Atheris ceratophora

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Subordo: Serpentes
Infraordo: Caenophidia
Superfamilia: Viperoidea

Familia: Viperidae
Subfamilia: Viperinae
Genus: Atheris
Species: Atheris ceratophora
Name

Atheris ceratophora Werner, 1896: 194

Holotype: BMNH 1946.1.18.23.

Type locality: “Usambara Mountains, Tanzania”.
References
Links

Uetz, P. & Hallermann, J. 2021. Atheris ceratophora. The Reptile Database. Accessed on 22 October 2018.

Vernacular names
English: Usambara Bush Viper

Atheris ceratophora is a venomous viper species endemic to a few mountain ranges in Tanzania. This used to be the only horned, arboreal viper known from Africa, until the discovery in 2011 of Atheris matildae, also found in Tanzania.[3] No subspecies are currently recognized.[6]

Description

It grows to a maximum total length (body + tail) of 54 cm (21 in). Females are slightly larger than males. The maximum total length for a male is reported to be 42 cm (17 in), the tail of which measured 8 cm (3.1 in).[7]

Easily recognized by a set of 3-5 horn-like superciliary scales above each eye. The rostral scale is more than twice as broad as high. There are 9 upper labials. The first 3 lower labials on each side are in contact with the one pair of chin shields.[7]

Midbody, the dorsal scales number 21-25. The ventral scale count is 142-152. There are 41-56 subcaudals.[7]

The color pattern consists of a yellowish-green, olive, gray or black ground color. This may or may not be overlaid by variable markings, sometimes in the form of irregular black spots or cross-bars that may be lined with yellow or white spots. The belly is dirty orange to almost black in color, sometimes with dark spots.[3]
Common names

Usambara bush viper,[3] horned bush viper,[4] eyelash bush viper,[5] Usambara mountain bush viper,[7] horned tree viper, Usambara tree viper.[8]
Geographic range

It is found in the Usambara and Uzungwe Mountains in Tanzania. It is probably also found in the Uluguru Mountains.[3]

The type locality is "Usambara" Mountains [Tanzania].[2]
Habitat

Found in grass and low bushes about 1 m (3.3 ft) above the ground[3] in woodlands and forests at altitudes of 700–2,000 m (2,300–6,600 ft).
Behavior

Like other Atheris species, it is probably active mainly at night, or at dawn or dusk.
References

Spawls, S. & Joger, U. (2010). "Atheris ceratophora". IUCN Red List of Threatened Species. 2010: e.T178563A7571666. doi:10.2305/IUCN.UK.2010-4.RLTS.T178563A7571666.en.
McDiarmid RW, Campbell JA, Touré T. 1999. Snake Species of the World: A Taxonomic and Geographic Reference, Volume 1. Washington, District of Columbia: Herpetologists' League. 511 pp. ISBN 1-893777-00-6 (series). ISBN 1-893777-01-4 (volume).
Spawls S, Branch B. 1995. The Dangerous Snakes of Africa. Ralph Curtis Books. Dubai: Oriental Press. 192 pp. ISBN 0-88359-029-8.
Atheris ceratophora at the Reptarium.cz Reptile Database. Accessed 2 August 2007.
Atheris ceratophora at The World Of Atheris. Accessed 8 September 2007.
"Atheris ceratophora". Integrated Taxonomic Information System. Retrieved 13 July 2006.
Mallow D, Ludwig D, Nilson G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Malabar, Florida: Krieger Publishing Company. ISBN 0-89464-877-2.

Brown JH. 1973. Toxicology and Pharmacology of Venoms from Poisonous Snakes. Springfield, Illinois: Charles C. Thomas. 184 pp. LCCCN 73-229. ISBN 0-398-02808-7.