis mirini Haseman, 1911.
o Imparfinis mishky Almiron, Casciotta, Bechara et al. 2007.[7]
o Imparfinis nemacheir (Eigenmann & Fisher, 1916).
o Imparfinis pijpersi (Hoedeman, 1961).
o Imparfinis piperatus Eigenmann & Norris, 1900.
o Imparfinis pristos Mees & Cala, 1989.
o Imparfinis pseudonemacheir Mees & Cala, 1989.
o Imparfinis schubarti (Gomes, 1956).
o Imparfinis spurrellii (Regan, 1913).
o Imparfinis stictonotus (Fowler, 1940).
* Genus Leptorhamdia Eigenmann, 1918.
o Leptorhamdia essequibensis (Eigenmann, 1912).
o Leptorhamdia marmorata Myers, 1928.
o Leptorhamdia schultzi (Miranda Ribeiro, 1964).
* Genus Mastiglanis Bockmann, 1994.
o Mastiglanis asopos Bockmann, 1994.
* Genus Myoglanis Eigenmann, 1912.
o Myoglanis aspredinoides Do Nascimiento & Lundberg, 2005.
o Myoglanis koepckei Chang, 1999.
o Myoglanis potaroensis Eigenmann, 1912.
* Genus Nannoglanis Boulenger, 1887.
o Nannoglanis fasciatus Boulenger, 1887.
* Genus Nemuroglanis Eigenmann & Eigenmann, 1889.
o Nemuroglanis furcatus Ribeiro, Pedroza & Py-Daniel, 2011[8]
o Nemuroglanis lanceolatus Eigenmann & Eigenmann, 1889.
o Nemuroglanis mariai (Schultz, 1944).
o Nemuroglanis pauciradiatus Ferraris, 1988.
* Genus Pariolius Cope, 1872.
o Pariolius armillatus Cope, 1872.
* Genus Phenacorhamdia Dahl, 1961.
o Phenacorhamdia anisura (Mees, 1987).
o Phenacorhamdia boliviana (Pearson, 1924).
o Phenacorhamdia hoehnei (Miranda Ribeiro, 1914).
o Phenacorhamdia macarenensis Dahl, 1961.
o Phenacorhamdia nigrolineata Zarske, 1998.
o Phenacorhamdia somnians (Mees, 1974).
o Phenacorhamdia tenebrosa (Schubart, 1964).
o Phenacorhamdia unifasciata Britski, 1993.
* Genus Pimelodella Eigenmann & Eigenmann, 1888.
o Pimelodella altipinnis (Steindachner, 1864).
o Pimelodella australis Eigenmann, 1917.
o Pimelodella avanhandavae Eigenmann, 1917.
o Pimelodella boliviana Eigenmann, 1917.
o Pimelodella boschmai Van der Stigchel, 1964.
o Pimelodella brasiliensis (Steindachner, 1877).
o Pimelodella breviceps (Kner, 1857).
o Pimelodella buckleyi (Boulenger, 1887).
o Pimelodella chagresi (Steindachner, 1876).
o Pimelodella chaparae Fowler, 1940.
o Pimelodella conquetaensis Ahl, 1925.
o Pimelodella cristata (Müller & Troschel, 1848).
o Pimelodella cruxenti Fernández-Yépez, 1950.
o Pimelodella cyanostigma (Cope, 1870).
o Pimelodella dorseyi Fowler, 1941.
o Pimelodella eigenmanni (Boulenger, 1891).
o Pimelodella eigenmanniorum (Miranda Ribeiro, 1911).
o Pimelodella elongata (Günther, 1860).
o Pimelodella enochi Fowler, 1941.
o Pimelodella eutaenia Regan, 1913.
o Pimelodella figueroai Dahl, 1961.
o Pimelodella geryi Hoedeman, 1961.
o Pimelodella gracilis (Valenciennes, 1835).
o Pimelodella griffini Eigenmann, 1917.
o Pimelodella grisea (Regan, 1903).
o Pimelodella hartii (Steindachner, 1877).
o Pimelodella hartwelli Fowler, 1940.
o Pimelodella hasemani Eigenmann, 1917.
o Pimelodella howesi Fowler, 1940.
o Pimelodella itapicuruensis Eigenmann, 1917.
o Pimelodella kronei (Miranda Ribeiro, 1907).
o Pimelodella lateristriga (Lichtenstein, 1823).
o Pimelodella laticeps Eigenmann, 1917.
o Pimelodella laurenti Fowler, 1941.
o Pimelodella linami Schultz, 1944.
o Pimelodella macrocephala (Miles, 1943).
o Pimelodella macturki Eigenmann, 1912.
o Pimelodella martinezi Fernández-Yépez, 1970.
o Pimelodella meeki Eigenmann, 1910.
o Pimelodella megalops Eigenmann, 1912.
o Pimelodella megalura Miranda Ribeiro, 1918.
o Pimelodella metae Eigenmann, 1917.
o Pimelodella modesta (Günther, 1860).
o Pimelodella montana Allen, 1942.
o Pimelodella mucosa Eigenmann & Ward, 1907.
o Pimelodella nigrofasciata (Perugia, 1897).
o Pimelodella notomelas Eigenmann, 1917.
o Pimelodella odynea Schultz, 1944.
o Pimelodella ophthalmica (Cope, 1878).
o Pimelodella pallida Dahl, 1961.
o Pimelodella pappenheimi Ahl, 1925.
o Pimelodella parnahybae Fowler, 1941.
o Pimelodella parva Güntert, 1942.
o Pimelodella pectinifer Eigenmann & Eigenmann, 1888.
o Pimelodella peruana Eigenmann & Myers, 1942.
o Pimelodella peruensis Fowler, 1915.
o Pimelodella procera Mees, 1983.
o Pimelodella reyesi Dahl, 1964.
o Pimelodella roccae Eigenmann, 1917.
o Pimelodella rudolphi Miranda Ribeiro, 1918.
o Pimelodella serrata Eigenmann, 1917.
o Pimelodella spelaea Trajano, Reis & Bichuette, 2004.
o Pimelodella steindachneri Eigenmann, 1917.
o Pimelodella taeniophora (Regan, 1903).
o Pimelodella taenioptera Miranda Ribeiro, 1914.
o Pimelodella tapatapae Eigenmann, 1920.
o Pimelodella transitoria Miranda Ribeiro, 1907.
o Pimelodella vittata (Lütken, 1874).
o Pimelodella wesselii (Steindachner, 1877).
o Pimelodella witmeri Fowler, 1941.
o Pimelodella yuncensis .
* Genus Rhamdella Eigenmann & Eigenmann, 1888.
o Rhamdella aymarae Miquelarena & Menni, 1999.
o Rhamdella cainguae Bockmann and Miquelarena, 2008.
o Rhamdella eriarcha (Eigenmann & Eigenmann, 1888).
o Rhamdella exsudans (Jenyns, 1842).
o Rhamdella gilli (Starks, 1906).
o Rhamdella ignobilis Steindachner, 1907.
o Rhamdella jenynsii (Günther, 1864).
o Rhamdella leptosoma Fowler, 1914.
o Rhamdella longipinnis Borodin, 1927.
o Rhamdella longiuscula Lucena & da Silva, 1991.
o Rhamdella montana Eigenmann, 1913.
o Rhamdella papariae Fowler, 1941.
o Rhamdella robinsoni Fowler, 1941.
o Rhamdella rusbyi Pearson, 1924.
o Rhamdella wolfi Fowler, 1941.
* Genus Rhamdia Bleeker, 1858.
o Rhamdia enfurnada Bichuette & Trajano, 2005.
o Rhamdia foina (Müller & Troschel, 1848).
o Rhamdia guasarensis Do Nascimiento, Provenzano & Lundberg, 2004.
o Rhamdia humilis (Günther, 1864).
o Rhamdia itacaiunas Silfvergrip, 1996.
o Rhamdia jequitinhonha Silfvergrip, 1996.
o Rhamdia laluchensis Weber, Allegrucci & Sbordoni, 2003.
o Rhamdia laticauda (Kner, 1857).
o Rhamdia laukidi Bleeker, 1858.
o Rhamdia macuspanensis Weber & Wilkens, 1998.
o Rhamdia muelleri (Günther, 1864).
o Rhamdia nicaraguensis (Günther, 1864).
o Rhamdia parryi Eigenmann & Eigenmann, 1888.
o Rhamdia poeyi Eigenmann & Eigenmann, 1888.
o Rhamdia quelen (Quoy & Gaimard, 1824).
o Rhamdia reddelli Miller, 1984.
o Rhamdia xetequepeque Silfvergrip, 1996.
* Genus Rhamdioglanis Ihering, 1907.
o Rhamdioglanis frenatus Ihering, 1907.
o Rhamdioglanis transfasciatus Miranda Ribeiro, 1908.
* Genus Rhamdiopsis Haseman, 1911.
o Rhamdiopsis microcephala (Reinhardt, 1874).
o Rhamdiopsis moreirai Haseman, 1911.
* Genus Taunayia Miranda Ribeiro, 1918.
o Taunayia bifasciata (Eigenmann & Norris, 1900).

References

1. ^ "Heptapteridae". Integrated Taxonomic Information System. http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=679381. Retrieved 07 April 2007.
2. ^ a b c d Nelson, Joseph, S. (2006). Fishes of the World. John Wiley & Sons, Inc.. ISBN 0471250317.
3. ^ Froese, Rainer, and Daniel Pauly, eds. (2007). "Heptapteridae" in FishBase. Apr 2007 version.
4. ^ Sullivan, JP; Lundberg JG; Hardman M (2006). "A phylogenetic analysis of the major groups of catfishes (Teleostei: Siluriformes) using rag1 and rag2 nuclear gene sequences". Mol Phylogenet Evol. 41 (3): 636–62. doi:10.1016/j.ympev.2006.05.044. PMID 16876440.
5. ^ Bichuette, Maria Elina; Trajano, Eleonora (2005). "A new cave species of Rhamdia (Siluriformes: Heptapteridae) from Serra do Ramalho, northeastern Brazil, with notes on ecology and behavior". Neotropical Ichthyology 3 (4): 587–595.
6. ^ Ferraris, Carl J., Jr. (2007). "Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes), and catalogue of siluriform primary types" (PDF). Zootaxa 1418: 180–202. http://mapress.com/zootaxa/2007f/zt01418p300.pdf.
7. ^ Almiron et al. (2007). "Imparfinis mishky (Siluriformes, Heptapteridae) a new species from the rios Parana and Uruguay basins in Argentina.". Revue Suisse de Zoologie 114 (4): 817–824.
8. ^ Ribeiro, F.R.V., Pedroza, W.S., & Py-Daniel, L.H.R. (2011). "A new species of Nemuroglanis (Siluriformes: Heptapteridae) from the rio Gauriba, rio Madeira basin, Brazil." Zootaxa 2799: 41-48.