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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Subsectio: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Euornithes
Cladus: Ornithuromorpha
Cladus: Ornithurae
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Ordo: Accipitriformes

Familia: Accipitridae
Subfamilia: Circaetinae
Genera: Circaetus - Spilornis - Terathopius - Pithecophaga

Name
Circaetinae

Circaetinae is a subfamily of the family Accipitridae which contains of a group of medium to large broad-winged birds of prey.[1][2] The group is sometimes treated as tribe Circaetini.[3][4] These birds mainly specialise in feeding on snakes and other reptiles, which is the reason most are referred to as "snake-eagles" or "serpent-eagles". The exceptions are the bateleur, a more generalised hunter, and the Philippine eagle, which preys on mammals and birds.

All but one of the subfamily are restricted to warmer parts of the Old World: Spilornis and Pithecophaga in south Asia, the others in Africa. The short-toed eagle Circaetus gallicus migrates between temperate Eurasia and Africa, as well as being resident in India.

They have hooked beaks for tearing flesh from their prey, strong legs and powerful talons. They also have extremely keen eyesight to enable them to spot potential prey from a distance.
Taxonomy

The subfamily Circaetinae was introduced in 1851 by the English zoologist Edward Blyth with Circaetus as the type genus.[5][6]

The genus level cladogram of the Circaetinae shown below is based on a molecular phylogenetic study of the Accipitridae by Therese Catanach and collaborators that was published in 2024.[7]

Circaetinae

Spilornis – serpent eagles (6 species)

Pithecophaga – Philippine eagle

Terathopius – Bateleur

Circaetus – snake eagles (6 species) + Dryotriorchis spectabilis Congo serpent eagle

The following taxonomy is based on the International Ornithological Congress.[8]

Image Genus Living Species
Spilornis G.R. Gray, 1840
  • Crested serpent eagle, S. cheela
    • Bawean serpent eagle, S. (cheela) baweanus
  • Great Nicobar serpent eagle, S. klossi
  • Mountain serpent eagle,S. kinabaluensis
  • Sulawesi serpent eagle, S. rufipectus
  • Philippine serpent eagle, S. holospilus
  • Andaman serpent eagle, S. elgini
Pithecophaga Ogilvie-Grant, 1896
  • Philippine eagle, P. jefferyi
Circaetus Vieillot, 1816
  • Short-toed snake eagle, C. gallicus
  • Beaudouin's snake eagle, C. beaudouini
  • Black-chested snake eagle, C. pectoralis
  • Brown snake eagle, C. cinereus
  • Southern banded snake eagle, C. fasciolatus
  • Western banded snake eagle, C. cinerascens
Dryotriorchis Shelley, 1874
  • Congo serpent eagle, D. spectabilis
Terathopius Lesson, 1830
  • Bateleur, T. ecaudatus
Eutriorchis Sharpe, 1875[note 1]
  • Madagascar serpent eagle, E. astur

Notes

Eutriorchis is genetically related to Gypaetus, and so might be classified with the Gypaetinae rather than Circaetinae. Lerner, Heather R. L.; Mindell, David P. (9 May 2006). "Accipitridae". The Tree of Life Web Project.

References

Lerner, Heather R.L.; Mindell, David P. (2005). "Phylogeny of eagles, Old World vultures, and other Accipitridae based on nuclear and mitochondrial DNA". Molecular Phylogenetics and Evolution. 37 (2): 327–346. Bibcode:2005MolPE..37..327L. doi:10.1016/j.ympev.2005.04.010. PMID 15925523.
Mindell, David P.; Fuchs, Jérôme; Johnson, Jeff A. (2018). "Phylogeny, taxonomy, and geographic diversity of diurnal raptors: Falconiformes, Accipitriformes, and Cathartiformes". Birds of Prey. pp. 3–32. doi:10.1007/978-3-319-73745-4_1. ISBN 978-3-319-73744-7.
Edward C., Dickinson; Remsen Jr., James V., eds. (2013). The Howard and Moore Complete Checklist of the Birds of the World. Vol. 1: Non-passerines (4th ed.). Aves Press.
"ACCIPITRIDAE - Kites, Hawks and Eagles". Trust for Avian Systematics. Retrieved 27 March 2024.
Blyth, Edward (1851). "Conspectus of the ornithology of India, Burma, and the Malayan Peninsular, inclusiveof Sindh, Asám, Ceylon, and the Nicobar Islands". Journal of the Asiatic Society of Bengal. 19: 317–342 [317, 327].
Bock, Walter J. (1994). History and Nomenclature of Avian Family-Group Names. Bulletin of the American Museum of Natural History. Vol. 222. New York: American Museum of Natural History. p. 132.
Catanach, T.A.; Halley, M.R.; Pirro, S. (2024). "Enigmas no longer: using ultraconserved elements to place several unusual hawk taxa and address the non-monophyly of the genus Accipiter (Accipitriformes: Accipitridae)". Biological Journal of the Linnean Society: blae028. doi:10.1093/biolinnean/blae028.
Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (December 2023). "Hoatzin, New World vultures, Secretarybird, raptors". IOC World Bird List Version 14.1. International Ornithologists' Union. Retrieved 3 April 2024.

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