Ptychozoon

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Subordo: Gekkota
Infraordo: Gekkomorpha
Superfamilia: Gekkonoidea

Familia: Gekkonidae
Subfamilia: Gekkoninae
Genus: Ptychozoon
Species (13): P. bannaense – P. borneoensis – P. cicakterbang – P. cliveevatti – P. engannoensis – P. horsfieldii – P. intermedium – P. johorensis – P. kabkaebin – P. kaengkrachanense – P. kuhli – P. lionotum – P. malayensis – P. nicobarensis – P. popaense – P. rhacophorus – P. sulawesiensis – P. sumatraensis – P. tokehos – P. trinotaterra – P. wallaceaensis
Name

Ptychozoon Kuhl, 1824: 82

Type species: Lacerta homalocephala Creveldt, 1809, by original designation.
Synonyms

Ptychozoon Kuhl & van Hasselt, 1822: 475 [nomen nudum]

References
Primary references

Kuhl, H. & van Hasselt, J.C. 1822. Aus einem Schreiben von Dr. Kuhl und Dr. van Hasselt auf Java an Professor Th. van Swideren zu Gröningen. Isis von Oken 6: 472–476. BHL
Kuhl, H. 1824. Sur les reptiles de Java. Extraitd’une lettre adressée de Java en Hollande, par M.Kuhl, datée de Pjihorjavor au pied du Pangerango, le 18 juillet 1821. Bulletin des sciences naturelles et de géologie. Deuxième section du Bulletin universel des sciences et de l'industrie 2: 79–83. BHL

Additional references

Hoser, R.T. 2018b. A revised taxonomy of the gecko genus Ptychozoon Kuhl and Van Hasselt, 1822, including the formal erection of two new genera to accommodate the most divergent taxa and description of ten new species. Australasian Journal of Herpetology 38: 19-31. Full article (PDF). Reference page.
Klaver, C. 2007. On the status and date of publication of the generic name Ptychozoon (Reptilia, Gekkonidae). Bibliotheca Herpetologica 7(1): 7–11.
Brown, R.M. et al. 2012. Phylogeny and cryptic diversification in Southeast Asian flying geckos. Molecular phylogenetics and evolution 65 (2): 351–361. DOI: 10.1016/j.ympev.2012.06.009
Sumontha, M., Pauwels, O.S.G., Kunya, K., Limlikhitaksorn, C., Ruksue, S., Taokratok, A., Ansermet, M. & Chanhome, L. 2012. A new species of parachute gecko (Squamata: Gekkonidae: genus Ptychozoon) from Kaeng Krachan National Park, western Thailand. Zootaxa 3513: 68–78. Preview. Paywall. Reference page.
Wang, Y-Y., Wang, J. & Liu, Z-Y. 2016. Description of a new species of the genus Ptychozoon (Squamata: Gekkonidae), representing a new national record of this genus from southern Yunnan Province, China. Zootaxa 4084(3): 406–420. DOI: 10.11646/zootaxa.4084.3.6.Reference page.
Grismer, L.L., Wood, Jr., P.L., Thura, M.K., Grismer, M.S., Brown, R.M. & Stuart, B.L. 2018. Geographically structured genetic variation in Ptychozoon lionotum (Squamata: Gekkonidae) and a new species from an isolated volcano in Myanmar. Zootaxa 4514(2): 202–214. DOI: 10.11646/zootaxa.4514.2.4 Reference page.
Grismer, L.L., Wood. Jr., P.L., Grismer, J.L., Quah, E.S.H., Thy, N., Phimmachak, S., Sivongxay, N., Seateun, S., Stuart, B.L., Siler, C.D., Mulcahy, D.G., Anamza, T. & Brown, R.M. 2019. Geographic structure of genetic variation in the Parachute Gecko Ptychozoon lionotum Annandale, 1905 across Indochina and Sundaland with descriptions of three new species. Zootaxa 4638(2): 151–198. DOI: 10.11646/zootaxa.4638.2.1. Reference page.

Vernacular names
English: Flying Geckos
ไทย: ตุ๊กแกบิน
中文: 飞行壁虎属

Ptychozoon was a genus of arboreal geckos, endemic to Southeast Asia, known commonly as flying geckos, gliding geckos, or parachute geckos.[1] They all are now placed in the genus Gekko in the family Gekkonidae. The biogeographic history of the genus Ptychozoon was deeply nested within that of the genus Gekko, the center of diversity of which is within Southeast Asia. Since dispersing into Southeast Asian rainforests, Pytochozoon, like other forest-dwelling vertebrates, adapted to facilitate gliding.[2] All species in the genus Ptychozoon are characterized by cryptic coloration and elaborate webs surrounding the neck, limbs, trunk, and tail.[1] These membranes help to conceal the gecko against trees. When the gecko leaps into the air, the flaps are used to generate lift and allow the gecko to control its fall. It can glide up to 200 feet (60 meters). Also it does a swoop at the end of its glide to land softly. A similar adaptation is found in geckos of the genus Cosymbotus. There were thirteen described species in the genus Ptychozoon.

Species

The following species were recognized as being valid.[3]

Ptychozoon bannaense Y. Wang, J. Wang & Liu, 2016 – Banna parachute gecko
Ptychozoon cicakterbang Grismer, Wood, Grismer, Quah, Thy, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza, & Brown, 2019 – Malaysia parachute gecko
Ptychozoon horsfieldii (Gray, 1827) – Horsfield's flying gecko, Horsfield's parachute gecko
Ptychozoon intermedium Taylor, 1915 – intermediate flying gecko, Philippine parachute gecko
Ptychozoon kabkaebin Grismer, Wood, Grismer, Quah, Thy, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza, & Brown, 2019 – Lao parachute gecko
Ptychozoon kaengkrachanense Sumontha, Pauwels, Kunya, Limlikhitaksorn, Ruksue, Taokratok, Ansermet & Chanhome, 2012 – Kaeng Krachan parachute gecko
Ptychozoon kuhli Stejneger, 1902 – Kuhl's flying gecko, Kuhl's parachute gecko
Ptychozoon lionotum Annandale, 1905 – smooth-backed gliding gecko, smooth-backed flying gecko, Burmese flying gecko, Burmese parachute gecko
Ptychozoon nicobarensis Das & Vijayakumar, 2009 – Nicobar gliding gecko
Ptychozoon popaense Grismer, Wood, Thura, M. Grismer, Brown, & Stuart, 2018 – Mt. Popa parachute gecko
Ptychozoon rhacophorus (Boulenger, 1899) – Sabah flying gecko
Ptychozoon tokehos Grismer, Wood, Grismer, Quah, Thy, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza, & Brown, 2019 – Cambodian parachute gecko
Ptychozoon trinotaterra R.M. Brown, 1999

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Ptychozoon.
References

Malkmus R, Manthey U (2002). Amphibians & Reptiles of Mount Kinabalu (North Borneo). Germany: A.R.G. 424 pp. ISBN 978-3904144834. (Ptychozoon, pp. 32, 267-268).
Heinicke MP, Greenbaum E, Jackman TR, Bauer AM (December 2012). "Evolution of gliding in Southeast Asian geckos and other vertebrates is temporally congruent with dipterocarp forest development". Biology Letters. 8 (6): 994–7. doi:10.1098/rsbl.2012.0648. PMC 3497132. PMID 22977067.

"Ptychozoon ". The Reptile Database. www.reptile-database.org.

Further reading
Temminck CJ, van Swinderen T, de Haan W (1822). Kuhl H, van Hasselt JC (eds.). "Uittreksels uit brieven ... aan de Heeren" [Ptychozoon, new genus]. Alg. Konst- en Letter-Bode (in Dutch). 1: 99–104.
Smith MA (1935). "Genus Ptychozoon". The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. Vol. II.—Sauria. London: Secretary of State for India in Council.: Taylor and Francis. p. 117.
Heinicke MP, Greenbaum E, Jackman TR, Bauer AM (December 2012). "Evolution of gliding in Southeast Asian geckos and other vertebrates is temporally congruent with dipterocarp forest development". Biology Letters. 8 (6): 994–7. doi:10.1098/rsbl.2012.0648. PMC 3497132. PMID 22977067.