Crocodylinae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Subsectio: Pseudosuchia
Cladus: Suchia
Cladus: Paracrocodylomorpha
Cladus: Loricata
Superordo: Crocodylomorpha
Ordo: Crocodilia
Cladus: Mesoeucrocodylia
Cladus: Neosuchia
Subordo: Eusuchia

Familia: Crocodylidae
Subfamilia: Crocodylinae
Genera: Crocodylus – Osteolaemus – †Aldabrachampsus – †Euthecodon – †Rimasuchus – †Voay

Name

Crocodylinae Cuvier, 1807

References

Cuvier 1807: Ann. Mus. Hist. Nat. Paris:64.
McAliley, L.R.; Willis, R.E.; Ray, D.A.; White, P.S.; Brochu, C.A.; Densmore, L.D., III 2006: Are crocodiles really monophyletic? — Evidence for subdivisions from sequence and morphological data. Molecular phylogenetics and evolution, 39(1): 16–32. DOI: 10.1016/j.ympev.2006.01.012

Vernacular names
ak: Denkyem
dansk: Krokodille
Deutsch: Echte Krokodile
eʋegbe: Elo
English: Crocodiles
español: Cocodrilos
suomi: Krokotiilit
français: Crocodiles
日本語: クロコダイル亜科
မြန်မာဘာသာ: မိကျောင်း
Nederlands: Krokodil
norsk: Krokodille
polski: Krokodyle
português: Crocodilo
svenska: Krokodiler
ไทย: จระเข้

Crocodylinae is a subfamily of true crocodiles within the family Crocodylidae, and is the sister taxon to Osteolaeminae (dwarf crocodiles and slender-snouted crocodiles).
Taxonomy

Crocodylinae was cladistically defined by Christopher Brochu in 1999 as Crocodylus niloticus (the Nile crocodile) and all crocodylians more closely related to it than to Osteolaemus tetraspis (the Dwarf crocodile).[1][2] This is a stem-based definition, and is the sister taxon to Osteolaeminae.

Crocodylinae contains the extant genus Crocodylus. It is disputed as to whether is also includes Mecistops (slender-snouted crocodiles),[2][3] or the extinct genus Voay.[4]
Phylogeny

Some morphological studies have recovered Mecistops as a basal member of Crocodylinae, more closely related to Crocodylus than to Osteolaemus and the other members of Osteolaeminae,[2][3] as shown in the cladogram below.[3]

Crocodylidae	Osteolaeminae Rimasuchus lloydi† Voay robustus† Osteolaemus osborni Osborn’s dwarf crocodile Osteolaemus tetraspis Dwarf crocodile "Crocodylus" gariepensis† Brochuchus parvidens† Brochuchus pigotti† Euthecodon arambourgi† Euthecodon brumpti† Crocodylinae Mecistops cataphractus West African slender-snouted crocodile Crocodylus thorbjarnarsoni† Crocodylus anthropophagus† Crocodylus niloticus Nile crocodile Crocodylus checchiai† Crocodylus moreletii Morelet's crocodile Crocodylus intermedius Orinoco crocodile Crocodylus acutus American crocodile Crocodylus rhombifer Cuban crocodile Crocodylus palaeindicus† Crocodylus palustris Mugger crocodile Crocodylus ossifragus† Crocodylus siamensis Siamese crocodile Crocodylus mindorensis Philippine crocodile Crocodylus johnstoni Freshwater crocodile Crocodylus porosus Saltwater crocodile Crocodylus raninus Borneo crocodile Crocodylus novaeguineae New Guinea crocodile	Paleoafrican Crocodylus Neotropical Crocodylus Indo-Pacific Crocodylus

The below cladogram is based on a 2021 study using paleogenomics that extracted DNA from the extinct Voay, recovering it as a member of Crocodylinae.[4]

Crocodylidae

Osteolaeminae Mecistops cataphractus West African slender-snouted crocodile Euthecodon† Brochuchus† Rimasuchus† Osteolaemus osborni Osborn’s dwarf crocodile Osteolaemus tetraspis Dwarf crocodile Crocodylinae Voay† Crocodylus Crocodylus anthropophagus† Crocodylus thorbjarnarsoni† Crocodylus palaeindicus† Crocodylus Tirari Desert† Crocodylus johnstoni Freshwater crocodile Crocodylus novaeguineae New Guinea crocodile Crocodylus mindorensis Philippine crocodile Crocodylus porosus Saltwater crocodile Crocodylus siamensis Siamese crocodile Crocodylus palustris Mugger crocodile Crocodylus checchiai† Crocodylus falconensis† Crocodylus suchus West African crocodile Crocodylus niloticus Nile crocodile Crocodylus moreletii Morelet's crocodile Crocodylus rhombifer Cuban crocodile Crocodylus intermedius Orinoco crocodile Crocodylus acutus American crocodile

(crown group)

References

Brochu, C. A.; Storrs, G. W. (2012). "A giant crocodile from the Plio-Pleistocene of Kenya, the phylogenetic relationships of Neogene African crocodylines, and the antiquity of Crocodylus in Africa". Journal of Vertebrate Paleontology. 32 (3): 587–602. doi:10.1080/02724634.2012.652324. S2CID 85103427.
Rio, Jonathan P.; Mannion, Philip D. (6 September 2021). "Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem". PeerJ. 9: e12094. doi:10.7717/peerj.12094. PMC 8428266. PMID 34567843.
Azarra, Beatrice; Boschian, Giovanni; Brochu, Christopher; Delfino, Massimo; Iurino, Dawid Adam; Kimambo, Jackson Stanley; Manzi, Giorgio; Masao, Fidelis T.; Menconero, Sofia; Njau, Jackson K; Cherin, Marco (2021). "A new cranium of Crocodylus anthropophagus from Olduvai Gorge, northern Tanzania". Rivista Italiana di Paleontologia e Stratigrafia (Research in Paleontology and Stratigraphy). 127 (2): 275–295. doi:10.13130/2039-4942/15771.
Hekkala, E.; Gatesy, J.; Narechania, A.; Meredith, R.; Russello, M.; Aardema, M. L.; Jensen, E.; Montanari, S.; Brochu, C.; Norell, M.; Amato, G. (2021-04-27). "Paleogenomics illuminates the evolutionary history of the extinct Holocene "horned" crocodile of Madagascar, Voay robustus". Communications Biology. 4 (1): 505. doi:10.1038/s42003-021-02017-0. ISSN 2399-3642. PMC 8079395. PMID 33907305.
"Species | the Reptile Database".