Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: †Allotheria
Ordo: †Multituberculata
Subordo: †Plagiaulacida
Familiae: †Albionbaataridae – †Allodontidae – †Eobaataridae – †Paulchoffatiidae – †Pinheirodontidae – †Plagiaulacidae – †Zofiabaataridae
Familiae incertae sedis: †Hahnodontidae
Genera incertae familiae: †Monobaatar
Name
Plagiaulacida Simpson, 1925
Plagiaulacida is a group of extinct multituberculate mammals. Multituberculates were among the most common mammals of the Mesozoic, "the age of the dinosaurs". Plagiaulacids are a paraphyletic grouping, containing all multituberculates that lie outside of the advanced group Cimolodonta. They ranged from the Middle Jurassic Period to the early Late Cretaceous of the northern hemisphere. During the Cenomanian, they were replaced by the more advanced cimolodontans.[1]
Kielan-Jaworowska and Hurum (2001) divides “Plagiaulacida” into three informal lineages, the paulchoffatiids, the plagiaulicids, and the allodontids.
Allodontid line
The Allodontid line may be a superfamily, Allodontoidea.
Both allodontids and paulchoffatiids (below) were among the most basal of the plagiaulacids. The Allodontid line contains:
The family Allodontidae is known from two genera from the Upper Jurassic Morrison Formation of North America.
The family Zofiabaataridae contains a single genus, Zofiabaatar and is also from the Morrison Formation. The affinities of a further Morrison Formation genus, Glirodon, are unclear, but it's also within the Allodontid line.
Paulchoffatiid line
The Paulchoffatiid line may be a superfamily, Paulchoffatioidea.
Some remains from the Middle Jurassic of England might belong within this group. Representatives are best known from the Upper Jurassic, (especially from Guimarota, Portugal), though some were still extant during the Lower Cretaceous.
The genera of the family Paulchoffatiidae are divided into two of subfamilies, plus a couple of harder-to-place individuals:
Subfamily Paulchoffatiinae includes Paulchoffatia and its relatives. This taxon contains nine genera.
Subfamily Kuehneodontinae consists solely of the genus Kuehneodon, though there are half-a-dozen named species.
Other genera include Galveodon and Sunnyodon, both based on teeth from the Lower Cretaceous of Spain and England respectively.
Also referable to the paulchoffatiid line, but not the family itself, is the family Pinheirodontidae, which is known from Early Cretaceous teeth found in Iberia and England. As well as Rugosodon from the Middle-Late Jurassic of China.
Plagiaulacid line (possibly Superfamily Plagiaulacoidea)
Family Plagiaulacidae is known from the Upper Jurassic (North America) to Lower Cretaceous (Europe), being represented by Plagiaulax, Bolodon, and Morrisonodon.
Family Albionbaataridae is known from the Upper Jurassic to Lower Cretaceous of Europe and Asia, (China – undescribed, 2001). These were shrew-sized Multituberculates, with some similarities to the paulchoffis.
Members of the family Eobaataridae display dental similarities with members of Paracimexomys group, (Cimolodonta). They are known from the Upper Jurassic to Lower Cretaceous of Europe and Asia.
Sinobaatar was described after the study by Kielan-Jaworowska and Hurum (2001). The Mongolian word ‘baatar’ is frequently employed in the nomenclature of Multituberculates. This reflects the fact that many of the most complete fossils have been recovered from sites in Mongolia, though this more applies to members of the more derived Cimolodonta.
A couple of further genera possibly fit somewhere within “Plagiaulacida”. This has been tentatively proposed for Janumys of the Middle Cretaceous. Its contemporary, Ameribaatar, is of uncertain affinities. Both were first described late in 2001.
Taxonomy
Subclass †Allotheria Marsh, 1880
Order †Multituberculata Cope, 1884:
suborder †Plagiaulacida Simpson 1925
Family †Paulchoffatiidae Hahn, 1969
subfamily †Paulchoffatiinae Hahn, 1971
Genus †Paulchoffatia Kühne, 1961
Species †P. delgador Kühne, 1961
Genus †Pseudobolodon Hahn, 1977
Species †P. oreas Hahn, 1977
Species †P. krebsi Hahn & Hahn, 1994
Genus †Henkelodon Hahn, 1987
Species †H. naias Hahn, 1987
Genus †Guimarotodon Hahn, 1969
Species †G. leiriensis Hahn, 1969
Genus †Meketibolodon (Hahn, 1978) Hahn, 1993
Species †M. robustus (Hahn, 1978) Hahn, 1993
Genus †Plesiochoffatia Hahn & Hahn, 1999
Species †P. thoas Hahn & Hahn, 1998
Species †P. peparethos Hahn & Hahn, 1998
Species †P. staphylos Hahn & Hahn, 1998
Genus †Xenachoffatia Hahn & Hahn, 1998
Species †X. oinopion Hahn & Hahn, 1998
Genus †Bathmochoffatia Hahn & Hahn, 1998
Species †B. hapax Hahn & Hahn, 1998
Genus †Kielanodon Hahn, 1987
Species †K. hopsoni Hahn, 1987
Genus †Meketichoffatia Hahn, 1993
Species †M. krausei Hahn, 1993
Genus †Galveodon Hahn & Hahn, 1992
Species †G. nannothus Hahn & Hahn, 1992
Genus †Sunnyodon Kielan-Jaworowska & Ensom, 1992
Species †S. notleyi Kielan-Jaworowska & Ensom, 1992
Subfamily †Kuehneodontinae Hahn, 1971
Genus †Kuehneodon Hahn, 1969
Species †K. dietrichi Hahn, 1969
Species †K. barcasensis Hahn & Hahn, 2001
Species †K. dryas Hahn, 1977
Species †K. guimarotensis Hahn, 1969
Species †K. hahni Antunes, 1988
Species †K. simpsoni Hahn, 1969
Species †K. uniradiculatus Hahn, 1978
Family †Pinheirodontidae Hahn & Hahn, 1999
Genus †Pinheirodon Hahn & Hahn, 1999
Species †P. pygmaeus Hahn & Hahn, 1999
Species †P. vastus Hahn & Hahn, 1999
Genus †Bernardodon Hahn & Hahn, 1999
Species †B. atlantica Hahn & Hahn, 1999
Genus †Gerhardodon Kielan-Jaworowska & Ensom, 1992
Species †G. purbeckensis Kielan-Jaworowska & Ensom, 1992
Genus †Iberodon Hahn & Hahn, 1999
Species †I. quadrituberculatus Hahn & Hahn, 1999
Genus †Lavocatia Canudo & Cuenca-Bescós, 1996
Species †L. alfambrensis Canudo & Cuenca-Bescós, 1996
Genus †Ecprepaulax Hahn & Hahn, 1999
Species †E. anomala Hahn & Hahn, 1999
Family †Allodontidae Marsh, 1889
Genus †Ctenacodon Marsh, 1879
Species †C. serratus Marsh, 1879
Species †C. nanus Marsh, 1881
Species †C. laticeps Marsh, 1881
Species †C. scindens Simpson, 1928
Genus †Psalodon Simpson, 1926
Species †P. potens Marsh, 1887
Species †P. fortis Marsh, 1887
Species †P. marshi Simpson, 1929
Family †Zofiabaataridae Bakker, 1992
Genus †Zofiabaatar Bakker & Carpenter, 1990
Species †Z. pulcher Bakker & Carpenter, 1990
Family Incertae sedis
Genus †Glirodon Engelmann & Callison, 2001
Species †G. grandis Engelmann & Callison, 2001
Family †Plagiaulacidae Gill, 1872
Genus †Morrisonodon Hahn and Hahn, 2004
Species †M. brentbaatar Bakker, 1998
Genus †Plagiaulax Falconer, 1857
Species †P. becklesii Falconer, 1857
Genus †Bolodon Owen, 1871
Species †B. crassidens Owen, 1871
Species †B. falconeri Owen, 1871
Species †B. minor Falconer, 1857
Species †B. osborni Simpson, 1928
Species †B. elongatus Simpson, 1928
Family †Eobaataridae Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
Genus †Eobaatar
Species †E. magnus Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
Species †E. minor Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
Species †E. hispanicus Hahn & Hahn, 1992
Species †E. pajaronensis Hahn & Hahn, 2001
Genus †Loxaulax Simpson, 1928
Species †L. valdensis Simpson, 1928
Genus †Monobaatar Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
Species †M. mimicus Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
Genus †Parendotherium Crusafont Pairó & Adrover, 1966
Species †Parendotherium herreroi Crusafont Pairó & Adrover, 1966
Genus †Sinobaatar Hu & Wang, 2002
Species †Sinobaatar lingyuanensis Hu & Wang, 2002
Genus †Heishanobaatar Kusuhashi et al., 2010
Species †H. triangulus Kusuhashi et al., 2010
Genus †Teutonodon Martin et al 2016
Species †Teutonodon langenbergensis Martin et al., 2016
Family †Albionbaataridae Kielan-Jaworowska & Ensom, 1994
Genus †Albionbaatar Kielan-Jaworowska & Ensom, 1994
Species †A. denisae Kielan-Jaworowska & Ensom, 1994
Genus †Proalbionbaatar Hahn & Hahn, 1998
Species †P. plagiocyrtus Hahn & Hahn, 1998
Genus †Kielanobaatar Kusuhashi et al., 2010
Species †K. badaohaoensis Kusuhashi et al., 2010
Family †Arginbaataridae Hahn & Hahn, 1983
Genus †Arginbaatar Trofimov, 1980
Species †Arginbaatar dmitrievae Trofimov, 1980
References
Weaver, Lucas N.; Wilson, Gregory P.; Krumenacker, L. J.; Mclaughlin, Kayla; Moore, Jason R.; Varricchio, David J. (2019-03-04). "New multituberculate mammals from the mid-Cretaceous (lower Cenomanian) Wayan Formation of southeastern Idaho and implications for the early evolution of Cimolodonta". Journal of Vertebrate Paleontology. 39 (2): e1604532. doi:10.1080/02724634.2019.1604532. ISSN 0272-4634. S2CID 196655261.
Hahn G & Hahn R (2000), Multituberculates from the Guimarota mine, p. 97-107 in
Martin T & Krebs B (eds), Guimarota - A Jurassic Ecosystem, Published by Dr Friedrich Pfeil, Münich, Germany.
Kielan-Jaworowska Z & Hurum JH (2001), Phylogeny and Systematics of multituberculate mammals. Paleontology 44, p. 389-429.
Much of this information has been derived from [1] Multituberculata Cope, 1884.
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