Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Chelicerata
Classis: Arachnida
Ordo: Araneae
Subordo: Opisthothelae
Infraordo: Araneomorphae
Taxon: Neocribellatae
Series: Entelegynae
Superfamilia: Lycosoidea
Familia: Pisauridae
Genera: Afropisaura – Archipirata – Architis – Bradystichus – Campostichommides – Caripetella – Charminus – Chiasmopes – Cispinilus – Cispius – Cladycnis – Conakrya – Dendrolycosa – Dianpisaura – Dolomedes – Eucamptopus – Euprosthenops – Euprosthenopsis – Eurychoera – Hala – Hygropoda – Ilipula – Inola – Maypacius – Megadolomedes – Nilus – Nukuhiva – Papakula – Paracladycnis – Perenethis – Phalaeops – Pisaura – Pisaurina – Polyboea – Qianlingula – Ransonia – Rothus – Sphedanus – Stoliczka – Tallonia – Tapinothele – Tapinothelella – Tapinothelops – Tetragonophthalma – Thalassiopsis – Thalassius – Thaumasia – Tinus – Tolma – Voraptipus – Vuattouxia – Walrencea
Name
Pisauridae Simon, 1890
Synonymy
Halidae Jocqué, 1994
References
Franenau, V.W. & Lehtinen, P.T. 2015. Nukuhiva Berland, 1935 is a troglobitic wolf spider (Araneae: Lycosidae), not a nursery-web spider (Pisauridae). Zootaxa 4028(1): 129–135. DOI: 10.11646/zootaxa.4028.1.6. Preview (PDF) Reference page.
Jäger, P. 2010: Papakula and Hesydrimorpha: how two spider genera were described from the same species collected from the same locality (Araneae: Pisauridae). Zootaxa, 2551: 65–68. Preview PDF
Jocqué, R. 1994: Halidae, a new spider family from Madagascar (Araneae). Bulletin of the British Arachnological Society 9(9): 281–289. Reference page.
Jocqué, R.& Dippenaar-Schoeman, A.S. 2006. Spider families of the world. Tervuren, Belgium: Musée royal de l'Afrique centrale; [Pretoria, South Africa]: ARC-PPRI. ISBN 978-9075-89485-1. Reference page. [See p. 212, synonymy of Halidae]
Paquin, P.; Vink, C.J.; Dupérré, N. 2010: Spiders of New Zealand: annotated family key & species list. Manaaki Whenua Press, Lincoln, New Zealand. ISBN 9780478347050
Silva, E.L.C.D.; Sierwald, P. 2013: On three monotypic nursery web spider genera from Madagascar with first description of the male of Tallonia picta Simon, 1889 and redescription of the type-species of Paracladycnis Blandin, 1979 and Thalassiopsis Roewer, 1955 (Araneae: Lycosoidea: Pisauridae). Zootaxa 3750(3): 277–288. DOI: 10.11646/zootaxa.3750.3.7 Reference page.
Tonini, L., da Silva, J.P., Filho, A.S. & Freitas, J. 2016. Replacement names for two preoccupied generic names in Arthropoda. Zootaxa 4097(1): 125–126. DOI: 10.11646/zootaxa.4097.1.7 Paywall. ResearchGate Open access. Reference page.
Links
Platnick, N. I. 2008. The World Spider Catalog, version 9.0. American Museum of Natural History. [1]
Vernacular names
English: Nursery web spider
한국어: 닷거미과
中文: 跑蛛科
Nursery web spiders (Pisauridae) is a family of araneomorph spiders first described by Eugène Simon in 1890.[1] They resemble wolf spiders (Lycosidae) except for several key differences. Wolf spiders have two very prominent eyes in addition to the other six, while a nursery web spider's eyes are all about the same size.[2] Additionally, female nursery web spiders carry their egg sacs with their jaws and pedipalps instead of attaching them to their spinnerets as wolf spiders do. When the eggs are about to hatch, a female spider builds a nursery "tent", places her egg sac inside, and stands guard outside, hence the family's common name. Like the wolf spiders, however, the nursery web spiders are roaming hunters that don't use webs for catching prey.
Species occur throughout the world except for extremely dry or cold environments, and are common just about everywhere. Many can walk on the surface of still bodies of water and may even dive beneath the surface temporarily to escape enemies. They can jump a distance of 5 to 6 inches (130 to 150 mm), but they have trouble climbing extremely smooth surfaces such as glass.
The name "nursery web spider" is especially given to the European species Pisaura mirabilis, but this family also includes fishing spiders and raft spiders. Adult female specimens may reach up to 15mm in length, excluding legs. The legs of the male are longer in relation to body size than those of the female.[3]
The female spider sometimes attempts to eat the male after mating. The male, to reduce the risk of this, often presents the female with a gift such as a fly when approaching in the hope that this will satisfy her hunger. Sometimes, this gift is a fake present intended to fool the female.[4] Males may wrap the fake gift in silk, to deceive the female to mate. Females can detect the fake gift and terminate mating, negating the male's deception in not giving a real gift.[5]
Genera
Main article: List of Pisauridae species
As of April 2019, the World Spider Catalog accepts the following genera:[6]
Afropisaura Blandin, 1976 — Africa
Archipirata Simon, 1898 — Turkmenistan, China
Architis Simon, 1898 — South America, Trinidad, Panama
Blandinia Tonini, Paulo da Silva, Serpa Filho & Freitas, 2016 — Madagascar
Bradystichus Simon, 1884 — New Caledonia
Caledomedes Raven & Hebron, 2018 — New Caledonia
Caripetella Strand, 1928 — Madagascar, Comoros
Charminus Thorell, 1899 — Africa
Chiasmopes Pavesi, 1883 — Ethiopia, Namibia, South Africa
Cispinilus Roewer, 1955 — Central Africa
Cispius Simon, 1898 — South Africa, Congo
Cladycnis Simon, 1898 — Canary Is.
Conakrya Schmidt, 1956 — Guinea
Dendrolycosa Doleschall, 1859 — Asia, Africa, Oceania
Dolomedes Latreille, 1804 — Africa, Oceania, South America, North America, Asia, Cuba
Eucamptopus Pocock, 1900 — India
Euprosthenops Pocock, 1897 — Africa, India
Euprosthenopsis Blandin, 1974 — Africa
Hala Jocqué, 1994 — Madagascar
Hygropoda Thorell, 1894 — Africa, Asia, Australia
Ilipula Simon, 1903 — Vietnam
Inola Davies, 1982 — Australia
Mangromedes Raven, 2018 — Australia
Maypacius Simon, 1898 — Africa
Megadolomedes Davies & Raven, 1980 — Australia
Nilus O. Pickard-Cambridge, 1876 — Asia, Africa
Ornodolomedes Raven & Hebron, 2018 — Australia
Papakula Strand, 1911 — Indonesia
Paracladycnis Blandin, 1979 — Madagascar
Perenethis L. Koch, 1878 — Asia, Comoros, Oceania
Phalaeops Roewer, 1955 — Mozambique, Djibouti
Pisaura Simon, 1886 — Asia
Pisaurina Simon, 1898 — United States, Canada, Cuba
Polyboea Thorell, 1895 — Asia
Qianlingula Zhang, Zhu & Song, 2004
Rothus Simon, 1898 — Israel, South Africa
Sphedanus Thorell, 1877 — Singapore, Malaysia, Indonesia
Stoliczka O. Pickard-Cambridge, 1885 — Pakistan
Tallonia Simon, 1889 — Madagascar
Tapinothele Simon, 1898 — Tanzania
Tapinothelella Strand, 1909 — South Africa
Tapinothelops Roewer, 1955 — Ethiopia
Tasmomedes Raven, 2018 — Australia
Tetragonophthalma Karsch, 1878 —
Thalassiopsis Roewer, 1955 — Madagascar
Thaumasia Perty, 1833 — Central America, South America, Mexico
Tinus F. O. Pickard-Cambridge, 1901 — India, Cuba, North America, Central America
Tolma Jocqué, 1994 — Madagascar
Voraptipus Roewer, 1955 — Mozambique
Vuattouxia Blandin, 1979 — Côte d'Ivoire
Walrencea Blandin, 1979 — South Africa
Some fossilized spiders have also been assigned to this family:[7]
†Eopisaurella Petrunkevitch, 1958 (Early Eocene; Baltic amber)
†Palaeoperenethis Seldon & Penney, 2009 (Ypresian, British Columbia, Canada)
Gallery
Nursery web spider carrying egg sac Nursery web spider carrying egg sac
Megadolomedes australianus with a dragonfly as prey
See also
List of Pisauridae species
References
Simon, E. (1890). Etudes arachnologiques.
Sierwald, P. (1997). "Phylogenetic analysis of Pisaurine nursery web spiders, with revisions of Tetragonophthalma and Perenethis (Araneae, Lycosidae, Pisauridae)" (PDF). The Journal of Arachnology. 25: 361–407.
Anderson, Alissa G.; Hebets, Eileen A. (2016). "Benefits of size dimorphism and copulatory silk wrapping in the sexually cannibalistic nursery web spider, Pisaurina mira". Biology Letters. 12 (2): 20150957. doi:10.1098/rsbl.2015.0957. PMC 4780555. PMID 26911340.
Male Spiders Scam Females with Gift-Wrapped Garbage
Albo, Maria J; Winther, Gudrun; Tuni, Cristina; Toft, Søren; Bilde, Trine (2011-11-14). "Worthless donations: male deception and female counter play in a nuptial gift-giving spider". BMC Evolutionary Biology. 11 (1): 329. doi:10.1186/1471-2148-11-329. PMC 3228764. PMID 22082300.
"Family: Pisauridae Simon, 1890". World Spider Catalog. Natural History Museum Bern. Retrieved 2019-04-23.
Dunlop, J.A.; Penney, D.; Jekel, D. (2015). "A summary list of fossil spiders and their relatives" (PDF). World Spider Catalog. Natural History Museum Bern. Retrieved 2016-03-15.
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