Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Cladus: Unidentata Episquamata
Subordo: Lacertoidea
Infraordo: Lacertibaenia
Cladus: Amphisbaenia
Familia: Blanidae
Genus: Blanus
Species: B. cinereus – B. mariae – B. mettetali – B. strauchi – B. tingitanus
Name
Blanus Wagler, 1830
References
Albert, E.M.; Fernández, A. 2009: Evidence of cryptic speciation in a fossorial reptile: description of a new species of Blanus (Squamata: Amphisbaenia: Blanidae) from the Iberian Peninsula. Zootaxa, 2234: 56–68. Abstract & excerpt
Vernacular names
English: Worm lizard
español: Culebrilla ciega
Blanus is a genus of amphisbaenians found in the Mediterranean region of Europe and North Africa.[2] Like other amphisbaenians, Blanus species are specialized for a subterranean existence, with long, slender bodies, reduced limbs, and rudimentary eyes. Their skulls are powerfully constructed, allowing them to push through soil to create a burrow. Their jaws are well-developed, with large, recurved teeth and a pair of canine-like teeth in the upper jaw.
Five extant species are currently known. The relationships of Blanus to other worm-lizards are not clear. The genus was formerly included in the Amphisbaenidae. More recent analyses suggest that blanids are more primitive, and are either related to Bipes[3] or represent an even more ancient lineage.[4]
A number of fossils from Europe have been referred either to Blanus or to the Blanidae.[5]
Species
Evolutionary tree of Blanidae; extinct species denoted with gray lines
The genus contains the following species:[6][7]
Blanus alexandri Sindaco, Kornilios, Sacchi & Lymberakis, 2014
Blanus aporus Werner, 1898
Blanus cinereus (Vandelli, 1797) – Iberian worm lizard
Blanus mariae Albert & Fernández, 2009
Blanus mendezi† Bolet et al., 2014
Blanus mettetali Bons, 1963 – Moroccan worm lizard
Blanus strauchi (Bedriaga, 1884) – Turkish worm lizard
Blanus tingitanus Busack, 1988
References
Wagler, J. 1830. Natürliches System der AMPHIBIEN, mit vorangehender Classification der SÄUGTHIERE und VÖGEL. Ein Beitrag zur vergleichenden Zoologie. Munich, Stuttgart, and Tübingen: J.G. Cotta. vi + 354 pp. + one plate. (Genus Blanus, p. 197).
Albert, E. M.; Zardoya, R.; García-París, M. (2007). "Phylogeographical and speciation patterns in subterranean worm lizards of the genus Blanus (Amphisbaenia: Blanidae)". Molecular Ecology. 16 (7): 1519–1531. doi:10.1111/j.1365-294X.2007.03248.x. PMID 17391273. S2CID 26071914.
Vidal, N.; Azvolinsky, A.; Cruaud, C.; Hedges, S. B. (2007-12-11). "Origin of tropical American burrowing reptiles by transatlantic rafting". Biology Letters. 4 (1): 115–118. doi:10.1098/rsbl.2007.0531. PMC 2412945. PMID 18077239.
Kearney, M.; Stuart, B. L. (2004). "Repeated evolution of limblessness and digging heads in worm lizards revealed by DNA from old bones". Proceedings of the Royal Society B: Biological Sciences. 271 (1549): 1677–1683. doi:10.1098/rspb.2004.2771. PMC 1691774. PMID 15306287.
Augé, M. L. (2012). "Amphisbaenians from the European Eocene: A biogeographical review". Palaeobiodiversity and Palaeoenvironments. 92 (4): 425–443. doi:10.1007/s12549-012-0104-6. S2CID 129023376.
Blanus at the Reptarium.cz Reptile Database. Accessed 18 January 2014.
Blanus Wikispecies.
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