Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Superfamilia: Curculionoidea
Familia: Curculionidae
Subfamilia: Entiminae
Tribus: Eupholini
Genus: Gymnopholus
Subgenera: G. (Gymnopholus) – G. (Niphetoscapha) – G. (Symbiopholus)
[list of subgenera after Riedel (2009: 3)]
Name
Gymnopholus Heller, 1901
Gender
masculine
Type species
Gymnopholus weiskei Heller, 1901, by monotypy
References
Template:Heller, 1901
Gressitt, J.L. 1966. Epizoic symbiosis: the Papuan weevil genus Gymnopholus (Leptopiinae) symbiotic with cryptogamic plants, oribatid mites, rotifers and nematodes. Pacific Insects 8(1): 221–280. PDF. Reference page.
Gressitt, J.L. 1977. Papuan weevil genus Gymnopholus: third supplement with studies in epizoic symbiosis. Pacific Insects 17(2–3): 179–195. PDF Reference page.
Gressitt, J.L. & Sedlacek, J. 1967. Papuan weevil genus Gymnopholus: supplement and further studies in epizoic symbiosis. Pacific Insects 9(3): 481–500. PDF. Reference page.
Gressitt, J.L. & Sedlacek, J. 1971. Papuan weevil genus Gymnopholus: second supplement with studies in epizoic symbiosis. Pacific Insects 12(4): 753–762. PDF. Reference page.
Riedel, A. 2009. Revision of the genus Penthoscapha Heller (Coleoptera, Curculionoidea, Entiminae, Eupholini) with notes on the genera of Eupholini from New Guinea. Zootaxa 2224: 1–29. Abstract & excerpt Reference page.
Riedel, A. & Kilmaskossu, A. 2017. Revision of the subgenus Niphetoscapha Heller of Gymnopholus Heller (Coleoptera, Curculionoidea, Entiminae, Eupholini) and a new species with epizoic symbiosis from West New Guinea. Zootaxa 4254(3): 339–356. DOI: 10.11646/zootaxa.4254.3.3. Reference page.
Gymnopholus is a genus of beetles in the family Curculionidae (true weevils) occurring in New Guinea. Many of them are notable for the growth of algae, diatoms, fungi, lichens and mosses on their backs[2](especially the subgenus Symbiopholus). The genus is divided into two subgenera (Gymnopholus and Symbiopholus) and contains the following species:[3]
Subgenus Gymnopholus Heller, 1901
Gymnopholus ajax
Gymnopholus angustus
Gymnopholus biformis
Gymnopholus brandti
Gymnopholus carinatus
Gymnopholus colmani
Gymnopholus cyphothorax
Gymnopholus divaricatus
Gymnopholus ellynae
Gymnopholus engabenae
Gymnopholus fulvospretus
Gymnopholus gemmifer
Gymnopholus glochidionis
Gymnopholus gressitti
Gymnopholus harti
Gymnopholus hornabrooki
Gymnopholus howcrofti
Gymnopholus integrirostris
Gymnopholus interpres
Gymnopholus ludificator
Gymnopholus magister
Gymnopholus mammifer
Gymnopholus macquarti
Gymnopholus marshalli
Gymnopholus muscosus
Gymnopholus nothofagi
Gymnopholus perspicax
Gymnopholus pulcher
Gymnopholus regalis
Gymnopholus rennii
Gymnopholus rostralis
Gymnopholus rubi
Gymnopholus sedlaceki
Gymnopholus seriatus
Gymnopholus setosus
Gymnopholus splendidus
Gymnopholus subnacreus
Gymnopholus suturalis
Gymnopholus suturalis suturalis
Gymnopholus suturalis angularis
Gymnopholus szentivanyi
Gymnopholus toxopei
Gymnopholus tricolor
Gymnopholus urticivorax
Gymnopholus vetustus
Gymnopholus weiskei
Gymnopholus acarifer
Gymnopholus algifer
Gymnopholus audax
Gymnopholus botanicus
Gymnopholus carolynae
Gymnopholus cheesemanae
Gymnopholus didiman
Gymnopholus euryae
Gymnopholus fallax
Gymnopholus fungifer
Gymnopholus hepaticus
Gymnopholus herbarius
Gymnopholus herbarius herbarius
Gymnopholus herbarius oribatifer
Gymnopholus huttoni
Gymnopholus kokodae
Gymnopholus lichenifer
Gymnopholus magnus
Gymnopholus nitidus
Gymnopholus nodifer
Gymnopholus piorae
Gymnopholus praecox
Gymnopholus rebeccae
Gymnopholus reticulatus
Gymnopholus rugicollis
Gymnopholus schefflerae
Gymnopholus schuurmannsiae
Gymnopholus senex
Gymnopholus shungolus
Gymnopholus symbioticus
Gymnopholus vegetatus
Gymnopholus zoarkes
References
Marshall, Guy A. K. Curculionid Genus Gymnopholus (Coleoptera)
Gressitt, J.L. (1977). "Papuan weevil genus Gymnopholus: third supplement with studies in epizoic symbiosis" (PDF). Pacific Insects. 17 (2–3): 179–195.
Setliff, G.P. (2007). "Annotated checklist of weevils from the Papuan region (Coleoptera, Curculionoidea)" (PDF). Zootaxa. 1536: 1–296.
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