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Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Scarabaeinae
Tribus: Scarabaeini
Genus: Kheper
Species (26): K. aegyptiorum – K. aeratus – K. asceticus – K. bonellii – K. clericus – K. cupreus – K. cuvieri – K. devotus – K. erichsoni – K. festivus – K. intermedius – K. kalaharicus – K. laevistriatus – K. lamarcki – K. nigroaeneus – K. paganus – K. platynotus – K. porosus – K. prodigiosus – K. pustulosus – K. rolciki – K. sanctus – K. subaeneus – K. vanderkelleni – K. venerabilis – K. zurstrasseni
Name

Kheper Janssens, 1940

Authority for current placement: Forgie, Philips & Scholtz, 2005: 69 ('stat. nov.')

Kheper dung beetle

References
Primary references

Janssens 1940: Mém. Mus. Hist. nat. Belg. (2), 16:

Additional references

Deschodt, C.M.; Davis, A.L.V.; Scholtz, C.H. 2011: New dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) taxa from arid south-western Africa. Insect systematics & evolution, 42(3): 277–294. DOI: 10.1163/187631211X568173 Reference page. [see pp. 288-289]
Forgie, S.A.; Philips, T.K.; Scholtz, C.H. 2005: Evolution of the Scarabaeini (Scarabaeidae: Scarabaeinae). Systematic entomology 30(1): 60–96. DOI: 10.1111/j.1365-3113.2004.00273.x Reference page.

Links

Schoolmeesters P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on March 27, 2017]

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