Hemidactylus frenatus

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Subordo: Gekkota
Infraordo: Gekkomorpha
Superfamilia: Gekkonoidea

Familia: Gekkonidae
Subfamilia: Gekkoninae
Genus: Hemidactylus
Species: Hemidactylus frenatus
Name

Hemidactylus frenatus Schlegel in Duméril & Bibron, 1836
Synonyms

Hemidactylus fragilis Calabresi, 1915
Hemidactylus hexaspis Cope, 1869
Hemidactylus javanicus Fitzinger, 1826 [nomen nudum]
Hemidactylus longiceps Cope, 1869
Hemidactylus nigriventris Lidth de Jeude, 1905
Hemidactylus okinavensis Okada, 1936
Hemidactylus punctatus Jerdon, 1853
Hemidactylus tristis Sauvage, 1879
Hemidactylus vandermeer-mohri Brongersma, 1928
Hemidactylus vittatus Gray, 1845
Pnoepus bojeri (Fitzinger, 1843)
Pnoepus frenatus (Schlegel in Duméril & Bibron, 1836)
Pnoepus papuensis (Macleay, 1877)
Pnoepus punctatus (Jerdon, 1853)

References

IUCN: Hemidactylus frenatus Schlegel in Duméril & Bibron, 1836 (Least Concern)

Vernacular names
অসমীয়া: জেঠী
Bikol Central: tagaw
বাংলা: টিকটিকি
Cebuano: tiki
English: Common House Gecko, Spiny-tailed House Gecko
magyar: Ázsiai házigekkó
Ilokano: alutiit
日本語: ホオグロヤモリ
ไทย: จิ้งจกบ้านหางหนาม
Tagalog: butiki
Tiếng Việt: Thạch sùng

The common house gecko (Hemidactylus frenatus) is a gecko native to South and Southeast Asia. It is also known as the Asian house gecko, Pacific house gecko, wall gecko, house lizard, tayoto, chipkali[3] or moon lizard.

Most geckos are nocturnal, hiding during the day and foraging for insects at night. They can be seen climbing walls of houses and other buildings in search of insects attracted to porch lights, and are immediately recognisable by their characteristic chirping.

They grow to a length of between 7.5–15 cm (3–6 in), and live for about 5 years. These small geckos are non-venomous and not harmful to humans. Most medium-sized to large geckos are docile, but may bite if distressed, which can pierce skin. The common house gecko is tropical, and thrives in warm, humid areas where it can crawl around on rotting wood in search of the insects it eats, as well as within urban landscapes in warm climates. The animal is very adaptable and may prey on insects and spiders, displacing other gecko species which are less robust or behaviourally aggressive. In parts of Australia and Papua New Guinea they are often confused with a similar native lizard, the dubious dtella.

Etymology

Like many geckos, this species can lose its tail when alarmed. Its call or chirp rather resembles the sound "gecko, gecko". However, this is an interpretation, and the sound may also be described as "tchak tchak tchak" (often sounded six to nine times in sequence). In Asia/Southeast Asia, notably Indonesia, Thailand, Singapore, and Malaysia, geckos have local names onomatopoetically derived from the sounds they make: Hemidactylus frenatus is called "chee chak" or "chi chak" (pr- chee chuck), said quickly, also commonly spelled as "cicak" in Malay dictionaries. In the Philippines, they are called "butiki" in Tagalog, "tiki" in Visayan, "alutiit" in Ilocano, and in Thailand, "jing-jok" (Thai: จิ้งจก[4]). In Myanmar, they are called "အိမ်မြှောင် - ain-mjong" ( "အိမ် - ain" means "house" and "မြှောင် - mjong" means "stick to"). In some parts of India and in Pakistan, they are called "chhipkali" (Urdu:چھپکلی, Hindi: छिपकली), from chhipkana, to stick. In Nepal, they are called "vhitti" (Nepali: भित्ती) or "mausuli" (Nepali: माउसुली). In other parts of India, they are called "kirli" (Punjabi: ਕਿੜਲੀ), "jhiti piti" (Oriya: ଝିଟିପିଟି), "zethi" (Assamese: জেঠী), "thikthikiaa" (Maithili: ठिकठिकिया), "paal" (Marathi: पाल), "gawli" or "palli" (Malayalam: ഗവ്ളി (gawli), പല്ലി (palli), Tamil: பல்லி (palli)), Telugu: బల్లి (balli), Kannada: ಹಲ್ಲಿ (halli), "ali" (Sylheti: ꠀꠟꠤ), "garoli" (Gujarati: ગરોળી). In West Bengal and Bangladesh, they are called "tiktiki" (Bengali: টিকটিকি) as the sound is perceived as "tik tik tik". In Sri Lanka, they are called "huna" in singular form (Sinhalese: හුනා). In Central America, they are sometimes called "limpia casas" (Spanish: "house cleaners") because they reduce the amount of insects and other arthropods in homes and are also called 'qui-qui' because of the sound they make.
Habitat and diet
Head profile
Ventral view of mating house geckos, with hemipenis inserted in the cloaca, and adhesive lamellae under the feet
House gecko catching a cockroach

The common house gecko is by no means a misnomer, displaying a clear preference for urban environments. The synanthropic gecko displays a tendency to hunt for insects in close proximity to urban lights.[5] They have been found in bushland, but the current evidence seems to suggest they have a preference for urban environments, with their distribution being mostly defined by areas within or in close proximity to city bounds.[6][failed verification]
A baby house gecko

The common house gecko appears to prefer areas in the light which are proximal to cracks, or places to escape. Geckos without an immediate opportunity to escape potential danger display behavioural modifications to compensate for this fact, emerging later in the night and retreating earlier in the morning.[7] Without access to the urban landscape, they appear to prefer habitat which is composed of comparatively dense forest or eucalypt woodland which is proximal to closed forest.[8]

The selection of primarily urban habitats makes available the preferred foods of the common house gecko. The bulk of the diet of the gecko is made up of invertebrates, primarily hunted around urban structures.[5] Primary invertebrate food sources include cockroaches, termites, some bees and wasps, butterflies, moths, flies, spiders, and several beetle groupings.[5] There is limited evidence that cannibalism can occur in laboratory conditions, but this is yet to be observed in the wild.[9]
Distribution

The common house gecko is prolific through the tropics and subtropics. It is able to exist in an ecologically analogous place with other Hemidactylus species.[10] Despite being native throughout Southeast Asia, recent introductions, both deliberate and accidental, have seen them recorded in the Deep South of the United States, large parts of tropical and sub-tropical Australia, and many other countries in South and Central America, Caribbean Dominican Republic, Africa, South Asia and the Middle East (Bahrain, Jordan, Qatar, Kuwait, Saudi Arabia, Oman and the United Arab Emirates). Most recently, this species has also invaded the Caribbean Lesser Antilles, and is now present on Dominica and Saint Lucia.[11][12] Their capacity to withstand a wide range of latitudes is also partially facilitated by their capacity to enter a state of brumation during colder months. The prospect of increased climate change interacts synergistically with increased urbanisation, greatly increasing the prospective distribution of the common house gecko.[10] Due to concerns over its potential capacity as an invasive species, there are efforts to limit their introduction and presence in locations where they could be a risk to native gecko species.

In Mexico, H. frenatus was first collected in Acapulco, Guerrero, in March of 1895 and found to be well established there and in the surrounding regions by the early 1940s. It was likely introduced through shipping and cargo. H. frenatus now occurs throughout the lowlands of Mexico on both the Atlantic and Pacific versants including the Yucatan Peninsula, and Baja California, with records from 21 of the 32 Mexican states. Most records of H. frenatus in Mexico are from buildings such as homes, hotels, and other structure in cities and towns, with only a few reports of the species in natural habitat, and its impact, if any, on native fauna there is unknown.[13]
As an invasive species

There is evidence to suggest that the presence of Hemidactylus frenatus has negatively impacted native gecko populations throughout tropical Asia, Central America and the Pacific.[10]

Some species which have been displaced include:

Lepidodactylus lugubris[14][15][16][17]
Hemidactylus garnotti[14][15][16][17]
The genus Nactus on the Mascarene Islands (three species in this genus are now considered to be extinct)[18]

As an introduced species, they pose a threat through the potential introduction of new parasites and diseases, but have potential negative impacts which extend beyond this.[19] The primary cause for concern appears to exist around their exclusionary behaviour and out-competition of other gecko species.[9][15] Mechanistically, three explanations have been derived to justify the capacity of H. frenatus to outcompete other gecko species:

Possessing a smaller body size. They fail to displace native species larger than themselves, such as the robust velvet gecko.[20]
Male H. frenatus displaying higher levels of aggression than females of other gecko species (particularly parthenogenic species with asexual females).
Sexual females displaying an increased capacity to compete in comparison to asexual females.[15]

These differences provide H. frenatus a competitive edge in the limited urban areas they preferentially inhabit, particularly those with high degrees of habitat fragmentation.[14] To compound this, they also are capable of operating on higher densities, which leads to an increase in gecko sightings and biomass in an area, even after reducing native species' density.[14] The common house gecko also displays a higher tolerance to high light levels, which may allow for an increased risk-reward pay off in hunting endeavours. There is also some limited evidence for cannibalism, hunting on other small gecko species, particularly juveniles.[9] Most of this evidence is in laboratory conditions, with several studies failing to find evidence of cannibalism in the wild for this species.[14][21]

Some males are more territorial than others. Territorial males will display larger heads, with a more pronounced head shape. This increase in head size incurs the cost of a poorer performance in escape sprint time. This suggests selective pressure prioritises the biting force capacity of the male, over their capacity to escape quickly. On the contrary, increases in female head size are met with a proportionate increase in hind limb length and no decrease in speed. Though both sexes use escape sprinting as a survival strategy, males are more likely to need to stop and fight using biting, due to the reduced mobility caused by disproportionate head to hind leg size, which in turn is correlated with localised territorial behaviours.[22]

The success of the common house gecko can also be explained through other elements of competition, such as postural displays and movement patterns. An example of this is how the common house gecko can trigger an "avoidance response" in the mourning gecko, causing it to avoid a specific area where food may become available.[15] Though triggering avoidance in other species, they themselves can tolerate the presence of other gecko species well, regardless of whether those species are smaller or larger, faster or slower, or more physically aggressive or not.[10][14][18] This allows them greater access to feeding areas and territories, making them a highly successful invasive species.
Physiology

The common house gecko is ectothermic ("cold-blooded") and displays a variety of means of thermoregulating through behaviour. Its physiology has ramifications for its distribution and nature of interaction with native species, as well as reproductive success as an introduced species.

Metabolically, the demand of the common house gecko is not significantly variable from other lizard species of a similar size, with oxygen consumption appearing congruent with trends observed in other tropical, subtropical and temperate species of gecko. Thermal independence exists between 26–35 degrees, with some capacity to self regulate temperature. This means that where the environmental temperature is 26–35 degrees, the common house gecko can modify body temperature through behavioural adaptations. Breathing rates of geckos are temperature dependent above this maximal heat, but independent as it grows colder.[23] There are behavioural mechanisms of thermoregulation present, such as the selection of sunlight[24] and the substrates on which they sit.

The common house gecko can be best defined as quinodiurnal. This means they thermoregulate during the daytime and forage at night.[25] An active form of this thermoregulation includes the presence of the Gecko in lighter environments, proximal to cracks in the substrate. As such, there is a close relationship between activity levels and correlated air temperature.[7] Timing of the circadian rhythm of the common house gecko is further impacted by light levels. This rhythm tends to involve the highest population presence around midnight, with highest activity levels just after sunset,[26] with a gradual reduction until dawn. Daily cycle differences from place to place can generally be explained by environmental factors such as human interaction, and structural features.[7] A peak in hunting activity after dark places them in an ideal spot to take advantage of invertebrate congregation around artificial lighting in the urban environment.

Due to this level of dependence on the environment, drops in temperature may act as a leading indicator for reduced gecko sightings in the medium term. Acute weather events such as rain or wind will result in acute decreases in Gecko sightings within that environment. It is unsure what impact these phenomena may have on the long term on distribution and the capacity of the common house gecko to compete with other gecko species.

There is some weak evidence to suggest a trend towards higher temperature for females, which has an evolutionary advantage of increasing the speed of egg development. However, there is no statistically significant data to support this.[25]

Due to them being a species which is adapted for tropical or subtropical environments, there appear to be few physiological adaptations designed to prevent water loss. This may limit their capacity to thrive in arid or semi-arid environments.[23]
Reproductive biology

H. frenatus has a similar gonad structure to the remainder of the gecko family. It is possible to differentiate the sex of larger common house geckos, with individuals which are larger than 40 mm (1.6 in) typically displaying differentiated gonads. Differentiated gonads are most clearly seen with a swelling at the entrance to the cloaca caused by the copulatory organs in males. Females lay a maximum of two hard-shelled eggs at any single time, with each descending from a single oviduct. Up to four eggs can exist within the ovaries in differing stages of development. This shortens the potential turn around between egg-laying events in gravid females.[27] Females produce a single egg per ovary per cycle. This means they are considered monoautochronic ovulatory.[28]

Within the testes, mature sperm are found in the male geckos year-round and are able to be stored within the oviduct of the female. Sperm can be stored for a period of time as long as 36 weeks. This provides a significantly increased chance of colonisation of new habitats, requiring smaller populations to be transplanted for a chance of success. However, longer storage time of sperm within the female is associated with negative survival outcomes and hatching, possibly due to sperm age. Sperm is specifically stored between the uterine and infundibular components of the oviduct. The capacity to store sperm enables a degree of asynchrony between ovulation, copulation and laying of eggs.[28] The capacity to store sperm is useful in island colonisation events, providing females which may be isolated the capacity to reproduce even if they have been separated from a male for some time.[29] In laboratories, one mating event may produce as many as seven viable egg clutches. This eliminates the need for parthenogenesis and allows the young to include both male and female offspring, with one mating event leading to multiple clutches of eggs being laid. This reduced need for asexual reproduction increases the fitness of young through hybrid vigour and increased diversity.[28] As well as this, sexually reproducing geckos are reported to be more robust and have higher survival rates than those which reproduce asexually.[10]

There is a positive correlation between size and viability of eggs, with larger geckos having eggs which were more likely to survive. There is also a correlation between warmer year-round temperatures and consistent food supply with reproductive seasonality, with Geckos with constant food and temperatures being less likely to develop fat deposits on their stomach, and more likely to be constantly reproductive.[27]
Genetics

Two distinct karyotypes of the common house gecko appear to exist, one with 40 chromosomes and one with 46 chromosomes.[30][31] This could be explained through an intraspecific variation of karyotype, or the possibility of two distinct species being misidentified. Morphological analysis seems particularly congruent with the suggestion that they indeed are different species.[31][32] Taxonomic revision may be required as a greater understanding of phylogenetic trees and population structures is developed.
House geckos in captivity
House gecko hunting the pointer of a computer mouse

House geckos can be kept as pets in a vivarium with a clean substrate, and typically require a heat source and a place to hide in order to regulate their body temperature, and a system of humidifiers and plants to provide them with moisture. [33]

The species will cling to vertical or even inverted surfaces when at rest. In a terrarium they will mostly be at rest on the sides or on the top cover rather than placing themselves on plants, decorations or on the substrate, thus being rather conspicuous.
Cultural beliefs
Two wall sculptures of geckos on the wall of the Mandapam of the Siva temple inside Vellore Fort, Tamil Nadu, India (2012)

In Southeast Asia, geckos are believed to be carriers of good omen.[citation needed] In the Philippines, geckos making a ticking sound are believed to indicate an imminent arrival of a visitor or a letter.[34]

An elaborate system of predicting good and bad omens based on the sounds made by geckos, their movement and the rare instances when geckos fall from roofs has evolved over centuries in India.[35][36] In some parts of India, the sound made by geckos is considered a bad omen; while in parts of India, Assam, West Bengal, Bangladesh and Nepal, it is considered to be an endorsement of the truthfulness of a statement made just before, because the sound "tik tik tik" resembles the expression "thik thik thik" (Assamese: ঠিক ঠিক ঠিক), which in many Indian languages (e.g. Bengali and Assamese) means "correct correct correct", i.e., a three-fold confirmation. The cry of a gecko from an east wall as one is about to embark on a journey is considered auspicious, but a cry from any other wall is supposed to be inauspicious.[citation needed] A gecko falling on someone's right shoulder is considered good omen, but a bad omen if it falls on the left shoulder. In Punjab, it is believed that contact with the urine of a gecko will cause leprosy.[37] In some places in India, it is believed that watching a lizard on the eve of Dhanteras is a good omen or a sign of prosperity.[citation needed]

In Sri Lanka, It's believed that it's inauspicious if a gecko makes a sound while someone is going out of the house. And, there is a art of divination based on gecko's falling into one's body, different body parts indicates different predictions. This art of divination can be observed throughout the Indian subcontinent.[38]
Notes

Ota, H. & Whitaker, A.H. 2010. Hemidactylus frenatus. The IUCN Red List of Threatened Species 2010: e.T176130A7184890. https://dx.doi.org/10.2305/IUCN.UK.2010-4.RLTS.T176130A7184890.en. Downloaded on 08 June 2018.
"THE REPTILE DATABASE". Retrieved 2022-03-09.
"Hemidactylus chipkali".
Tiyapan, Kittisak Nui (20 April 2018). Thai Grammar, Poetry and Dictionary, in a New Romanised System. Lulu.com. p. 102. ISBN 9789741718610 – via Google Books.
Newbery, Brock; Jones, Darryl N. (2007). Lunney, Daniel (ed.). Presence of Asian House Gecko Hemidactylus frenatus across an urban gradient in Brisbane: influence of habitat and potential for impact on native gecko species. Pest or Guest: The Zoology of Overabundance. Royal Zoological Society of New South Wales. pp. 59–65. doi:10.7882/fs.2007.009. hdl:10072/18554. ISBN 9780980327212.
Keim, Lauren (2002). The spatial distribution of the introduced Asian House Gecko (Hemidactylus frenatus) across suburban/forest edges (BSc Honors thesis). Unpublished Honours Thesis, Department of Zoology and Entomology, the University of Queensland.
Marcellini, Dale (1971). "Activity Patterns of the Gecko Hemidactylus frenatus". Copeia. 1971 (4): 631–635. doi:10.2307/1442631. JSTOR 1442631.
McKay, J. Lindley; Griffiths, Anthony D.; Crase, Beth (December 2009). "Distribution and habitat Use by Hemidactylus frenatus Duméril and Bibron (Gekkonidae) in the Northern Territory, Australia". The Beagle: Records of the Museums and Art Galleries of the Northern Territory. 25: 111–116. doi:10.5962/p.287454. S2CID 130714348.
Gallina-Tessaro, Patricia; Ortega-Rubio, Alfredo; Alvarez-Cardenas, Sergio; Arnaud, Gustavo (1998). "Colonization of Socorro Island (Mexico), by the tropical house gecko Hemidactylus frenatus (Squamata:Gekkonidae)". Review of Tropical Biology. 47: 237–238.
Rödder, Dennis; Solé, Micro; Böhme, Wolfgang (December 2008). "Predicting the potential distributions of two alien invasive Housegeckos (Gekkonidae: Hemidactylus frenatus, Hemidactylus mabouia)" (PDF). North-Western Journal of Zoology. 4 (2): 236–246.
Brisbane, Jeanelle L. K.; Dewynter, Maël; Angin, Baptiste; Questel, Karl; van den Burg, Matthijs P. (17 June 2021). "Importation of ornamental plants facilitates establishment of the CommonHouse Gecko, Hemidactylus frenatus Duméril & Bibron, in the Lesser Antilles". Caribbean Herpetology: 1–5. doi:10.31611/ch.77. eISSN 2333-2468.
Griffing, Aaron H.; Griffing, David H.; Lesmond, Stephen; Gamble, Tony (21 February 2022). "Establishment of the Common House Gecko, Hemidactylus frenatus Duméril & Bibron, on Saint Lucia". Caribbean Herpetology: 1–3. doi:10.31611/ch.83. eISSN 2333-2468.
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References
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