Hemiphyllodactylus

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Subordo: Gekkota
Infraordo: Gekkomorpha
Superfamilia: Gekkonoidea

Familia: Gekkonidae
Subfamilia: Gekkoninae
Genus: Hemiphyllodactylus
Species (52): H. arakuensis – H. aurantiacus – H. banaensis – H. bintik – H. bonkowskii – H. changningensis – H. chiangmaiensis – H. cicak – H. dalatensis – H. dupanglingensis – H. dushanensis – H. engganoensis – H. flaviventris – H. ganoklonis – H. goaensis – H. harterti – H. hongkongensis – H. huishuiensis – H. indosobrinus – H. insularis – H. jinpingensis – H. jnana – H. khlonglanensis – H. kiziriani – H. kolliensis – H. kyaiktiyoensis – H. larutensis – H. linnwayensis – H. longlingensis – H. margarethae – H. minimus – H. montawaensis – H. nahangensis – H. ngocsonensis – H. ngwelwini – H. nilgiriensis – H. pardalis – H. peninsularis – H. pinlaungensis – H. serpispecus – H. tehtarik – H. titiwangsaensis – H. tonywhitteni – H. typus – H. uga – H. yunnanensis – H. ywanganensis – H. zalonicus – H. zayuensis – H. zhutangxiangensis – H. zugi – H. zwegabinensis
Name

Hemiphyllodactylus Bleeker, 1860: 327

Type species: Hemiphyllodactylus typus Bleeker, 1860, by monotypy.

References
Primary references

Bleeker, P. 1860. Conspectus systematis cyprinorum. Natuurkundig Tijdschrift voor Nederlandsch-Indië 20: 421–441. BHL Reference page.

Additional references

Zug, G.R. 2010. Speciation and Dispersal in a Low Diversity Taxon: The Slender Geckos Hemiphyllodactylus (Reptilia, Gekkonidae). Smithsonian Contributions Museum 631: 1–70.
Grismer, L.L., Wood, Jr., P.L., Anuar Mohd Sah, S., Muin, M.A., Quah, E.S.H., McGuire, J.A., Brown, R.M., Ngo, V.T. & Thai, P.H. 2013. Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia. Zoological Journal of the Linnean Society 169(4): 849–880. DOI: 10.1111/zoj.12064 Open access Reference page.
Nguyen, T.Q., Lehmann, T., Le, M.D., Duong, H.T., Bonkowski, M. & Ziegler, T. 2013. A new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from northern Vietnam. Zootaxa 3736(1): 89–98. DOI: 10.11646/zootaxa.3736.1.5 Paywall Reference page.
Grismer, L.L., Riyanto, A., Iskandar, D.T. & McGuire, J.A. 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Pulau Enggano, southwestern Sumatra, Indonesia. Zootaxa 3821(4): 485–495. DOI: 10.11646/zootaxa.3821.4.7 Paywall PDF ResearchGate Open accessReference page.
Nguyen, T.Q., Botov, A., Le, M.D., Nophaseud, L., Zug, G., Bonkowski, M. & Ziegler, T. 2014. A new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from northern Laos. Zootaxa 3827(1): 45–56. DOI: 10.11646/zootaxa.3827.1.4 Reference page.
Tri, N.V., Grismer, L.L., Thai, P.H. & Wood Jr., P.L. 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Ba Na–Nui Chua Nature Reserve, central Vietnam. Zootaxa 3760(4): 539–552. DOI: 10.11646/zootaxa.3760.4.3 Paywall Reference page.
Grismer, L.L., Wood, P.L. Jr, Anuar Mohd Sah, S., Quah, E.S.H., Muin, M.A., Onn, C.K., Sumarli, A.X. & Loredo, A.I. 2015. Repeated evolution of sympatric, palaeoendemic species in closely related, co-distributed lineages of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) across a sky-island archipelago in Peninsular Malaysia. Zoological Journal of the Linnean Society 174(4): 859–876. DOI: 10.1111/zoj.12254 Reference page.
Guo, W-B., Zhou, K-Y., Yan, J. & Li, P. 2015. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from western Yunnan, China. Zootaxa 3974(3): 377–390. DOI: 10.11646/zootaxa.3974.3.5. Paywall. Preview (PDF). Reference page
Cobos, A.J., Grismer, L.L., Wood Jr., P.L., Quah, E.S.H., Anuar Mohd Sah, S. & Muin, M.A. 2016. Phylogenetic relationships of geckos of the Hemiphyllodactylus harterti group, a new species from Penang Island, Peninsular Malaysia, and a likely case of true cryptic speciation. Zootaxa 4107(3): 367–380. DOI: 10.11646/zootaxa.4107.3.5 Paywall Reference page.
Yan, J., Lin, Y., Guo, W-B., Li, P. & Zhou, K-Y. 2016. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Guizhou, China. Zootaxa 4117(4): 543–554. DOI: 10.11646/zootaxa.4117.4.6 Paywall Reference page.
Grismer, L.L., Wood Jr., P.L., Thura, M.K., Zin, T., Quah, E.S.H., Murdoch, M.L., Grismer, M.S., Lin, A., Kyaw, H. & Lwin, N. 2017. Phylogenetic taxonomy of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) with descriptions of three new species from Myanmar. Journal of Natural History 52(13–16): 881–915. DOI: 10.1080/00222933.2017.1367045 Paywall Reference page.
Grismer, L.L., Wood, Jr., P.L., Zug, G.R., Thura, M.K., Grismer, M.S., Murdoch, M.L., Quah, E.S.H. & Lin, A. 2018. Two more new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from the Shan Hills of eastern Myanmar (Burma). Zootaxa 4483(2): 295–316. DOI: 10.11646/zootaxa.4483.2.4. Reference page.
Sukprasert, A., Sutthiwises, S., Lauhachinda, V. & Taksintum, W. 2018. Two new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from Thailand. Zootaxa 4369(3): 363–376. DOI: 10.11646/zootaxa.4369.3.4 Reference page.
Sung, Y-H., Lee, W-H., Ng, H-N., Zhang, Y-J. & Yang, J-H. 2018. A new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hong Kong. Zootaxa 4392(2): 361–373. DOI: 10.11646/zootaxa.4392.2.8 Reference page.
Agarwal, I., Khandekar, A., Giri, V.B., Ramakrishnan, U. & Karanth, K.P. 2019. The hills are alive with geckos! A radiation of a dozen species on sky islands across peninsular India (Squamata: Gekkonidae, Hemiphyllodactylus) with the description of three new species. Organisms Diversity & Evolution 19: 341–361. DOI: 10.1007/s13127-019-00392-5 Open access Reference page.
Eliades, S.J., Phimmachak, S., Sivongxay, N., Siler, C.D. & Stuart, B.L. 2019. Two new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from Laos. Zootaxa 4577(1): 131–147. DOI: 10.11646/zootaxa.4577.1.8 Paywall Reference page.
Agarwal, I., Bauer, A.M., Pal, S., Srikanthan, A.N. & Khandekar, A. 2020. Two more new Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Tamil Nadu, India. Zootaxa 4729(2): 249–265. DOI: 10.11646/zootaxa.4729.2.6 Paywall Reference page.
Do, Q.H., Pham, C.T., Phan, T.Q., Le, M.D., Ziegler, T. & Nguyen, T.Q. 2020. A new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Tuyen Quang Province, Vietnam. Zootaxa 4821(3): 511–532. DOI: 10.11646/zootaxa.4821.3.3 Paywall Reference page.
Grismer, L.L., Yushchenko, P.V., Pawangkhanant, P., Naiduangchan, M., Nazarov, N.A., Orlova, V.F., Suwannapoom, C. & Poyarkov, N.A. 2020. A new species of Hemiphyllodactylus Bleeker (Squamata; Gekkonidae) from Peninsular Thailand that converges in morphology and color pattern on Pseudogekko smaragdinus (Taylor) from the Philippines. Zootaxa 4816(2): 171–190. DOI: 10.11646/zootaxa.4816.2.2 Paywall Reference page.
Mohapatra, P.P., Khandekar, A., Dutta, S.K., Mahapatra, C. & Agarwal, I. 2020. A novel, diminutive Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from a sacred grove in Odisha, eastern India. Zootaxa 4852(4): 485–499. DOI: 10.11646/zootaxa.4852.4.6 Paywall Reference page.
Nguyen, T.Q., Do, Q.H., Ngo, H.T., Pham, A.V., Pham, C.T., Le, M.D. & Ziegler, T. 2020. Two new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hoa Binh Province, Vietnam. Zootaxa 4801(3): 513–536. DOI: 10.11646/zootaxa.4801.3.5 Paywall Reference page.
Zhang, B., Qian, T.-Y., Jiang, X.-J., Cai, B., Deng, X.-J. & Yang, D. 2020. A New Species of Hemiphyllodactylus Bleeker, 1860 (Reptilia: Squamata) from Hunan Province, China. Asian Herpetological Research 11(3): 183–193. DOI: 10.16373/j.cnki.ahr.190066 Paywall Reference page.
Agung, A.P., Grismer, L.L., Grismer, J.L., Quah, E.S.H., Chornelia, A., Lu, J.-M. & Hughes, A.C. 2021. A new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from Yunnan, China and its phylogenetic relationship to other congeners. Zootaxa 4980(1): 1–27. DOI: 10.11646/zootaxa.4980.1.1 Paywall Reference page.
Do, Q.H., Nguyen, K.V., Le, M.D., Ziegler, T. & Nguyen, T.Q. 2021. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Da Lat Plateau, Vietnam. Zootaxa 5023(1): 93–106. DOI: 10.11646/zootaxa.5023.1.5 Paywall Reference page.
Grismer, L.L., Chit, M.T., Pawangkhanant, P., Nazarov, N.A., Zaw, T. & Poyarkov, N.A. 2021. The phylogeny of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) with a description of a new species from the Mangin Range, Sagaing Region, northern Myanmar. Journal of Natural History 54(29-30): 1913–1931. DOI: 10.1080/00222933.2020.1833095 Paywall Reference page.
Khandekar, A., Parmar, D.S., Sawant, N. & Agarwal, I. 2021. A new species of the genus Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Goa, India. Zootaxa 5027(2): 254–268. DOI: 10.11646/zootaxa.5027.2.6 Paywall Reference page.

Links

Uetz, P. & Hallermann, J. 2022. Hemiphyllodactylus . The Reptile Database. Accessed on 5 April 2022.

Vernacular names
English: Slender Geckos

Hemiphyllodactylus is a genus of geckos ranging from India and China southward to Southeast Asia and Oceania. Species of Hemiphyllodactylus are commonly known as half leaf-fingered geckos. Many species are known as dwarf geckos or slender geckos.
Species

Hemiphyllodactylus arakuensis Agarwal, Khandekar, Giri, Ramakrishnan, & Karanth, 2019
Hemiphyllodactylus aurantiacus (Beddome, 1870) – Southern Ghats slender gecko
Hemiphyllodactylus banaensis Ngo Van Tri et al., 2014 [1] – Ba Na dwarf gecko
Hemiphyllodactylus bintik Grismer et al., 2015 [2] – spotted dwarf gecko
Hemiphyllodactylus bonkowskii Nguyen, Do, Ngo, Pham, Pham, Le, & Ziegler, 2020 [2] – Bonkowski’s slender gecko
Hemiphyllodactylus changningensis Guo et al., 2015 [2] – Changning slender gecko
Hemiphyllodactylus chiangmaiensis Grismer, Wood & Cota, 2014 [3] – Chiang Mai dwarf gecko
Hemiphyllodactylus cicak Grismer et al., 2016 [2] – Penang Island slender gecko
Hemiphyllodactylus dalatensis Do, Nguyen, Le, Pham, Ziegler, & Nguyen, 2021 [2]
Hemiphyllodactylus dupanglingensis Zhang, Qian, & Yang, 2020 [2] – Dupangling slender gecko
Hemiphyllodactylus dushanensis (Zhou & Liu, 1981) [2] – Dushan slender gecko, Dushan gypsy gecko, or Dushan dwarf gecko
Hemiphyllodactylus engganoensis Grismer et al., 2014 [2] – Pulau Enggano dwarf gecko
Hemiphyllodactylus flaviventris Sukprasert, Sutthiwises, Lauhachinda, & Taksintum, 2018 – yellow-bellied dwarf gecko
Hemiphyllodactylus ganoklonis Zug, 2010 [2] – Palauan slender gecko
Hemiphyllodactylus goaensis Khandekar, Parmar, N Sawant, & Agarwal, 2021 [2] – Goan slender gecko
Hemiphyllodactylus harterti (F. Werner, 1900) [2] – Bintang slender gecko, Hartert's slender gecko
Hemiphyllodactylus hongkongensis Sung, Lee, Ng, Zhang, & Yang, 2018 [2] – Hong Kong slender gecko
Hemiphyllodactylus huishuiensis Yan, Lin, Guo, Li, & Zhou, 2016 [2]
Hemiphyllodactylus indosobrinus Eliades, Phimmachak, Sivongxay, Siler, & Stuart, 2019 [2]
Hemiphyllodactylus insularis Taylor, 1918 – Philippine slender gecko
Hemiphyllodactylus jinpingensis (Zhou & Liu, 1981) [2] – Jinping slender gecko, Jinping gypsy gecko, or Jinping dwarf gecko
Hemiphyllodactylus jnana Agarwal, Khandekar, Giri, Ramakrishnan, & Karanth, 2019
Hemiphyllodactylus khlonglanensis Sukprasert, Sutthiwises, Lauhachinda, & Taksintum, 2018 [2] – Khlong Lan dwarf gecko
Hemiphyllodactylus kiziriani Nguyen et al., 2014 [2] – Kizirian's slender gecko
Hemiphyllodactylus kolliensis Agarwal, Khandekar, Giri, Ramakrishnan, & Karanth, 2019
Hemiphyllodactylus kyaiktiyoensis Grismer, Wood, Quah, Thura, Oaks, & Lin, 2020 – Kyaiktiyo Mountain slender gecko
Hemiphyllodactylus larutensis (Boulenger, 1900) – Larut dwarf gecko
Hemiphyllodactylus linnwayensis Grismer et al., 2017 – Lynn-Way dwarf gecko
Hemiphyllodactylus longlingensis (Zhou & Liu, 1981) [2] – Longling slender gecko, Longling gypsy gecko, or Longling dwarf gecko
Hemiphyllodactylus margarethae Brongersma, 1931 [2] – Sumatran slender gecko
Hemiphyllodactylus minimus Mohapatra, Khandekar, Dutta, Mahapatra & Agarwal, 2020[4] – Ganjam slender gecko
Hemiphyllodactylus montawaensis Grismer et al., 2017 – Montawa dwarf gecko
Hemiphyllodactylus nahangensis Do, Pham, Phan, Le, Ziegler, & Nguyen, 2020 [2] – Nahang slender gecko
Hemiphyllodactylus ngwelwini Grismer, Wood, Quah, Thura, Oaks, & Lin, 2020 – Ngwe Lwin’s slender gecko
Hemiphyllodactylus ngocsonensis Nguyen, Do, Ngo, Pham, Pham, Le, & Ziegler, 2020 [2] – Ngocson slender gecko
Hemiphyllodactylus nilgiriensis Agarwal, Bauer, Pal, Srikanthan, & Khandekar, 2020 – Nilgiris slender gecko
Hemiphyllodactylus pardalis Grismer, Yushchenko, Pawangkhanant, Naiduangchan, Nazarov, Orlova, Suwannapoom, & Poyarkov, 2020 [2] – spotted slender gecko
Hemiphyllodactylus peninsularis Agarwal, Bauer, Pal, Srikanthan, & Khandekar, 2020 – KMTR slender gecko
Hemiphyllodactylus pinlaungensis Grismer, Wood, Quah, Thura, Oaks, & Lin, 2020 – Pinlaung slender gecko
Hemiphyllodactylus serpispecus Eliades, Phimmachak, Sivongxay, Siler, & Stuart, 2019 [2]
Hemiphyllodactylus tehtarik Grismer et al., 2013 [5] – Tebu Mountain slender-toed gecko
Hemiphyllodactylus titiwangsaensis Zug, 2010 [2] – Titiwangsa slender gecko
Hemiphyllodactylus tonywhitteni Grismer et al., 2017 – Phapant dwarf gecko
Hemiphyllodactylus typus Bleeker, 1860 – Indopacific tree gecko, Indopacific slender gecko, or common dwarf gecko
Hemiphyllodactylus typus chapaensis Bourret, 1937
Hemiphyllodactylus typus pallidus Auffenberg, 1980
Hemiphyllodactylus typus typus Bleeker, 1860
Hemiphyllodactylus uga Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Li, Kyaw, & Lwin, 2018 – Uga's slender gecko
Hemiphyllodactylus yunnanensis (Boulenger, 1903) – Asian slender gecko, Yunnan gypsy gecko, or Yunnan dwarf gecko
Hemiphyllodactylus ywanganensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Li, Kyaw, & Lwin, 2018 – Ywangan slender gecko
Hemiphyllodactylus zalonicus Grismer, Chit, Pawangkhanant, Nazarov, Zaw, & Poyarkov, 2021 [2] – Mt. Zalon slender gecko
Hemiphyllodactylus zayuensis Jiang, Wang, & Che, 2020 [2]
Hemiphyllodactylus zugi Nguyen et al., 2013 [6] – Zug's slender gecko
Hemiphyllodactylus zhutangxiangensis Agung, Grismer, Grismer, Quah, Chornelia, Lu, & Hughes, 2021
Hemiphyllodactylus zwegabinensis Grismer, Wood, Quah, Thura, Oaks, & Lin, 2020 – Zwegabin Mountain slender gecko

Nota bene: In the above list, a binomial authority in parentheses indicates that the species was originally described in a genus other than Hemiphyllodactylus.
Notes

Ngo Van Tri et al. (2014). "A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Ba Na–Nui Chua Nature Reserve, Central Vietnam". Zootaxa 3760 (4): 539-552.
"Hemiphyllodactylus ". The Reptile Database. www.reptile-database.org.
Grismer et al. (2014). "A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from northwestern Thailand". Zootaxa 3760 (1): 67-78.
Mohapatra, Pratyush P.; Khandekar, Akshay; Dutta, Sushil K.; Mahapatra, Cuckoo; Agarwal, Ishan (2020-09-17). "A novel, diminutive Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from a sacred grove in Odisha, eastern India". Zootaxa. 4852 (4): 485–499. doi:10.11646/zootaxa.4852.4.6. ISSN 1175-5334. PMID 33056409. S2CID 222830832.
Grismer et al. (2013). "Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia". Zool. J. Linnean Soc. 169 (4): 849-880.

Nguyen et al. (2013). "A new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from northern Vietnam". Zootaxa 3736 (1): 89-98.