Typophorini

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Cladus: Phytophaga
Superfamilia: Chrysomeloidea

Familia: Chrysomelidae
Subfamilia: Eumolpinae
Tribus: Typophorini

Genera assigned to informal sections (after Seeno & Wilcox, 1982):

Section Callisinites: Callisina – Edistus
Section Metachromites: Ambohitsitondrona – Chrysopida – Clypeolaria – Coniomma – Deretrichia – Gressittana – Gressittella – Ivongionymus – Jansonius – Labasa – Lindinia – Metachroma – Metacolaspis – Micromolpus – Nakanaia – Phainodina – Phytorus – Phytorellus – Pyropida – Rhynchomolpus – Rhyparida – Rhyparidella – Sahantaha – Stizomolpus – Stygnobia – Vitibia
Section Nodostomites: Basilepta – Berliozita – Brachymolpus – Cheiridea – Chrysonopa – Epinodostoma – Mireditha – Nodina – Nodostella – Phaedroides – Pseudolpus – Pseudostonopa – Scotosus – Sedlacekia – Sybriacus – Weiselina – Zavadilia
Section Pagriites: Pagria
Section Typophorites: Afroeurydemus – Amblynetes – Ambodiriana – Andrahomana – Androyinus – Bastrhembus – Bedelia – Chloropterus – Cleoparida – Cleoporus – Cleorina – Colasita – Entreriosa – Eulychius – Eurydemus – Gaberella – Hemigymna – Hemyloticus – Ivongius – Keeta – Lebisiella – Lepidocolaspis – Mahatsinjoa – Majungaeus – Mandollia – Medvedemolpus – Meniellus – Menioporus – Menius – Microeurydemus – Microsyagrus – Mouhotina – Navanites – Nossioecus – Paraivongius – Paria – Periparia – Pheloticus – Phytoparia – Proliniscus – Pseudivongius – Pseudosyagrus – Rhembastichus – Rhembastus – Rhembivongius – Rhyparidula – Sandrananta – Sarum – Selmania – Stethotes – Syagrus – Thyrasia – Tijucana – Tricliona – Tricliophora – Trypocolaspis – Typophorus – Zohrana
Section incertae sedis: Iviva – Vadoniella

Genera not assigned to a section: Aphilenia – Atomyria – Graphops – Pagellia – Parascela
Name

Typophorini Baly, 1865

Type genus: Typophorus Chevrolat in Dejean, 1836
Synonymy

Typophorinae Baly, 1865: 433
Typophorites: Chapuis, 1874: 329
Typophoritae: Lefèvre, 1885: 129
Typophorini: Jacoby, 1908: 468
Callisinites Chapuis, 1874: 263
Callisinitae: Lefèvre, 1885: 63
Callisini: Jacoby, 1908: 362
Callisinini: Clavareau, 1914: 62
Nodostomites Chapuis, 1874: 261
Nodostomitae: Lefèvre, 1885: 56
Nodostomini: Jacoby, 1908: 291
Metachromites Chapuis, 1874: 295
Metachromitae: Lefèvre, 1885: 92
Metachromini: Jacoby, 1908: 378
Pagriites Lefèvre, 1884: lxvii
Pagriitae: Lefèvre, 1885: 62
Pagrini: Jacoby, 1908: 355
Pagriini: Clavareau, 1914: 61
Cheirideitae Lefèvre, 1885: 67
Nodinini Chen, 1940: 487
Basileptini Chûjô, 1956: 9
Nodini Selman, 1965: 145 [permanently invalid (Art. 39): based on preoccupied type genus]

References
Primary references

Baly, J.S. 1865. Attempt at a classification of the Eumolpidae. (Cont.). The Journal of Entomology 2: 433–442. BHL
Chapuis, F. 1874. Tome dixième. Famille des phytophages. In: Lacordaire, J.T. & Chapuis, F. (eds) Histoire naturelle des insectes. Genera des coléoptères, ou exposé méthodique et critique de tous les genres proposés jusqu'ici dans cet ordre d'insectes. Librairie Encyclopédique de Roret, Paris. iv + 445 pp. BHL Reference page.
Chen, S.H. 1940. Notes on Chinese Eumolpidae. Sinensia 11(5–6): 483–528.
Chûjô, M. 1956. A Taxonomic Study on the Chrysomelidae (Insecta: Coleoptera) from Formosa. Part VIII. Subfamily Eumolpinae. The Philippine Journal of Science 85(1): 1–180. Full article (PDF).
Clavareau, H. 1914. Pars 59: Chrysomelidae: 11. Eumolpinae. In: Junk, W. & Schenkling, S. (eds.). Coleopterorum Catalogus. Volumen XXIV. Chrysomelidae I. Berlin: W. Junk. 215 pp. BHL. Reference page.
Jacoby, M. 1908. The Fauna of British India, including Ceylon and Burma. Coleoptera. Chrysomelidae. Vol. 1. London: Francis & Taylor. xx + 534 pp., 2 pls. Internet Archive. BHL Reference page.
Lefèvre, É. 1884. [new taxa]. Bulletin de la Société entomologique de France 1884: lxv–lxvii. BHL [Published 8 Oct 1884.]
Lefèvre, É. 1885. Eumolpidarum hucusque cognitarum catalogus, sectionum conspectu systematico, generum sicut et specierum nonnullarum novarum descriptionibus adjunctis. Mémoires de la Société Royale des Sciences de Liège (2) 11(16): 1–172. BHL Reference page.
Selman, B.J. 1965. A revision of the Nodini and a key to the genera of Eumolpidae of Africa (Coleoptera: Eumolpidae). Bulletin of the British Museum (Natural History). Entomology 16(3): 141–174. BHL. Full article (PDF). Reference page.

Additional references

Askevold, I.S. & LeSage, L. 1990. A taxonomic revision of the genus Jansonius Baly 1878: Taxonomic confusion and tribal reclassification (Coleoptera: Chrysomelidae: Eumolpinae). Revista Chilena de Entomología 18: 29–37. PDF. BHL. Reference page. [as "Nodinini"]
Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page.
Chaboo, C.S. & Flowers, R.W. 2015. Beetles (Coleoptera) of Peru: A Survey of the Families. Chrysomelidae: Eumolpinae Hope, 1840. Journal of the Kansas Entomological Society 88(3): 375–379. DOI: 10.2317/kent-88-03-375-379.1 Paywall. JSTOR Hybrid open access journal. Reference page.
Gressitt, J.L. 1969. Chrysomelid beetles from the Papuan subregion, 6 (Eumolpinae, 4). Pacific Insects 11(1): 1–31. PDF. Reference page. [as "Nodinini"]
Gómez-Zurita, J. 2017. Insights on the genus Acronymolpus Samuelson with new synonymies and exclusion of Stethotes Baly from the fauna of New Caledonia (Coleoptera, Chrysomelidae, Eumolpinae). Pp 65–75 in: Chaboo, C.S. & Schmitt, M. (Eds) 2017. Research on Chrysomelidae 7. Zookeys 720: 1–137. Reference page. ZooKeys 720: 65–75. DOI: 10.3897/zookeys.720.13582 Open access. Reference page.
Gómez-Zurita, J., Jolivet, P. & Vogler, A.P. 2005. Molecular systematics of Eumolpinae and the relationships with Spilopyrinae (Coleoptera, Chrysomelidae). Molecular phylogenetics and evolution 34(3): 584–600. DOI: 10.1016/j.ympev.2004.11.022 PDF.
Medvedev, L.N. 2009. New genera and species of Zeugophorinae and Eumolpinae (Coleoptera: Chrysomelidae) from New Guinea. Stuttgarter Beiträge zur Naturkunde A, Neue Serie 2: 371–408. PDF. Reference page. [Metachromini and Typophorini]
Medvedev, L.N. & Moseyko, A.G. 2003. A revision of the genus Phytorus Jacoby, 1884 (Chrysomelidae: Eumolpinae). Russian Entomological Journal 11(4): 401–407. PDF. Reference page. [as Metachromini and Typophorini]
Moseyko, A.G. 2010. Medvedemolpus gen.n. — a new genus of Eumolpinae (Coleoptera: Chrysomelidae) from Philippines. Russian Entomological Journal 19(1): 3–8. PDF. Reference page. [as "Nodinini"]
Moseyko, A.G. 2011. К познанию жуков-листоедов трибы Nodinini (Coleoptera, Chrysomelidae, Eumolpinae) с Филиппинских островов и О. Калимантан [To the knowledge of the leaf-beetle tribe Nodinini (Coleoptera, Chrysomelidae, Eumolpinae) from the Philippines and Borneo]. Entomologicheskoe Obozrenie 90(2): 358–374. (in Russian) [English transl. in: 2012. Entomological Review 92(3): 315–328. DOI: 10.1134/S0013873812030086 Paywall.] Reference page. [as "Nodinini"]
Moseyko, A.G. 2012. Замечания о роде Aphilenia Weise in Reitter, 1889 (Coleoptera, Chrysomelidae, Eumolpinae) с описанием нового вида из России [Comments on the genus Aphilenia Weise in Reitter, 1889 (Coleoptera, Chrysomelidae, Eumolpinae), with a description of a new species from Russia]. Entomologicheskoe Obozrenie 91(1): 99–105. (in Russian). Abstract. [English transl. in: Entomological Review 92(7): 773–781. DOI: 10.1134/S0013873812070044 Paywall.] Reference page. [as "Nodinini"]
Moseyko, A.G. & Sprecher-Uebersax, E. 2010. Subfamily Eumolpinae Hope, 1840. Pp. 619–643. In: Löbl, I. & Smetana, A. (eds.) Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea. Apollo Books, Stenstrup, Denmark. 924 pp. ISBN 978-87-88757-84-2. Google Books. Reference page. [as "Nodinini"]
Romantsov, P.V. & Moseyko, A.G. 2016. To the knowledge of the leaf-beetle genera Rhyparida and Tricliona (Coleoptera: Chrysomelidae: Eumolpinae) from Indochina and Malay Peninsula. Zoosystematica Rossica 25(2): 299–313. DOI: 10.31610/zsr/2016.25.2.299 Open access Reference page.
Romantsov, P.V. & Moseyko, A.G. 2019. New and little known species of Eumolpinae (Coleoptera: Chrysomelidae) from Northern Vietnam. Zootaxa 4609(2): 321–334. DOI: 10.11646/zootaxa.4609.2.7 Paywall Reference page.
Seeno, T.N. & Wilcox, J.A. 1982. Leaf beetle genera (Coleoptera: Chrysomelidae). Entomography 1: 1–221. Google Books (Snippet view only). Reference page. [as "Nodinini"]
Zoia, S. 2019. New data on African Eumolpinae with description of new taxa from the collections of the Natural History Museum of Erfurt (Insecta: Coleoptera: Chrysomelidae). Vernate 38: 305–333. PDF. Reference page.

Typophorini is a tribe of leaf beetles in the subfamily Eumolpinae. The tribe contains approximately 100 genera, which are found worldwide. Members of the tribe are mainly characterized by notches on the tibiae of the middle and hind legs, which are sometimes referred to as antenna cleaners. They also generally have a subglabrous body, as well as bifid pretarsal claws.[4][5]

Taxonomy

Following the leaf beetle classification of Seeno and Wilcox (1982), the genera of Typophorini are largely divided into five informal groups or "sections": Callisinites, Metachromites, Nodostomini, Pagriites and Typophorites, with some genera having incertae sedis placement within the tribe.[6]

In the Catalog of the leaf beetles of America North of Mexico, published in 2003, the section Scelodontites of Bromiini was transferred to Typophorini.[7]
Genera

These 76 genera belong to the tribe Typophorini:[8][9][10][11][12][13][14][15][16][17][18]

Afroeurydemus Selman, 1965[3]
Amblynetes Weise, 1904[19]
Aphilenia Weise in Reitter, 1889
Atomyria Jacobson, 1894[20]
Basilepta Baly, 1860[21]
Bedelia Lefèvre, 1875
Callisina Baly, 1860[21]
Cheiridea Baly, 1878[22]
Chloropterus Morawitz, 1861[23]
Chrysonopa Jacoby, 1908[24]
Chrysopida Baly, 1861[25]
Cleoparida Gressitt, 1967[26]
Cleoporus Lefèvre, 1884[27]
Cleorina Lefèvre, 1885[28]
Clypeolaria Lefèvre, 1885[29]
Coniomma Weise, 1922
Deretrichia Weise, 1913[30]
Edistus Lefèvre, 1884[31]
Entreriosa Bechyné, 1953[32]
Epinodostoma Bryant & Gressitt, 1957
Eurydemus Chapuis, 1874[2]
Gaberella Selman, 1965[3]
Graphops LeConte, 1859 (formerly in Adoxini)
Gressittana Medvedev, 2009[33]
Gressittella Medvedev, 2009[33]
Iviva Gressitt, 1969[16]
Jansonius Baly, 1878[22][34]
Keeta Maulik, 1931[35]
Labasa Bryant, 1925
Lindinia Lefèvre, 1893
Mandollia Selman, 1965[3]
Medvedemolpus Moseyko, 2010[12]
Meniellus Weise, 1903[36]
Menius Chapuis, 1874[2]
Metachroma Chevrolat in Dejean, 1836
Microeurydemus Pic, 1938
Micromolpus Gressitt, 1969[16]
Microsyagrus Pic, 1952
Mireditha Reitter, 1913
Mouhotina Lefèvre, 1885[28]
Nakanaia Gressitt, 1969[16]
Nodina Motschulsky, 1858
Nodostella Jacoby, 1908[24]
Pagellia Lefèvre, 1885[28]
Pagria Lefèvre, 1884[37]
Paraivongius Pic, 1936
Parascela Baly, 1878[22]
Paria LeConte, 1858
Periparia Bechyné, 1951[38]
Phaedroides Lefèvre, 1885[29]
Phainodina Gressitt, 1969[16]
Phytoparia Bechyné, 1957[15]
Phytorellus Medvedev & Moseyko, 2003[39]
Phytorus Jacoby, 1884[40]
Proliniscus Selman, 1965[3]
Pseudivongius Jacoby, 1897[41]
Pseudolpus Jacoby, 1884[40]
Pseudostonopa Jacoby, 1903
Rhembastus Harold, 1877[42]
Rhynchomolpus Gressitt, 1969[16]
Rhyparida Baly, 1861[25]
Rhyparidella Gressitt, 1969[16]
Sarum Selman, 1965[3]
Sedlacekia Gressitt, 1969[16]
Selmania Zoia, 2019[43] (= Massartia Selman, 1965[3])
Stethotes Baly, 1867[44]
Stizomolpus Gressitt, 1969[16]
Stygnobia Weise, 1922
Syagrus Chapuis, 1874[2]
Thyrasia Jacoby, 1884[40]
Tijucana Bechyné, 1957[15]
Tricliona Lefèvre, 1885[45]
Tricliophora Jacoby, 1904
Typophorus Chevrolat in Dejean, 1836
Vitibia Fairmaire, 1882
Zohrana Aslam, 1968[46]

Gallery

Typophorus nigritus

References

Baly, J. S. (1865). "Attempt at a classification of the Eumolpidae. (Cont.)". The Journal of Entomology. 2: 433–442.
Chapuis, F. (1874). "Tome dixième. Famille des phytophages". In Lacordaire, J.T.; Chapuis, F. (eds.). Histoire naturelle des Insectes. Genera des coléoptères. Paris: Librairie Encyclopédique de Roret. pp. i–iv, 1–455.
Selman, B. J. (1965). "A revision of the Nodini and a key to the genera of Eumolpidae of Africa (Coleoptera: Eumolpidae)" (PDF). Bulletin of the British Museum (Natural History) Entomology. 16 (3): 141–174. doi:10.5962/bhl.part.21864.
Gómez-Zurita, Jesús; Jolivet, Pierre; Vogler, Alfried P. (2005). "Molecular systematics of Eumolpinae and the relationships with Spilopyrinae (Coleoptera, Chrysomelidae)". Molecular Phylogenetics and Evolution. 34 (3): 584–600. doi:10.1016/j.ympev.2004.11.022. PMID 15683931. S2CID 439719.
Jolivet, Pierre; Lawrence, John F.; Verma, Krishna K.; Ślipiński, Adam (2014). "2.7.3 Eumolpinae C. G. Thomson, 1859". In Leschen, R.A.B.; Beutel, R.G. (eds.). Handbook of Zoology. Arthropoda: Insecta: Coleoptera: Volume 3: Morphology and Systematics (Phytophaga). Berlin - Boston: Walter de Gruyter. pp. 217–225. doi:10.1515/9783110274462.189. ISBN 978-3-11-027370-0.
Seeno, T.N.; Wilcox, J.A. (1982). "Leaf beetle genera (Coleoptera, Chrysomelidae)". Entomography. 1: 1–221.
Riley, Edward G.; Clark, Shawn M.; Seeno, Terry N. (2003). Catalog of the leaf beetles of America north of Mexico (Coleoptera: Megalopodidae, Orsodacnidae and Chrysomelidae, excluding Bruchinae). Special Publication No. 1. The Coleopterists' Society. ISBN 978-0-9726087-1-8.
"Typophorini Report". Integrated Taxonomic Information System. Retrieved 2018-05-04.
"Typophorini Tribe Information". BugGuide.net. Retrieved 2018-05-04.
Moseyko, A. G.; Sprecher-Uebersax, E. (2010). "Eumolpinae". In Löbl, I.; Smetana, A. (eds.). Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea. Stenstrup, Denmark: Apollo Books. pp. 619–643. ISBN 978-87-88757-84-2.
Moseyko, A. G. (2012) [2011]. "To the knowledge of the leaf-beetle tribe Nodinini (Coleoptera, Chrysomelidae, Eumolpinae) from the Philippines and Borneo". Entomological Review. 92 (3): 315–328. doi:10.1134/S0013873812030086. S2CID 15106058.
Moseyko, A.G. (2010). "Medvedemolpus gen.n. — a new genus of Eumolpinae (Coleoptera: Chrysomelidae) from Philippines" (PDF). Russian Entomological Journal. 19 (1): 3–8. doi:10.15298/rusentj.19.1.02.
Moseyko, A. G. (2012). "Comments on the genus Aphilenia Weise in Reitter, 1889 (Coleoptera, Chrysomelidae, Eumolpinae), with a description of a new species from Russia". Entomological Review. 92 (7): 773–781. doi:10.1134/S0013873812070044. S2CID 15377841.
Chaboo, Caroline S.; Flowers, R. Wills (2015). "Beetles (Coleoptera) of Peru: A Survey of the Families. Chrysomelidae: Eumolpinae Hope, 1840". Journal of the Kansas Entomological Society. 88 (3): 375–379. doi:10.2317/kent-88-03-375-379.1. S2CID 87900852.
Bechyné, J. (1957). "Eumolpides neo-tropicaux de la collection du Museo civico di Storia naturale "Giacomo Doria" di Genova (Col. Phytophaga)". Annali del Museo Civico di Storia Naturale Giacomo Doria. 69: 226–247.
Gressitt, J. L. (1969). "Chrysomelid beetles from the Papuan subregion, 6 (Eumolpinae, 4)" (PDF). Pacific Insects. 11 (1): 1–31.
Sekerka, L. (16 September 2015). "Eumolpinae". Catálogo Taxonômico da Fauna do Brasil. PNUD. Retrieved 24 October 2021.
Bryant, G. E.; Gressitt, J. L. (1957). "Chrysomelidae of Fiji (Coleoptera)". Pacific Science. 11 (1): 2–91. hdl:10125/8611.
Weise, J. (1904). "Chrysomeliden und Coccinelliden aus Afrika". Archiv für Naturgeschichte. 70 (1): 35–62.
Jacobson, G. G. (1894). "Zur Chrysomeliden - Fauna der Umgegend vom See Issyk-kul". Deutsche Entomologische Zeitschrift. 1894 (1): 97–107.
Baly, J. S. (1860). "Descriptions of New Genera and Species of Eumolpidae". The Journal of Entomology. 1 (1): 23–36.
Baly, J. S. (1878). "Description of New Species and Genera of Eumolpidæ" (PDF). Zoological Journal of the Linnean Society. 14 (75): 246–265. doi:10.1111/j.1096-3642.1878.tb01834.x.
Morawitz, F. (1861). "Zur Kenntniss der russischen Eumolpiden". Horae Societatis Entomologicae Rossicae. 1: 159–164.
Jacoby, M. (1908). Bingham, C. T. (ed.). Coleoptera. Chrysomelidae. Vol. 1. The Fauna of British India, Including Ceylon and Burma. London: Taylor & Francis.
Baly, J. S. (1861). "Descriptions of new genera and species of Phytophaga". The Journal of Entomology. 1: 275–302.
Gressitt, J. L. (1967). "Chrysomelid beetles from the Papuan subregion, 4 (Eumolpinae, 2)" (PDF). Pacific Insects. 9 (2): 295–340.
Lefèvre, E. (1884). "[Cleoporus]". Bulletin de la Société Entomologique de France: LXXVI–LXXVII.
Lefèvre, É. (1885). "Eumolpidarum hucusque cognitarum catalogus, sectionum conspectu systematico, generum sicut et specierum nonnullarum novarum descriptionibus adjunctis". Mémoires de la Société Royale des Sciences de Liège. 2. 11 (16): 1–172.
Lefèvre, E. (1885). "Deux genres nouveaux d'Eumolpides [Phaedroides et Clypeolaria]". Bulletin de la Société Entomologique de France. 6. 5 (2): CXXV–CXXVI.
Weise, J. (1913). "Chrysomeliden und Coccinelliden" (PDF). Nova Guinea: Résultats de l'Expédition Scientifique Néerlandaiso à la Nouvelle Guinée. 9: 423–446.
Lefèvre, E. (1884). "Descriptions de quatre genres nouveaux et de plusieurs espèces nouvelles de coléoptères de la famille des eumolpides". Bulletin de la Société Entomologique de Belgique. 1884: cxciii–ccvi.
Bechyné, J. (1953). "Katalog der neotropischen Eumolpiden (Col. Phytoph. Chrysomeloidea)". Entomologische Arbeiten aus dem Museum G. Frey (in German). 4: 26–303.
Medvedev, Lev N. (2009). "New genera and species of Zeugophorinae and Eumolpinae (Coleoptera: Chrysomelidae) from New Guinea" (PDF). Stuttgarter Beiträge zur Naturkunde A, Neue Serie. 2: 371–408.
Askevold, I. S.; LeSage, L. (1990). "A taxonomic revision of the genus Jansonius Baly 1878: Taxonomic confusion and tribal reclassification (Coleoptera: Chrysomelidae: Eumolpinae)" (PDF). Revista Chilena de Entomología. 18: 29–37.
Maulik, S. (1931). "Coleoptera, Chrysomelidæ: Eumolpinæ, Galerucinæ and Halticinæ". Transactions of the Linnean Society of London. 2, Zoology. 19 (2): 241–260. doi:10.1111/j.1096-3642.1931.tb00128.x.
Weise, J. (1903). "Afrikanische Chrysomeliden". Archiv für Naturgeschichte. 69 (1): 197–226.
Lefèvre, E. (1884). "Melindea, Phascus, Pagria nn. genn". Bulletin de la Société Entomologique de France: LXV–LXVII.
Bechyné, J. (1951). "Liste provisoire des Eumolpides de Bolivie et observations diverses sur les espèces de l'Amérique du Sud (Col. Phytophaga)". Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei München. 2: 227–352.
Medvedev, L. N.; Moseyko, A. G. (2003). "A revision of the genus Phytorus Jacoby, 1884 (Chrysomelidae: Eumolpinae)" (PDF). Russian Entomological Journal. 11 (4): 401–407.
Jacoby, M. (1884). "Description of new genera and species of Phytophagous Coleoptera from the Indo-Malayan and Austro-Malayan sub-regions, contained in the Genoa Civic Museum. First Part". Annali del Museo Civico di Storia Naturale di Genova. 20: 188–233. doi:10.5962/bhl.part.6180.
Jacoby, M. (1897). "Further contributions to the knowledge of the phytophagous Coleoptera of Africa, including Madagascar, Part II". Journal of Zoology. 65 (3): 527–577. doi:10.1111/j.1096-3642.1897.tb03108.x.
Harold, E. von (1877). "Coleopterorum species novae". Mitteilungen der Münchener Entomologischen Vereins. 1: 97–111.
Zoia, S (2019). "New data on African Eumolpinae with description of new taxa from the collections of the Natural History Museum of Erfurt (Insecta: Coleoptera: Chrysomelidae)" (PDF). Vernate. 38: 305–333.
Baly, J. S. (1867). "Phytophaga Malayana; a revision of the phytophagous beetles of the Malay Archipelago, with descriptions of the new species collected by Mr. A. R. Wallace". Transactions of the Entomological Society of London. 3. 4: 1–300. doi:10.1111/j.1365-2311.1866.tb01857.x.
Lefèvre, É. (1885). "[New Taxa]". Bulletin de la Société entomologique de France. 1885: cxlvii.

Aslam, N. A. (1968). "Nomenclatorial notes on Chrysomeloidea (Coleoptera)". Journal of Natural History. 2 (1): 127–129. doi:10.1080/00222936800770691.