Anser caerulescens

Anser caerulescens (*)

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Pangalloanserae
Cladus: Galloanseres
Ordo: Anseriformes

Familia: Anatidae
Subfamilia: Anserinae
Genus: Anser
Species: Anser caerulescens
Subspecies: A. c. atlanticus – A. c. caerulescens
Name

Anser caerulescens (Linnaeus, 1758)
Synonyms

Anas caerulescens (protonym)
Chen caerulescens
Anser hyperboreus Pallas, 1769
Chen hyperborea

References

Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Tomus I. Holmiæ (Stockholm): impensis direct. Laurentii Salvii. 824 pp. DOI: 10.5962/bhl.title.542 BHL p. 124 BHL Reference page.
IOC

Vernacular names
български: Снежна гъска
brezhoneg: Gwaz-erc'hegi
čeština: Husa sněžní
dansk: Snegås
Deutsch: Schneegans
English: Snow Goose
Esperanto: Neĝansero
español: Ansar nival
suomi: Lumihanhi
Nordfriisk: Snägus
français: Oie des neiges
galego: Ganso das neves
magyar: Sarki lúd
Iñupiak: Kafuq
íslenska: Snjógæs
italiano: Oca delle nevi
日本語: ハクガン
한국어: 흰기러기
lietuvių: Snieginė žąsis
Nederlands: Sneeuwgans
norsk: Snøgås
polski: Śnieżyca większa
Piemontèis: Òca dla fiòca
português: Ganso-das-neves
русский: Белый гусь
саха тыла: Үрүҥ хаас
davvisámegiella: Muohtačuonjá
slovenčina: Hus snežná
slovenščina: Snežna gos
svenska: Snögås
Türkçe: Kar kazı
Zeêuws: Snieuwhanze
中文: 雪雁

The snow goose (Anser caerulescens) is a species of goose native to North America. Both white and dark morphs exist, the latter often known as blue goose. Its name derives from the typically white plumage. The species was previously placed in the genus Chen, but is now typically included in the "gray goose" genus Anser.[2][3]

Snow geese breed north of the timberline in Greenland, Canada, Alaska, and the northeastern tip of Siberia, and spend winters in warm parts of North America from southwestern British Columbia through parts of the United States to Mexico.[4] Snow goose populations increased dramatically in the 20th century.

Taxonomy

In 1750 the English naturalist George Edwards included an illustration and a description of the snow goose in the third volume of his A Natural History of Uncommon Birds. He used the English name "The blue-winged goose". Edwards based his hand-coloured etching on a preserved specimen that had been brought to London from the Hudson Bay area of Canada by James Isham.[5] When in 1758 the Swedish naturalist Carl Linnaeus updated his Systema Naturae for the tenth edition, he placed the snow goose with the ducks and geese in the genus Anas. Linnaeus included a brief description, coined the binomial name Anas caerulescens and cited Edwards' work.[6] The snow goose is now placed in the genus Anser that was introduced by the French zoologist Mathurin Jacques Brisson in 1760.[7][8] The scientific name is from the Latin anser, "goose", and caerulescens, "bluish", derived from caeruleus, "dark blue".[9] The snow goose is the sister species to Ross's goose (Anser rossii).[10]

Two subspecies are recognised:[8][11]

A. c. caerulescens (Linnaeus, 1758) – lesser snow goose – breeds in northeast Siberia, north Alaska and northwest Canada, winters in south USA, north Mexico and Japan
A. c. atlanticus (Kennard, 1927) – greater snow goose – breeds in northeast Canada and northwest Greenland, winters in northeast USA

The greater snow goose is distinguished from the nominate form by being slightly larger. It nests farther north and east. The lesser snow goose can be found in two colour phases, the normal white-coloured animals and a dark grey-coloured "blue" phase. The greater snow goose is rarely seen in a blue phase.[12]
Description
Snow goose at the Gray Lodge Wildlife Area in Northern California

The snow goose has two color plumage morphs, white (snow) or gray/blue (blue), thus the common description as "snows" and "blues". White-morph birds are white except for black wing tips, but blue-morph geese have bluish-grey plumage replacing the white except on the head, neck and tail tip. The immature blue phase is drab or slate-gray with little to no white on the head, neck, or belly. Both snow and blue phases have rose-red feet and legs, and pink bills with black tomia ("cutting edges"), giving them a black "grin patch". The colors are not as bright on the feet, legs, and bill of immature birds. The head can be stained rusty-brown from minerals in the soil where they feed. They are very vocal and can often be heard from more than a mile away.

White- and blue-morph birds interbreed and the offspring may be of either morph. These two colors of geese were once thought to be separate species; since they interbreed and are found together throughout their ranges, they are now considered two color phases of the same species. The color phases are genetically controlled. The dark phase results from a single dominant gene and the white phase is homozygous recessive. When choosing a mate, young birds will most often select a mate that resembles their parents' coloring. If the birds were hatched into a mixed pair, they will mate with either color phase.

The species is divided into two subspecies on the basis of size and geography. Size overlap has caused some to question the division.[4] The smaller subspecies, the lesser snow goose (C. c. caerulescens), lives from central northern Canada to the Bering Straits area. The lesser snow goose stands 64 to 79 cm (25 to 31 in) tall and weighs 2.05 to 2.7 kg (4.5 to 6.0 lb). The larger subspecies, the greater snow goose (C. c. atlanticus), nests in northeastern Canada. It averages about 3.2 kg (7.1 lb) and 79 cm (31 in), but can weigh up to 4.5 kg (9.9 lb). The wingspan for both subspecies ranges from 135 to 165 cm (53 to 65 in).
Breeding

Long-term pair bonds are usually formed in the second year, although breeding does not usually start until the third year. Females are strongly philopatric, meaning they will return to the place they hatched to breed.

Snow geese often nest in colonies. Nesting usually begins at the end of May or during the first few days of June, depending on snow conditions. The female selects a nest site and builds the nest on an area of high ground. The nest is a shallow depression lined with plant material and may be reused from year to year. After the female lays the first of three to five eggs, she lines the nest with down. The female incubates for 22 to 25 days, and the young leave the nest within a few hours of hatching.

The young feed themselves, but are protected by both parents. After 42 to 50 days they can fly, but they remain with their family until they are two to three years old.

Where snow geese and Ross's geese breed together, as at La Pérouse, they hybridize at times, and hybrids are fertile. Rare hybrids with the greater white-fronted goose, Canada goose, and cackling goose have been observed.[4]
Migration

Snow geese breed from late May to mid-August, but they leave their nesting areas and spend more than half the year on their migration to-and-from warmer wintering areas. During spring migration (the reverse migration), large flocks of snow geese fly very high and migrate in large numbers along narrow corridors, more than 3,000 mi (4,800 km) from traditional wintering areas to the tundra.

The lesser snow goose travels through the Central Flyway, Mississippi Flyway, and Pacific Flyway across prairie and rich farmland to their wintering grounds on grassland and agricultural fields across the United States and Mexico, especially the Gulf coastal plain. The larger and less numerous greater snow goose travels through the Atlantic Flyway and winters on a relatively more restricted range on the Atlantic coastal plain. Traditionally, lesser snow geese wintered in coastal marsh areas where they used their short but strong bills to dig up the roots of marsh grasses for food. However, they have also since shifted inland towards agricultural areas, likely the cause behind the unsustainable population increase in the 20th century. This shift may help to contribute to increased goose survival rates, leading to overgrazing on tundra breeding grounds.[13]

In March 2015, 2,000 snow geese were killed in northern Idaho from an avian cholera epidemic while flying their spring migration to northern Canada.[14]
Vagrancy

The snow goose is a rare vagrant to Europe but for a frequent escape from collections and an occasional feral breeder. Snow geese are visitors to the British Isles where they are seen regularly among flocks of brant, barnacle goose, and greater white-fronted goose. There is also a feral population in Scotland from which many vagrant birds in Britain seem to derive.

In Central America, vagrants are frequently encountered during winter.[15]
Ecology

Outside of the nesting season, they usually feed in flocks. In winter, snow geese feed on left-over grain in fields. They migrate in large flocks, often visiting traditional stopover habitats in spectacular numbers. Snow geese frequently travel and feed alongside greater white-fronted geese; in contrast, the two tend to avoid travelling and feeding alongside Canada geese, which are often heavier birds.[citation needed]

The population of greater snow geese was in decline at the beginning of the 20th century, but has now recovered to sustainable levels. Snow geese in North America have increased to the point where the tundra breeding areas in the Arctic and the saltmarsh wintering grounds are both becoming severely degraded,[16] and this affects other species using the same habitat.

Major nest predators include Arctic foxes and skuas.[17] The biggest threat occurs during the first couple of weeks after the eggs are laid and then after hatching. The eggs and young chicks are vulnerable to these predators, but adults are generally safe. They have been seen nesting near snowy owl nests, which is likely a solution to predation. Their nesting success was much lower when snowy owls were absent, leading scientists to believe that the owls, since they are predatory, were capable of keeping competing predators away from the nests. A similar association as with the owls has been noted between geese and rough-legged hawks.[17] Additional predators at the nest have reportedly included wolves, coyotes and all three North American bear species.[18][19] Few predators regularly prey on snow geese outside of the nesting season, but bald eagles (as well as possibly golden eagles) will readily attack wintering geese.
Population

The breeding population of the lesser snow goose exceeds 5 million birds, an increase of more than 300% since the mid-1970s. The population is increasing at a rate of more than five percent per year. Non-breeding geese (juveniles or adults that fail to nest successfully) are not included in this estimate, so the total number of geese is likely higher. Lesser snow goose population indices are the highest they have been since population records have been kept, and evidence suggests that large breeding populations are spreading to previously untouched sections of the Hudson Bay coastline. The cause of this overpopulation may be the heavy conversion of land from forest and prairie to agricultural usage in the 20th century.

Since the late 1990s, efforts have been underway in the U.S. and Canada to reduce the North American population of lesser snow and Ross's geese to sustainable levels due to the documented destruction of tundra habitat in Hudson Bay and other nesting areas. The Light Goose Conservation Order was established in 1997 and federally mandated in 1999. Increasing hunter bag limits, extending the length of hunting seasons, and adding new hunting methods have all been successfully implemented, but have not reduced the overall population of snow geese in North America.[20][21]
Conservation order for light geese

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The late 1990s was when the mid-continent population of snow geese was recognized as causing significant damage to the arctic and sub-arctic breeding grounds which was also causing critical damage to other varieties of waterfowl species and other wildlife that uses the arctic and sub-arctic grounds for home habitat. The increase in population in substantial amounts raised concern to then DU chief biologist Dr. Bruce Batt who was part of a committee that put together various data and submitted it to the U.S. Fish and Wildlife Service and the Canadian Wildlife Service with the recommendation on ways to combat the growing population and the damage that the snow geese were creating in the arctic breeding grounds.

The committee recommended relaxing hunting restrictions and giving hunters a better opportunity to harvest more snow geese on their way back to the breeding grounds in the spring. The suggested restrictions were to allow the use of electronic callers, unplugged shotguns, extended shooting hours, and no bag limits. Two years after the Light Goose Conservation Order was introduced it was federally mandated in 1999.

References

BirdLife International (2018). "Anser caerulescens". IUCN Red List of Threatened Species. 2018: e.T22679896A131908897. doi:10.2305/IUCN.UK.2018-2.RLTS.T22679896A131908897.en. Retrieved 19 November 2021.
Ogilvie, Malcolm A.; Young, Steve (2002). Wildfowl of the World. London: New Holland Publishers. p. 38. ISBN 978-1-84330-328-2.
Kear, Janet (2005). Ducks, Geese and Swans. Vol. 1. Oxford: Oxford University Press. p. 297. ISBN 978-0-19-861008-3.
Mowbray, Thomas B.; Fred, Cooke; Barbara, Ganter (2000). "Snow Goose (Chen caerulescens)". In Poole, A. (ed.). The Birds of North America Online. Ithaca: Cornell Lab of Ornithology. Retrieved 10 April 2009.
Edwards, George (1743). A Natural History of Uncommon Birds. Vol. Part 3. London: Printed for the author at the College of Physicians. p. 152, Plate 152.
Linnaeus, Carl (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1 (10th ed.). Holmiae (Stockholm): Laurentii Salvii. p. 124.
Brisson, Mathurin Jacques (1760). Ornithologie, ou, Méthode Contenant la Division des Oiseaux en Ordres, Sections, Genres, Especes & leurs Variétés (in French and Latin). Paris: Jean-Baptiste Bauche. Vol. 1, p. 58, Vol. 6, p. 261.
Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (2020). "Screamers, ducks, geese & swans". IOC World Bird List Version 10.2. International Ornithologists' Union. Retrieved 6 October 2020.
Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 48, 83. ISBN 978-1-4081-2501-4.
Ottenburghs, J.; Megens, H.-J.; Kraus, R.H.S.; Madsen, O.; van Hooft, P.; van Wieren, S.E.; Crooijmans, R.P.M.A.; Ydenberg, R.C.; Groenen, M.A.M.; Prins, H.H.T. (2016). "A tree of geese: A phylogenomic perspective on the evolutionary history of True Geese". Molecular Phylogenetics and Evolution. 101: 303–313. doi:10.1016/j.ympev.2016.05.021. PMID 27233434.
Kenneth F. Abraham, Robert L. Jefferies (1997). Arctic Ecosystems in Peril: Report of the Arctic Goose Habitat Working Group. Part II High Goose Populations: causes, impacts and implications (PDF) (Report). U.S. Fish and Wildlife Service and Canadian Wildlife Service. p. 1. Retrieved 15 November 2020.
McKelvey, Rick; Leafloor, Jim; Alisauskas, Ray (2010). Lesser Snow Goose. Hinterland Who's Who. Canadian Wildlife Federation. ISBN 978-0-662-17199-7. Retrieved 15 November 2020.
"Light Goose Dilemma". www.ducks.org. Retrieved 2018-01-01.
"2,000 snow geese drop dead from the sky in Idaho". 18 March 2015.
Herrera, Néstor; Rivera, Roberto; Ibarra Portillo, Ricardo; Rodríguez, Wilfredo (2006). "Nuevos registros para la avifauna de El Salvador" [New records for the avifauna of El Salvador] (PDF). Boletín de la Sociedad Antioqueña de Ornitología (in Spanish and English). 16 (2): 1–19.
"Snow geese degrade tundra". The New York Times. 22 September 2014.
Tremblay, J.-P.; Gauthier, G.; Lepage, D.; Desrochers, A. (1997). "Factors Affecting Nesting Success in Greater Snow Geese: Effects of Habitat and Association with Snowy Owls". Wilson Bulletin. 109 (3): 449–461. JSTOR 4163840.
Coakley, Amber (3 March 2009). "Duck Duck Goose – Snow Goose". Birders Lounge.
Johnson, Stephen R.; Noel, Lynn E. (2005). "Temperature and Predation Effects on Abundance and Distribution of Lesser Snow Geese in the Sagavanirktok River Delta, Alaska". Waterbirds. 28 (3): 292. doi:10.1675/1524-4695(2005)028[0292:TAPEOA]2.0.CO;2. ISSN 1524-4695. JSTOR 4132542.
"Too Many Snow Geese | Central Flyways". central.flyways.us. Retrieved 2017-12-31.

"Light Goose Dilemma". www.ducks.org. Retrieved 2017-12-31.

Further reading
Johnson, Mike (16 July 1997). "The snow goose population problem". Northern Prairie Wildlife Research Center Online.