Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea
Familia: Scarabaeidae
Subfamilia: Scarabaeinae
Tribus: Scarabaeini
Genus: Kheper
Species (26): K. aegyptiorum – K. aeratus – K. asceticus – K. bonellii – K. clericus – K. cupreus – K. cuvieri – K. devotus – K. erichsoni – K. festivus – K. intermedius – K. kalaharicus – K. laevistriatus – K. lamarcki – K. nigroaeneus – K. paganus – K. platynotus – K. porosus – K. prodigiosus – K. pustulosus – K. rolciki – K. sanctus – K. subaeneus – K. vanderkelleni – K. venerabilis – K. zurstrasseni
Name
Kheper Janssens, 1940
Authority for current placement: Forgie, Philips & Scholtz, 2005: 69 ('stat. nov.')
References
Primary references
Janssens 1940: Mém. Mus. Hist. nat. Belg. (2), 16:
Additional references
Deschodt, C.M.; Davis, A.L.V.; Scholtz, C.H. 2011: New dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) taxa from arid south-western Africa. Insect systematics & evolution, 42(3): 277–294. DOI: 10.1163/187631211X568173 Reference page. [see pp. 288-289]
Forgie, S.A.; Philips, T.K.; Scholtz, C.H. 2005: Evolution of the Scarabaeini (Scarabaeidae: Scarabaeinae). Systematic entomology 30(1): 60–96. DOI: 10.1111/j.1365-3113.2004.00273.x Reference page.
Links
Schoolmeesters P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on March 27, 2017]
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